Paranomina hendeli, Mcalpine, 2019

Mcalpine, David K., 2019, Review of the Species of Paranomina (Diptera: Lauxaniidae), Records of the Australian Museum 71 (3), pp. 71-85 : 77-78

Paranomina hendeli sp. nov.

http://zoobank. org/NomenclaturalActs/ 71CCEB73-4254-41A3-ABE1-49B495F4A848

Figs 10 –12

Holotype ♂. New South Wales: Royal National Park [eastern parts, probably near Flat Rock Creek], near Sydney , 13.vii.1971, D.K.M., G.A.H. ( AM K.456341) . Paratypes. New South Wales: 105 (34♂♂ 26♀♀ in AM K.515447–506, 25♂♂ 20♀♀ in QM), Royal National Park (including Flat Rock Creek, Cascade Creek , Gundamain ), v vi vii viii x xi xii. 1963–1977 , G.D., D.K.M.; 3 (2♂♂ 1♀ in QM), Engadine , Oct. 1977 , G.D.; 11 (3♂♂ 1♀ in AM K.515507–510, 2♂♂ 1♀ in ANIC, 3♂♂ 1♀ in QM) Ku-ring-gai Chase National Park (including near Terrey Hills, McCarr’s Creek , West Head ), i viii ix. 1960–1986 , G.D., D.H.C., D.K.M.; 1♂ ( ANIC), Deep Creek, near Narrabeen , xii 1961 , D.H.C.; 15 (7♂♂, 8♀♀ in AM K.556212–226) Halfway Creek , 18 miles [c. 30 km] S of Grafton, v.1972 , D.K.M.

Other material examined (localities only given). Queensland: 3♂♂ 1♀ ( QM) Cholmondeley Creek, 11 km SW of Heathlands; 1♀ ( QM) Gunshot Ck , 13 km NW Heathlands ; 2♂♂ ( AM) 4 miles [c. 6 km] NE of Mount Lamond , Iron Range district ; 3♂♂ 2♀♀ ( QM) near Coen Aerodrome ; 2♂♂ ( AM) Bald Hills Station , 6 km NW of Isabella Falls ; 3♂♂ 3♀♀ ( AM) Clohesy Road , near Kuranda ; 3♂♂ 2♀♀ ( AM) 9 miles N [probably more accurately 14 km NW] of Rollingstone ; 1♂ 1♀ ( AM) 30 miles [c. 48 km] N of Marlborough ; 1♂ 1♀ ( AM) Deception Bay ; 1♂ ( QM) Tibrogargen Ck ; 1♀ ( QM) Brown Lake , North Stradbroke Island ; 1♂ ( QM) 13 km N Dunwich, North Stradbroke Island ; 1♂ ( QM) Acacia Ridge .

Description (male, female). Agreeing with generic description given above; size slightly smaller than that of most Paranomina species.

Coloration (typical New South Wales populations) generally tawny with brown tinge (a few specimens paler). Head: parafacial and anterior cheek region paler, often white to creamy. Antenna largely dark brown to greyish brown, arista blackish. Prelabrum tawny to brown; palpus brownish, often darker distally. Thorax without definite dark brown zones; mesoscutum tawny-brown, usually with pair of longitudinal tawny-yellow zones; pleura usually largely tawny brown with greyish pruinescence; humeral callus yellow to tawny yellow on c. dorsal half, becoming greyish brown below, this somewhat darker tone extending on to pleura. Legs tawny yellow. Wing hyaline to faintly smoky; halter yellow. Abdomen normally tawny-yellow, often discoloured in preserved specimens.

Head slightly shorter and more rotund in profile than that of P. stuckenbergi and related species; cephalic chaetotaxy typical of genus.

Thorax structurally typical of genus.

Male postabdomen (New South Wales population). Surstylar-epandrial complex somewhat smaller than in related species; surstylus distally simple in outline, slightly narrowly rounded, with rather coarse, irregularly placed setulae; paired basal hypandrial sclerites stout and strictly transverse ( Fig. 11 ); each aedeagal rod almost straight, somewhat shorter and stouter than in related species, with stout, subterminal lateral tooth, more narrowed terminally and joined to other of its pair by transparent membrane almost to apex; aedeagal apodeme long and slender.

Dimensions. Total length, males 3.7–4.2 mm, females 3.4–4.2 mm; length of thorax, males 1.5–1.7 mm, females 1.5–1.6 mm; length of wing, males 4.0– 4.4 mm, females 3.9–4.2 mm.

= 0.2 mm. (11) Hypandrium and aedeagus, anterior view, scale = 0.2 mm.

Distribution. New South Wales: coastal districts, as far south as Royal National Park (c. 34°12'S). Queensland: eastern coastal districts, as far north as Heathlands (c. 11°45'S).

Notes. Paranomina hendeli is, on the average, smaller than other species of the genus, and is probably consistently smaller than P. unicolor . The usually more greyish toning of the thoracic pleura (at least for the non-tropical populations) is somewhat different from that of the tawny-yellow pleural coloration of the partly sympatric P. danielsi and P. stuckenbergi , and the slightly darker greyish brown shading of the ventral half of the humeral callus of P. hendeli is absent in those two species. This condition of P. hendeli is distinct from the dark brown spot on the lower part of the humeral callus present in P. mouldsorum . As indicated above, the form of the surstylus and the aedeagal complex enables accurate identification of males of P. hendeli , including those from tropical populations.

The specific epithet refers to Friedrich Hendel, the original author of Paranomina and basic contributor to the systematic study of Lauxaniidae and other dipterous families.

Figures

Figures 10, 11. Paranomina hendeli, male, Royal National Park. (10) Epandrium with surstyli, posterior view, scale