Phortica (Ashima) speculum ( Máca & Lin, 1993 )

Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit & Katoh, Toru, 2020, Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species, Zootaxa 4789 (1), pp. 1-54 : 21

16) Phortica (Ashima) speculum ( Máca & Lin, 1993)

Amiota (Phortica) speculum Máca & Lin, 1993b: 172 .

Phortica speculum: Chen et al., 2005b: 420 .

Phortica (Phortica) speculum: Cheng et al., 2008: 621 .

Phortica (Ashima) speculum: Chen & Máca, 2012: 507 .

Diagnosis. Arista apically with distinct, leaf-like expansion narrower than long and apically tapering and pointed and 2–3 long and 1–3 minute branches dorsally and ventrally, respectively; longest, dorsal branch longer than longest seta on pedicel [“Fig. 19 (mislabeled as 18 on the plate)” in Cheng et al. 2008]; posterior postgonite apically blunt.

Supplementary description (not repeating characters common to P. pavriarista ). Supracervical setae 8–11. Dorsomedial, tentorial apodeme slightly longer than 1/2 basal, parallel portion of dorsolateral, tentorial apodeme. Cibarial, medial sensilla approximately 8 per side; posterior sensilla 6–7 per side. All tarsi with gray tarsomeres IV and V. Abdominal sternite III wider than long (ch.22-1), widest among sternites (ch.23-1), with dense setulae on medial surface (ch.26-1).

The antisymmetry is observed in the postgonites: in A-type, the left lateral lobe bears 2 anterior postgonites, and the right lobe 1 posterior and 1 anterior postgonites (the holotype; “ Fig. 3 ” in Máca & Lin 1993b); but in B-type, vice versa (“ Fig. 4 ” in Máca & Lin 1993b).

Specimens examined. Japan: 1♂ (A-type), Iriomote , Ryukyu Is., x.1994, M. T . Kimura leg. ( SEHU) . China: 1♂ (A-type), Mt. Emei , Sichuan, 550 m a.s.l., 18.vii.1992, M.J. Toda leg. ( SEHU) .

Distribution. Ryukyu Is., Taiwan, China (Shaanxi, Jiangxi, Fujian, Guangdong, Guangxi, Guizhou, Sichuan, Yunnan).

Remarks. This species resembles P. pavriarista in the morphology of the aedeagal sheath and postgonites, but can be distinguished from it by the diagnostic characters.

Figures

FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm.

FIGURE 4. Phortica (Ashima) andreagigoni Toda & Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm.