Phanolinopsis fassli ( Bernhauer, 1917 ) Chatzimanolis, 2017

Chatzimanolis, Stylianos, 2017, And then there were six: a revision of the genus Phanolinopsis Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae), Zootaxa 4323 (1), pp. 49-67 : 59-60

Phanolinopsis fassli ( Bernhauer, 1917) , new combination

( Figs. 3 , 9 , 15 , 31–33 , 44 )

Trigonopselaphus fassli Bernhauer, 1917: 113 .

Phanolinus fassli (Bernhauer) ; Scheerpeltz, 1933: 1415. Trigonopselaphus fassli Bernhauer ; Scheerpeltz, 1972: 43. Torobus fassli (Bernhauer) ; Herman 2001: 29.

Type material. Holotype, female, with labels: “ Columbia occ. Cali. Fassl” / “ Cañon del Monte Tolima, 1700m, iiiii.1910” / “ fassli Bern. Typus ” / “ Chicago NHMus, M. Bernhauer Collection ” / HOLOTYPE teste A . Westricht 2015 GDI imaging Project ” / “PHOTOGRAPHED Kelsey Keaton 2015 Emu Catalog ” / “FMNHINS3047816 Field Museum Pinned ” / “ Phanolinopsis fassli (Bernhauer) det. Chatzimanolis 2017”. In the collection of FMNH. Bernhauer (1917) added the following to the locality label on the specimen: “ Umgebung von Cali am Cauca ”, and mentioned that he had only a single female specimen, which is the holotype .

Additional Material. Unknown Country: Coll. Kraatz (1 ♂ SDEI).

Diagnosis. Among all species of Phanolinopsis , P. fassli and P. norahae are easily distinguished from other species due to the lack of punctures on the disc of the pronotum in these two species. Phanolinopsis fassli can be distinguished from P. norahae based on the following: head and pronotum with weak micropunctation ( Fig. 9 ; much starker in P. norahae , Fig. 12 ); reduced microsculpture between eyes and postmandibular ridge ( Fig. 15 ; not reduced in P. norahae , Fig. 18 ); tip of paramere flat ( Figs. 31, 33 ; more rounded in P. norahae , Figs. 40, 42 ). The two species also differ in coloration ( Figs. 3 , 6 ).

Description. Body length 12.6–14.0mm. Head, pronotum and elytra metallic purple-brown with green overtones. Mouthparts, antennae, mesoscutellum, ventral surface of thorax, legs and abdomen brown. Posterior half of abdominal terga slightly lighter color; posterior 1/4 of segment VII and segment VIII orange. Head transverse, width: length ratio = 1.32. Epicranium with transverse and polygon-shaped microsculpture and sparse micropunctures; with large to medium-sized punctures around margin of head (becoming more numerous near posterior margin), but with no other punctures on epicranium. Eyes medium-sized, length of eyes / length of head ratio = 0.39, distance between eyes as wide as 2.67 times length of eye. Area between postmandibular ridge and eye (lateral side of head) wide, without polygon-shaped microsculpture but with sparse transverse microsculpture; posterolateral corner of head not pointed. Antennomeres 1–8, longer than wide; antennomeres 9–11 missing. Neck with micropunctures, microsculpture, and with dense small punctures. Pronotum subquadrate, width: length ratio = 0.98; surface of pronotum uniformly covered with sparse micropunctures and dense polygon-shaped microsculpture; appearing matte due to microsculpture. Pronotum with few large punctures around margin; disc of pronotum impunctate. Elytra with medium-sized punctures (about 12–13 punctures / elytron width); punctures almost confluent. Elytra appearing shiny; with sparse longitudinal microsculpture. Abdominal terga with 1–2 rows of small punctures. Male secondary sexual structures with sternum VIII having shallow V-shaped emargination medially; sternum IX with deep V-shaped emargination medially. Female without obvious sexual structures. Aedeagus as in Figs. 31–33 ; in dorsal view paramere converging to flat tip; paramere shorter and narrower (apically) than median lobe; in lateral view paramere concave apically; paramere with peg setae as in Fig. 33 . Median lobe in dorsal view wide, converging to rounded apex, with two broad teeth apically; in lateral view becoming narrower near apex.

Distribution. Known from the type locality of west Colombia in the mountainous surrounding of Cali.

Habitat. Unknown, but given the high altitude (1700m), the species is probably found in cloud forests.

Figures

FIGURE 44. Distribution map of P. fassli (Bernhauer), P. metaksenios Chatzimanolis and P. norahae Chatzimanolis.

FIGURES 3 – 4. Habitus photographs of species of Phanolinopsis Scheerpeltz. 3. The holotype of P. fassli (Bernhauer), total length = 14.0 mm. 4. The holotype of P. goniakos Chatzimanolis, total length = 12.1 mm.

FIGURES 7 – 12. Heads and pronota of Phanolinopsis Scheerpeltz. 7. P. discedens (Sharp). 8. P. erythros Chatzimanolis. 9. P. fassli (Bernhauer). 10. P. goniakos Chatzimanolis. 11. P. metaksenios Chatzimanolis. 12. P. norahae Chatzimanolis. Not to scale.

FIGURES 13 – 18. Lateral view of heads of Phanolinopsis Scheerpeltz indicating the differences in the temple area and specifically the distance between the eye and postmandibular ridge. White arrows point to the extent of the postmandibular ridge. 13. P. discedens (Sharp). 14. P. erythros Chatzimanolis. 15. P. fassli (Bernhauer). 16. P. goniakos Chatzimanolis. 17. P. metaksenios Chatzimanolis. 18. P. norahae Chatzimanolis. Not to scale.

FIGURES 31 – 33. Aedeagus of P. fassli (Bernhauer). 31. Dorsal view. 32. Lateral view. 33. Detail of paramere, ventral view.

FIGURES 40 – 42. Aedeagus of P. norahae Chatzimanolis. 40. Dorsal view. 41. Lateral view. 42. Detail of paramere, ventral view.

FIGURES 5 – 6. Habitus photographs of species of Phanolinopsis Scheerpeltz. 5. The holotype of P. metaksenios Chatzimanolis, total length = 12.5 mm. 6. The holotype of P. norahae Chatzimanolis, total length = 13.5 mm.