Varunidae, Taxa Only, Exact Match: 80 Treatments

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Decapoda   new species  Hwang, Jiuan-Jiuan & Takeda, Masatsune, 1986, A new freshwater crab of the family Graspidae from Taiwan, Proceedings of the Japanese Society of Systematic Zoology 33, pp. 11-18 : 12 12
Varunidae     Brösing, Andreas, 2010, Recent developments on the morphology of the brachyuran foregut ossicles and gastric teeth, Zootaxa 2510, pp. 1-44 : 33-34 33-34
Orcovita     Davie, Peter J. F. & Ng, Peter K. L., 2012, Two New Species Of Orcovita (Crustacea: Decapoda: Brachyura: Varunidae) From Anchialine Caves On Christmas Island, Eastern Indian Ocean, Raffles Bulletin of Zoology 60 (1), pp. 57-70 : 58-62 58-62
Cyclograpsus     In, First Published, 2022, Brachyuran crabs (Crustacea, Decapoda) from the remote oceanic Archipelago Trindade and Martin Vaz, South Atlantic Ocean, Zootaxa 5146, pp. 1-129 : 104 104
Varunidae     Castro, Peter, 2011, Catalog of the anomuran and brachyuran crabs (Crustacea: Decapoda: Anomura, Brachyura) of the Hawaiian Islands 2947, Zootaxa 2947 (1), pp. 1-154 : 121 121
Eriocheir ogasawaraensis   sp. nov.  Komai, T., Yamasaki, I., Kobayashi, S., Yamamoto, T. & Watanabe, S., 2006, Eriocheir ogasawaraensis Komai, a new species of mitten crab (Crustacea: Decapoda: Brachyura: Varunidae) from the Ogasawara Islands, Japan, with notes on the systematics of Eriocheir De Haan, 1835, Zootaxa 1168, pp. 1-20 : 3-19 3-19
Cyclograpsus henshawi     Castro, Peter, 2011, Catalog of the anomuran and brachyuran crabs (Crustacea: Decapoda: Anomura, Brachyura) of the Hawaiian Islands 2947, Zootaxa 2947 (1), pp. 1-154 : 121 121
Cyclograpsus integer     Mantelatto, Fernando L., Tamburus, Ana Francisca, Magalhães, Tatiana, Buranelli, Raquel C., Terossi, Mariana, Negri, Mariana, Castilho, Antonio L., Costa, Rogério C. & Zara, Fernando J., 2020, Checklist of decapod crustaceans from the coast of the São Paulo state (Brazil) supported by integrative molecular and morphological data: III. Infraorder Brachyura Latreille, 1802, Zootaxa 4872 (1), pp. 1-108 : 25-26 25-26
Gopkittisak gallardoi   comb. nov.  Naruse, Tohru & Clark, Paul F., 2009, Establishment of a new genus for Asthenognathus gallardoi Serène & Soh, 1976 within Gaeticinae Davie & N. K. Ng, 2007 (Crustacea: Decapoda: Brachyura: Varunidae), Zootaxa 1987, pp. 61-68 : 63-67 63-67
Pseudograpsus crassus     Lagare, Nickel Jean S., Mapi-Ot, Emmarie F., Molina, Ziljih S., Neri, Jemateo B., Nuñeza, Olga M. & Mendoza, Jose Christopher E., 2020, On a collection of freshwater and estuarine crabs (Crustacea: Brachyura) from Mindanao Island, the Philippines, Zootaxa 4868 (3), pp. 301-330 : 314 314
Hemigrapsus nudus     Wicksten, Mary K., 2012, Decapod Crustacea of the Californian and Oregonian Zoogeographic Provinces 3371, Zootaxa 3371, pp. 1-307 : 265 265
Brachynotus     McLay, C. L. & Schubart, C. D., 2004, Short Communication On the correct name for Hemigrapsus edwardsii (Hilgendorf, 1882) (Brachyura: Varunidae) from New Zealand, Journal of Natural History 38 (6), pp. 695-704 : 697-698 697-698
Thalassograpsus harpax     Naderloo, Reza & Türkay, Michael, 2012, Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: Faunistics, Biodiversity and Zoogeography 3374, Zootaxa 3374 (1), pp. 1-67 : 7 7
Sestrostoma balssi     Kornienko, Elena S. & Korn, Olga M., 2011, Morphology of the complete larval development of the symbiotic crab Sestrostoma balssi (Shen, 1932) (Varunidae: Gaeticinae), Zootaxa 2990, pp. 45-58 : 46-55 46-55
Hemigrapsus penicillatus     McDermott, John J., 2011, Parasites of shore crabs in the genus Hemigrapsus (Decapoda: Brachyura: Varunidae) and their status in crabs geographically displaced: a review, Journal of Natural History 45 (39 - 40), pp. 2419-2441 : 2425-2426 2425-2426
Pyxidognathus granulosus     Lagare, Nickel Jean S., Mapi-Ot, Emmarie F., Molina, Ziljih S., Neri, Jemateo B., Nuñeza, Olga M. & Mendoza, Jose Christopher E., 2020, On a collection of freshwater and estuarine crabs (Crustacea: Brachyura) from Mindanao Island, the Philippines, Zootaxa 4868 (3), pp. 301-330 : 315-318 315-318
Metaplax elegans     Shih, Hsi-Te, Hsu, Jhih-Wei, Wong, Kingsley J. H. & Ng, Ngan Kee, 2019, Review of the mudflat varunid crab genus Metaplax (Crustacea, Brachyura, Varunidae) from East Asia and northern Vietnam, ZooKeys 877, pp. 1-29 : 1 1
Cyclograpsus integer     In, First Published, 2022, Brachyuran crabs (Crustacea, Decapoda) from the remote oceanic Archipelago Trindade and Martin Vaz, South Atlantic Ocean, Zootaxa 5146, pp. 1-129 : 104-105 104-105
Varunidae     GUINOT, DANIÈLE, TAVARES, MARCOS & CASTRO, PETER, 2013, Significance of the sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura), with new nomina for higher-ranked podotreme taxa, Zootaxa 3665 (1), pp. 1-414 : 143-144 143-144
Pseudograpsus setosus     Tasso, Vicente, El Haddad, Mustapha, Assadi, Carolina, Canales, Remy, Aguirre, Luis & Velez-Zuazo, Ximena, 2018, Macrobenthic fauna from an upwelling coastal area of Peru (Warm Temperate South-eastern Pacific province - Humboldtian ecoregion), Biodiversity Data Journal 6, pp. 28937-28937 : 28937 28937
Gopkittisak   gen. nov.  Naruse, Tohru & Clark, Paul F., 2009, Establishment of a new genus for Asthenognathus gallardoi Serène & Soh, 1976 within Gaeticinae Davie & N. K. Ng, 2007 (Crustacea: Decapoda: Brachyura: Varunidae), Zootaxa 1987, pp. 61-68 : 62-63 62-63
Proexotelson tokoroi   sp. nov.  Naruse, Tohru, 2015, Description of a new genus and a new species of gaeticine crab (Crustacea: Brachyura: Varunidae) from the Ryukyu Islands, and a review of Acmaeopleura Stimpson, 1858, and Sestrostoma Davie & N. K. Ng, 2007, Zootaxa 3925 (2), pp. 211-228 : 222-224 222-224
Pseudohelice latreillii     Hsu, Jhih-Wei, Shih, Hsi-Te & Innocenti, Gianna, 2022, A review of the mud crab genus Pseudohelice Sakai, Türkay & Yang, 2006 (Crustacea: Brachyura: Varunidae), with redescription of Cyclograpsus latreillii H. Milne Edwards, 1837, from the western Indian Ocean, Raffles Bulletin of Zoology 70, pp. 94-107 : 100-104 100-104
Varunidae     Alves, Maria De Lourdes, Ramos-Porto, Marilena & Viana, Girlene Fábia Segundo, 2008, Checklist of the Decapods (Crustacea) from the Fernando de Noronha Archipelago, Brazil, Zootaxa 1881, pp. 43-68 : 65 65
Orcovita hicksi   sp. nov.  Davie, Peter J. F. & Ng, Peter K. L., 2012, Two New Species Of Orcovita (Crustacea: Decapoda: Brachyura: Varunidae) From Anchialine Caves On Christmas Island, Eastern Indian Ocean, Raffles Bulletin of Zoology 60 (1), pp. 57-70 : 62-65 62-65
Utica gracilipes     Lagare, Nickel Jean S., Mapi-Ot, Emmarie F., Molina, Ziljih S., Neri, Jemateo B., Nuñeza, Olga M. & Mendoza, Jose Christopher E., 2020, On a collection of freshwater and estuarine crabs (Crustacea: Brachyura) from Mindanao Island, the Philippines, Zootaxa 4868 (3), pp. 301-330 : 318 318
Decapoda   new species  Naruse, Tohru & Tamura, Hisao, 2006, A first record of anchialine crab of the genus Orcovita Ng and Tomascik, 1994 (Decapoda: Brachyura: Varunidae) from Japan, with description of the species., Limnology 7, No. 3, pp. 147-151 : 147 147
Hemigrapsus sanguineus     McDermott, John J., 2011, Parasites of shore crabs in the genus Hemigrapsus (Decapoda: Brachyura: Varunidae) and their status in crabs geographically displaced: a review, Journal of Natural History 45 (39 - 40), pp. 2419-2441 : 2429-2430 2429-2430
Thalassograpsus harpax     Naderloo, Reza & Türkay, Michael, 2012, Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: Faunistics, Biodiversity and Zoogeography 3374, Zootaxa 3374 (1), pp. 1-67 : 48 48
Metaplax indica     Naderloo, Reza & Türkay, Michael, 2012, Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: Faunistics, Biodiversity and Zoogeography 3374, Zootaxa 3374 (1), pp. 1-67 : 48 48
Eriocheir sinensis     Wicksten, Mary K., 2012, Decapod Crustacea of the Californian and Oregonian Zoogeographic Provinces 3371, Zootaxa 3371, pp. 1-307 : 264-265 264-265
Neohelice granulata     Mantelatto, Fernando L., Tamburus, Ana Francisca, Magalhães, Tatiana, Buranelli, Raquel C., Terossi, Mariana, Negri, Mariana, Castilho, Antonio L., Costa, Rogério C. & Zara, Fernando J., 2020, Checklist of decapod crustaceans from the coast of the São Paulo state (Brazil) supported by integrative molecular and morphological data: III. Infraorder Brachyura Latreille, 1802, Zootaxa 4872 (1), pp. 1-108 : 26 26
Cyclograpsus integer     Coelho, Petrônio Alves, Almeida, Alexandre Oliveira De & Bezerra, Luis Ernesto Arruda, 2008, Checklist of the marine and estuarine Brachyura (Crustacea: Decapoda) of northern and northeastern Brazil, Zootaxa 1956 (1), pp. 1-58 : 41 41
Ptychognathus key    Hsu, Jhih-Wei, Shih, Hsi-Te & Li, Jheng-Jhang, 2022, Description of a new species of brackish-water crab of the genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) from southern Taiwan, Nauplius (e 2022002) 30, pp. 1-11 : 8 8
Pseudohelice subquadrata     Hsu, Jhih-Wei, Shih, Hsi-Te & Innocenti, Gianna, 2022, A review of the mud crab genus Pseudohelice Sakai, Türkay & Yang, 2006 (Crustacea: Brachyura: Varunidae), with redescription of Cyclograpsus latreillii H. Milne Edwards, 1837, from the western Indian Ocean, Raffles Bulletin of Zoology 70, pp. 94-107 : 96-100 96-100
Ptychognathus altimanus     Lagare, Nickel Jean S., Mapi-Ot, Emmarie F., Molina, Ziljih S., Neri, Jemateo B., Nuñeza, Olga M. & Mendoza, Jose Christopher E., 2020, On a collection of freshwater and estuarine crabs (Crustacea: Brachyura) from Mindanao Island, the Philippines, Zootaxa 4868 (3), pp. 301-330 : 314-315 314-315
Hemigrapsus oregonensis     Wicksten, Mary K., 2012, Decapod Crustacea of the Californian and Oregonian Zoogeographic Provinces 3371, Zootaxa 3371, pp. 1-307 : 265-267 265-267
Metaplax tredecim     Shih, Hsi-Te, Hsu, Jhih-Wei, Wong, Kingsley J. H. & Ng, Ngan Kee, 2019, Review of the mudflat varunid crab genus Metaplax (Crustacea, Brachyura, Varunidae) from East Asia and northern Vietnam, ZooKeys 877, pp. 1-29 : 1 1
Ptychognathus glaber     Komai, Tomoyuki, Maenosono, Tadafumi, Saeki, Toshifumi & Naruse, Tohru, 2021, Redescription of the brackish water crab Ptychognathus glaber Stimpson, 1858 based on the topotypic material from the Ogasawara Islands and new record of P. lipkei N. K. Ng, 2010 from Japan (Decapoda: Brachyura: Varunidae), Zootaxa 5048 (1), pp. 58-76 : 60-66 60-66
Metaplax sheni     Shih, Hsi-Te, Hsu, Jhih-Wei, Wong, Kingsley J. H. & Ng, Ngan Kee, 2019, Review of the mudflat varunid crab genus Metaplax (Crustacea, Brachyura, Varunidae) from East Asia and northern Vietnam, ZooKeys 877, pp. 1-29 : 1 1
Eriocheir sinensis     Chatterjee, Tapas, 2021, A checklist of halacarid mites (Acari: Halacaridae) associated with decapod crustaceans (Arthropoda, Crustacea, Decapoda), Persian Journal of Acarology 10 (2), pp. 155-166 : 161 161
Ptychognathus lipkei     Komai, Tomoyuki, Maenosono, Tadafumi, Saeki, Toshifumi & Naruse, Tohru, 2021, Redescription of the brackish water crab Ptychognathus glaber Stimpson, 1858 based on the topotypic material from the Ogasawara Islands and new record of P. lipkei N. K. Ng, 2010 from Japan (Decapoda: Brachyura: Varunidae), Zootaxa 5048 (1), pp. 58-76 : 66-70 66-70
Acmaeopleura parvula     Naruse, Tohru, 2015, Description of a new genus and a new species of gaeticine crab (Crustacea: Brachyura: Varunidae) from the Ryukyu Islands, and a review of Acmaeopleura Stimpson, 1858, and Sestrostoma Davie & N. K. Ng, 2007, Zootaxa 3925 (2), pp. 211-228 : 218-221 218-221
Parahelice pilimana     Shih, Hsi-Te, Hsu, Jhih-Wei, Li, Jheng-Jhang, Ng, Ngan Kee & Lee, Jung-Hsiang, 2020, The identities of three species of Parahelice Sakai, Türkay & Yang, 2006 (Crustacea: Brachyura: Varunidae) from the western Pacific, based on morphological and molecular evidence, Zootaxa 4728 (2), pp. 249-265 : 255-257 255-257
Varunidae     Hernández-Orts, Jesús S., Kuchta, Roman, Semenas, Liliana, Crespo, Enrique A., González, Raúl A. & Aznar, Francisco J., 2019, An annotated list of the Acanthocephala from Argentina, Zootaxa 4663 (1), pp. 1-64 : 46 46
Decapoda   new species  Ng, Ngan Kee, Naruse, Tohru & Shih, Hsi-Te, 2018, Helice epicure, A New Species of Varunid Mud Crab (Brachyura, Decapoda, Grapsoidea) from the Ryukyus, Japan, Zoological Studies 57 : -1 -1
Parahelice daviei     Shih, Hsi-Te, Hsu, Jhih-Wei, Li, Jheng-Jhang, Ng, Ngan Kee & Lee, Jung-Hsiang, 2020, The identities of three species of Parahelice Sakai, Türkay & Yang, 2006 (Crustacea: Brachyura: Varunidae) from the western Pacific, based on morphological and molecular evidence, Zootaxa 4728 (2), pp. 249-265 : 252-254 252-254
Decapoda   new species  Komai, Tomoyuki, 2011, A new species of the varunid crab genus Gopkittisak (Crustacea: Decapoda: Brachyura: Grapsoidea) from the Ryukyu Islands, Species Diversity 16, No. 3, pp. 103-111 : 104-106,108-111 104-106,108-111
Varunidae     Castro, Peter, 2011, Catalog of the anomuran and brachyuran crabs (Crustacea: Decapoda: Anomura, Brachyura) of the Hawaiian Islands 2947, Zootaxa 2947 (1), pp. 1-154 : 144 144
Hemigrapsus nudus     McDermott, John J., 2011, Parasites of shore crabs in the genus Hemigrapsus (Decapoda: Brachyura: Varunidae) and their status in crabs geographically displaced: a review, Journal of Natural History 45 (39 - 40), pp. 2419-2441 : 2430-2431 2430-2431
Metaplax longipes     Shih, Hsi-Te, Hsu, Jhih-Wei, Wong, Kingsley J. H. & Ng, Ngan Kee, 2019, Review of the mudflat varunid crab genus Metaplax (Crustacea, Brachyura, Varunidae) from East Asia and northern Vietnam, ZooKeys 877, pp. 1-29 : 1 1
Ptychognathus sakaii   sp. nov.  Hsu, Jhih-Wei, Shih, Hsi-Te & Li, Jheng-Jhang, 2022, Description of a new species of brackish-water crab of the genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) from southern Taiwan, Nauplius (e 2022002) 30, pp. 1-11 : 3-7 3-7
Chasmagnathus granulata     Cremonte, Florencia & Drago, Fabiana B., 2007, Checklist of digenean parasites of birds from Argentina, Zootaxa 1403, pp. 1-36 : 29 29
Varunidae     Wicksten, Mary K., 2012, Decapod Crustacea of the Californian and Oregonian Zoogeographic Provinces 3371, Zootaxa 3371, pp. 1-307 : 262 262
Gaeticinae key    Naruse, Tohru, 2015, Description of a new genus and a new species of gaeticine crab (Crustacea: Brachyura: Varunidae) from the Ryukyu Islands, and a review of Acmaeopleura Stimpson, 1858, and Sestrostoma Davie & N. K. Ng, 2007, Zootaxa 3925 (2), pp. 211-228 : 226 226
Cyclograpsus granulatus     Castro, Peter, 2011, Catalog of the anomuran and brachyuran crabs (Crustacea: Decapoda: Anomura, Brachyura) of the Hawaiian Islands 2947, Zootaxa 2947 (1), pp. 1-154 : 121 121
Cyclograpsinae     Castro, Peter, 2011, Catalog of the anomuran and brachyuran crabs (Crustacea: Decapoda: Anomura, Brachyura) of the Hawaiian Islands 2947, Zootaxa 2947 (1), pp. 1-154 : 121 121
Eriocheir sinensis     GUINOT, DANIÈLE, TAVARES, MARCOS & CASTRO, PETER, 2013, Significance of the sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura), with new nomina for higher-ranked podotreme taxa, Zootaxa 3665 (1), pp. 1-414 : 110 110
Cyclograpsus integer     Parasram, Nadeshinie, Santana, William & Vallès, Henri, 2021, Updated checklist of semi-terrestrial and estuarine crabs (Crustacea: Decapoda: Brachyura) of Barbados, West Indies, Zootaxa 5052 (4), pp. 451-485 : 473-474 473-474
Gaeticinae     Naruse, Tohru, 2015, Description of a new genus and a new species of gaeticine crab (Crustacea: Brachyura: Varunidae) from the Ryukyu Islands, and a review of Acmaeopleura Stimpson, 1858, and Sestrostoma Davie & N. K. Ng, 2007, Zootaxa 3925 (2), pp. 211-228 : 212-215 212-215
Varunidae     González, José A., 2018, Checklists of Crustacea Decapoda from the Canary and Cape Verde Islands, with an assessment of Macaronesian and Cape Verde biogeographic marine ecoregions, Zootaxa 4413 (3), pp. 401-448 : 429-430 429-430
Ptychognathus insolitus   sp. nov.  Osawa, Masayuki & Ng, Ngankee, 2006, A new species of Ptychognathus Stimpson, 1858 (Crustacea: Decapoda: Brachyura: Varunidae) from the Ryukyu Islands, southwestern Japan, Zootaxa 1260, pp. 57-66 : 58-65 58-65
Orcovita key    Davie, Peter J. F. & Ng, Peter K. L., 2012, Two New Species Of Orcovita (Crustacea: Decapoda: Brachyura: Varunidae) From Anchialine Caves On Christmas Island, Eastern Indian Ocean, Raffles Bulletin of Zoology 60 (1), pp. 57-70 : 66 66
Chhapgarus   gen. nov.  Ng, Ngan Kee, Trivedi, Jigneshkumar & Bhat, Mithila, 2022, Redescription of Pseudograpsus intermedius Chhapgar, 1955 (Decapoda, Brachyura, Varunidae) from India, Zootaxa 5209 (1), pp. 127-138 : 130 130
Chhapgarus intermedius   comb. nov.  Ng, Ngan Kee, Trivedi, Jigneshkumar & Bhat, Mithila, 2022, Redescription of Pseudograpsus intermedius Chhapgar, 1955 (Decapoda, Brachyura, Varunidae) from India, Zootaxa 5209 (1), pp. 127-138 : 131 131
Metaplax indica     Naderloo, Reza, 2011, Grapsoid crabs (Decapoda: Brachyura: Thoracotremata) of the Persian Gulf and the Gulf of Oman, Zootaxa 3048, pp. 1-43 : 31-34 31-34
Gaeticinae     Ng, Peter K. L., 2012, The systematic status of two enigmatic ocypodoid crabs, " Paracleistostoma " dentatum Tesch, 1918, and " Paracleistostoma " fossulum Barnard, 1955 (Crustacea: Decapoda: Brachyura), Zootaxa 3206, pp. 58-68 : 62-67 62-67
Acmaeopleura     Naruse, Tohru, 2015, Description of a new genus and a new species of gaeticine crab (Crustacea: Brachyura: Varunidae) from the Ryukyu Islands, and a review of Acmaeopleura Stimpson, 1858, and Sestrostoma Davie & N. K. Ng, 2007, Zootaxa 3925 (2), pp. 211-228 : 215-218 215-218
Varuninae     Castro, Peter, 2011, Catalog of the anomuran and brachyuran crabs (Crustacea: Decapoda: Anomura, Brachyura) of the Hawaiian Islands 2947, Zootaxa 2947 (1), pp. 1-154 : 121 121
Austrohelice crassa     McDermott, John J., 2011, Parasites of shore crabs in the genus Hemigrapsus (Decapoda: Brachyura: Varunidae) and their status in crabs geographically displaced: a review, Journal of Natural History 45 (39 - 40), pp. 2419-2441 : 2426-2427 2426-2427
Varunidae     Martin, Joel W. & Haney, Todd A., 2005, Decapod crustaceans from hydrothermal vents and cold seeps: a review through 2005, Zoological Journal of the Linnean Society 145 (4), pp. 445-522 : 498 498
Hemigrapsus sexdentatus     McLay, C. L. & Schubart, C. D., 2004, Short Communication On the correct name for Hemigrapsus edwardsii (Hilgendorf, 1882) (Brachyura: Varunidae) from New Zealand, Journal of Natural History 38 (6), pp. 695-704 : 699-700 699-700
Ptychognathus riedelii     Lagare, Nickel Jean S., Mapi-Ot, Emmarie F., Molina, Ziljih S., Neri, Jemateo B., Nuñeza, Olga M. & Mendoza, Jose Christopher E., 2020, On a collection of freshwater and estuarine crabs (Crustacea: Brachyura) from Mindanao Island, the Philippines, Zootaxa 4868 (3), pp. 301-330 : 315 315
Parahelice pilosa     Shih, Hsi-Te, Hsu, Jhih-Wei, Li, Jheng-Jhang, Ng, Ngan Kee & Lee, Jung-Hsiang, 2020, The identities of three species of Parahelice Sakai, Türkay & Yang, 2006 (Crustacea: Brachyura: Varunidae) from the western Pacific, based on morphological and molecular evidence, Zootaxa 4728 (2), pp. 249-265 : 257-261 257-261
Decapoda   new species  Stasolla, Gianluca & Innocenti, Gianna, 2014, A new species of cavernicolous crab from Coron Island, Palawan, the Philippines (Decapoda: Brachyura: Varunidae), Raffles Bulletin of Zoology 62, pp. 591-599 : 596 596
Proexotelson   gen. nov.  Naruse, Tohru, 2015, Description of a new genus and a new species of gaeticine crab (Crustacea: Brachyura: Varunidae) from the Ryukyu Islands, and a review of Acmaeopleura Stimpson, 1858, and Sestrostoma Davie & N. K. Ng, 2007, Zootaxa 3925 (2), pp. 211-228 : 221-222 221-222
Varuna litterata     Lagare, Nickel Jean S., Mapi-Ot, Emmarie F., Molina, Ziljih S., Neri, Jemateo B., Nuñeza, Olga M. & Mendoza, Jose Christopher E., 2020, On a collection of freshwater and estuarine crabs (Crustacea: Brachyura) from Mindanao Island, the Philippines, Zootaxa 4868 (3), pp. 301-330 : 318-319 318-319
Varunidae key    Wicksten, Mary K., 2012, Decapod Crustacea of the Californian and Oregonian Zoogeographic Provinces 3371, Zootaxa 3371, pp. 1-307 : 264 264
Ptychognathus pusillus     Lagare, Nickel Jean S., Mapi-Ot, Emmarie F., Molina, Ziljih S., Neri, Jemateo B., Nuñeza, Olga M. & Mendoza, Jose Christopher E., 2020, On a collection of freshwater and estuarine crabs (Crustacea: Brachyura) from Mindanao Island, the Philippines, Zootaxa 4868 (3), pp. 301-330 : 315 315
Thalassograpsus harpax     Naderloo, Reza, 2011, Grapsoid crabs (Decapoda: Brachyura: Thoracotremata) of the Persian Gulf and the Gulf of Oman, Zootaxa 3048, pp. 1-43 : 34-36 34-36

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