Taurohelops incultus ( Allard, 1877 ) Keskin, Bekir & Nabozhenko, Maxim, 2015

Keskin, Bekir & Nabozhenko, Maxim, 2015, The New Genus Taurohelops Keskin and Nabozhenko (Coleoptera: Tenebrionidae) from Anatolia, Turkey, The Coleopterists Bulletin (mo 14) 69, pp. 83-92 : 84-88

Taurohelops incultus ( Allard, 1877) , new combination

( Figs. 1–14 , 23–25 )

Stenomax incultus Allard 1877: 132 . Seidlitz 1896:724, 728 ( Helops ); Reitter 1922: 136 ( Cylindronotus

3) Coeloconic pronotal sensillum; 4) Trichoid strial sensillum; 5) Trichoid and coeloconic (arrow) sensillae on pronotum. ( Odocnemis )); Nabozhenko 2008: 37 ( Odocnemis ); Nabozhenko and Löbl 2008: 244 ( Odocnemis ). Holotype female at DEI labeled “ incultus // Külek//coll. Kraatz//coll. DEI Müncheberg// Holotypus ”.

Helops lordiscelis Reitter 1900: 157 . Lectotype male (here designated) at HNHM labeled “ Asia minor, Bulghar Magara, v. Bodemeyer//Coll. Reitter// Holotypus 1900, Odocnemis lordiscelis Reitter // incultus All, Coll Reitter ”.

Material Examined. 58♂♂, 45♀♀ ( ZIN, CN): Turkey, Içel Province, above Arslanköy, N 37°02′242″ E 34°17′191″, 1800 m, 16.iv.2007, on Cedrus libani , Juniperus excelsa , leg. M.V. Nabozhenko ; 3♂♂, 1♀ ( ZDEU, dry collection) , 1♂, 6♀♀ ( ZDEU, ethanol): same locality, 18.05.2009, leg. M.V. and S.V. Nabozhenko, B. Keskin ; 2♂♂ ( CN): Turkey, Karaman Province, Ayranci District , 10 km S Kıraman, 37°11′851″N 33°59′208″E, 1962 m, 19.iv.2008, leg. M.V. Nabozhenko ; 2♂♂ ( CN): Turkey, Içel Province, Güzeloluk , 1400 m, 16–18. v.2008, leg. I.V. Shokhin ; 49♂♂, 40♀♀ ( ZDEU, dry collection) , 2♂♂, 21♀♀ ( ZDEU, ethanol) , 9♂♂, 4♀♀ ( ZIN, CN): Turkey, Içel Province, near Çamlıyayla, N 37°10′307″/E 34°31′239″, 1700 m, on Cedrus libani , 18.v.2009, leg. M.V. and S.V. Nabozhenko, B. Keskin ; 1♀ ( ZDEU, dry collection): Turkey, Içel Province, Gülekboğazı , 37°20′17″ N 34°45′48.8″ E, 1520 m, 17.v.2010, leg. M.V. and S.V. Nabozhenko, B. Keskin ; 4♂♂ 15 ♀♀ ( ZDEU, dry collection) 3♂♂ 13 ♀♀ ( ZDEU, ethanol): Turkey, Içel Province, Toros (above town), N 36°54′28″, E 34°05′45″, 1709 m, 31.05.2011, leg. B. Keskin ; 1♂ 2 ♀♀ ( ZDEU, dry collection): Turkey, Içel Province, Sertavul Pass , 1441m, N 36°51′14″, E 33°17′50″, 30.05.2011, leg. B. Keskin.

Redescription. Male. Body length 13–19 mm, width 3.7–4.5 mm. Body slender, subparallel, strongly elongate, brown, with dull shine. Head: Widest at eye level. Eyes large, convex. Head width 1.8 times width of interocular space. Genae strongly rounded. Outer margin of head between genae and clypeus with deep rectangular sinuation. Frontoclypeus strongly depressed. Punctation of head moderately coarse, sparse on frons (puncture diameter less than distance between punctures), dense on other surfaces (puncture diameter 1.5–2.0 times as long as distance between punctures). Antennae long, with 5 apical antennomeres extending beyond base of pronotum, reaching middle of elytra. Thorax: Pronotum s ubquadrate (1.04 times as wide as long), widest before middle. Lateral margins weakly rounded in widest place and straight to base and anterior margin; anterior margin widely emarginate, base emarginate in middle. Anterior angles weakly obtuse, with rounded apex; posterior angles almost right, distinct on apex. All margins narrowly regularly beaded. Disc weakly convex, often with middle line. Punctation of disc fine and sparse, distance between punctures 2 or more times as long as puncture diameter. Prothoracic hypomera with fine microwrinkles and clear sparse punctation. Prosternal process moderately convex. Hind wings absent. Elytra: Strongly elongate, 2.8 times as long as wide, 1.55 times as wide as head, 1.2 times as wide and 3.5 times as long as pronotum. Striae with round punctures not merged in entire furrows. Punctation of interstriae fine and sparse, with 2 punctures placed in transverse interval. Interstriae (except at middle of elytra) with granules with celoconic sensilla. Interstria 8 keel-shaped on apex, connected with elytral margin. Epipleura strongly narrowing apically, extending to elytral apex. Elytra without mucro. Dorsal epipleural carina visible only near base and apex of elytra where it abutts interstria 8. Venter: Mesepimera with coarse and dense punctation. Metepisterna and metaventrite with sparse fine punctation; metaventrite with depression in middle and convex intercoxal process. Abdominal ventrites with fine and sparse punctation and fine longitudinal wrinkles on sides. Abdominal ventrites 3 and 4 depressed on sides, their posterior margin strongly elevated. Membranes between ventrites 3–5 very wide. Abdominal ventrite 5 strongly convex, inwardly curved, with weakly bisinuate apical margin. Legs: Trochanters (all legs) without hair brush, with 1 long seta. Protibiae bent and flattened (viewed from flexion side), depressed and densely pubescent on inner side. Mesotibiae bent (in dorsal view). Metatibiae bent and thickened in basal third (in dorsal view) and weakly bent from flexion side, flattened and strongly pubescent on inner side of apical 2/3. Tarsi long, not widened, with dense brush on plantar surface.

Female. Body length 12–17 mm, width 4.5–6 mm. Body more robust. Head width 1.4 times width of interocular space. Antennae shorter, with only 3 apical antennomeres extending beyond base of pronotum, reaching basal quarter of elytra. Pronotum more transverse (1.2 times as wide as long), widest in middle (rarely after middle), narrowed to anterior margin. Punctation of pronotum moderately coarse and dense (puncture diameter subequal to distance between them). Elytra more wide (1.8 times as long as wide), 2 times as wide as head, 1.3 times as wide and 2.90–2.95 times as long as elytra. Protibiae simple, straight, not bent, not flattened; meso- and metatibiae weakly bent, with simple pubescence.

Distribution. Turkey: central part of Central Taurus Mountains ( Fig. 28 ).

Bionomics. Adult specimens of this species were collected on trunks of C. libani and rarely Juniperus excelsa Pursh (Cupressaceae) between 20:00 and 00:00 hrs in April and May. They feed on lichens on the trunks.

Figures

Figs. 1–5. Taurohelops incultus. 1) Apex of right elytron with convex interstriae 8; 2) Celoconic elytral sensillum;

Figs. 6–13. Taurohelops incultus, male structures. 6) Abdominal ventrite 5, posterior view; 7) Abdominal ventrites 3–5, lateral view; 8) Sternite VIII, ventral view; 9) Sternite VIII, lateral view; 10) Aedeagus, ventral view; 11) Aedeagus, lateral view; 12) Gastral spicula, vetral view; 13) Gastral spicula, lateral view. Scale bars = 1 mm.

Fig. 14. Taurohelops incultus, female genital tubes. bds = basal duct of spermatheca; g = spermathecal gland; ov = oviduct; s = spermatheca.

Figs. 23–27. Taurohelops species, habitus. 23) T. incultus, female, holotype; 24) T. incultus, male; 25) T. incultus, female; 26) T. triplehorni, male; 27) T. triplehorni, female.

Fig. 28. Distribution map of Taurohelops species. Circle = T. incultus; square = T. triplehorni.