Pinodytes isabella Peck & Cook, 2011

Peck, Stewart B. & Cook, Joyce, 2011, Systematics, distributions and bionomics of the Catopocerini (eyeless soil fungivore beetles) of North America (Coleoptera: Leiodidae: Catopocerinae) 3077, Zootaxa 3077 (1), pp. 1-118 : 46-47

Pinodytes isabella Peck & Cook , new species

( Figs. 39 , 351–359 )

Type material. Holotype: male ( WFBM). UNITED STATES. Idaho: Clearwater Co., Isabella Creek , V-29-1985, (B.F. cedar litter), F.W. Merickel . Paratypes (8): same data as holotype ( WFBM) .

Material examined. We have examined 9 specimens.

Distribution. Specimens ( Fig. 359 ) are known only from Clearwater County, in northern Idaho.

Diagnostic description. Total length 1.62–1.74 mm; greatest width 0.74–0.86 mm. Reddish brown; elongateoval in shape ( Fig. 39 ). Head. Finely, sparsely punctate; shining; microsculpture reticulate to substriate. Eyes absent. Antenna ( Fig. 351 ) with antennomere 2 larger than 3; antennomere 5 larger than 4 and 6; antennomere 7 clearly larger than 8; antennomeres 9 and 10 lack visible sensory vesicles. Pronotum. Finely punctate, punctures separated by 3–5 diameters, occasional larger punctures near margins; shining, with reticulate microsculpture. Widest at about basal one-fourth; sides rounded; apical margin emarginate, basal margin nearly straight; apical angles narrowly rounded, basal angles nearly right-angled. Elytra. Punctures mainly larger than on pronotum, irregularly arranged, finer apically; punctures joined transversely by fine strioles. Joined elytra slightly wider than pronotum; widest in basal one-third, then narrowing to apex. Legs. Protibia ( Fig. 352 ) widened apically; apex sparsely spinose; apical one-half of inner margin finely, densely spinose. Mesotibia ( Fig. 353 ) straight, evenly widened to apex; strongly spinose apically and on outer margin. Metatibia ( Fig. 354 ) weakly curved, expanded apically; spinose on apical one-half. Metafemur ( Fig. 354 ) narrow. Male protarsomeres ( Fig. 352 ) not expanded; protarsomere 1 about as long as 2 and 3 combined; with elongate setae laterally and a double row of thin, colorless, transverse phanerae ventrally. Mesotarsomeres without phanerae. Venter. Mesoventrite ( Fig. 358 ) carinate; longitudinal carina shallowly emarginate, bearing a small sharp tooth; excavation behind transverse carina. Male genitalia. Median lobe of aedeagus ( Fig. 355, 356 ) strongly curved dorsoventrally beyond midpoint; broad and flattened before narrow apex. Inverted internal sac with a mass of broad, flattened setae. Parameres ( Figs. 355, 356 ) slender, not reaching apex of median lobe; each bearing two apical setae. Spermatheca. Cylindrical ( Fig. 357 ), curved.

Etymology. The name isabella , a noun in apposition, refers to the type locality of this species.

Figures

FIGURE 359. Distributions of species in the Pinodytes cryptophagoides species group in northwestern United States. P. isabella, open triangle in northern Idaho. P.orca, closed square in southwestern Oregon. P.setosus, closed triangles in southwestern Oregon and northwestern California. P. shoshone, closed circles in northern Idaho. P. sinuatus, open squares in southwestern Oregon.

FIGURES 37–43. Pinodytes habitus photos. 37. P.garibaldi, total length 2.80 mm. 38. P. haidagwaii, total length 3.04 mm. 39. P. isabella, total length 2.18 mm. 40. P.orca, total length 4.20 mm. 41. P.setosus, total length 1.96 mm. 42. P. shoshone, total length 2.18 mm. 43. P. sinuatus, total length 2.28 mm.

FIGURES 351–358. Pinodytes isabella. 351. Male right antenna, dorsal. 352. Male right protibia and protarsus. 353. Male right mesotibia. 354. Male right metafemur and metatibia. 355. Aedeagus, lateral. 356. Aedeagus, dorsal. 357. Spermatheca. 358. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 357), 0.3 mm (Figs. 355, 356, 358), 0.4 mm (Figs. 351–354).