Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae)
Author
Locke, Michelle M.
michellemlocke@gmail.com
Author
Skevington, Jeffrey H.
michellemlocke@gmail.com
text
Zootaxa
2013
2013-05-22
3660
1
1
80
http://dx.doi.org/10.11646/zootaxa.3660.1.1
journal article
10.11646/zootaxa.3660.1.1
1175-5326
5265855
95ADD39C-98BE-4879-B070-34A5D86BD67B
Dasysyrphus pauxillus
(Williston)
Figures: 14A–D, 18A & D, 21D
Synonyms and References:
Syrphus pauxillus
Williston 1887: 74
(original description)
Metasyrphus pauxillus
:
Fluke 1933: 115
(key, redescription and transfer to
Metasyrphus
)
Dasysyrphus pauxillus
:
Stone
et al.
1965: 564
(catalogue and transfer to
Dasysyrphus
)
Vockeroth 1986: 203
(discussion)
Vockeroth 1992: 68
(key and redescription)
Speight 2011: 70
(discussion)
Diagnosis: Abdominal maculae of tergites 3&4 slightly arcuate, uniform in thickness and do not cross abdominal margin (
Figs 14A & B
); very similar in appearance to
D. pacificus
(
Fig. 13A
),
D. nigricornis
(
Figs 11A & B
) and
D. laticaudus
(
Fig. 8A
). Wing typically densely microtrichose, sometimes with bare areas in cells r
1
, br, bm and/or c (
Fig. 14C
). Male gonostylus with projection on posterodorsal surface straight (
Fig. 14D
). Due to an extremely similar habitus, it is difficult to distinguish
D. pacificus
from
D. pauxillus
.
Male genitalia should be used to conclusively identify specimens, especially in the overlap zone of these species’ ranges in southern
British Columbia
.
Redescription:
Body Length:
5.3–9.7mm
; Wing Length:
5.2–9.5mm
Head: Frons dark with light pollenose fascia (in males it runs along ventral edge of where eyes meet, in females it is ⅓–½ the length between the antennae and ocelli with a separation medially and edges triangular to rounded (
Fig. 18A
)) and light to dark pile; face light with dark vitta ¼–⅓ the width of the face, coming close to antennal socket if not reaching it, face with dark pile lateral to the eye, some females with varying amounts of light pile interspersed with dark pile; gena with light to dark pile; occiput dark, covered in light pollen, pile light; scape and pedicel light to dark, flagellomere dark with lighter base.
Thorax: Scutum dark, may appear shiny or metallic, pile light to dark; scutellum light to dark with darker lateral edges, pile on female mostly light with some dark, male mostly dark pile some with light interspersed throughout; wing cell r
1
with the base bare or very small areas at base bare, br, bm and c may all be entirely microtrichose or they may have bare areas along spurious vein, along the middle of the cell
form the
base to ¾ or small areas at base, respectively (
Fig. 14C
); haltere light; pro- and mesofemora ⅓–⅔ dark basally, ⅓–⅔ light apically, metafemur ⅔–¾ dark basally, ¼–⅓ light apically, tibiae light, meso- and metatibiae with dark band ¼ of the way from the apex, tarsi light with dark anterior side.
Abdomen: Maculae on tergite 2 large and oval, lateral edge tends to be more flattened than medial edge, female usually with very small, pointed, anterolateral projection, but it never reaches the edge of the tergite, maculae on tergites 3&4 are oblique, fairly straight and uniform in width, sometimes with medial edge slightly swollen, maculae do not reach abdominal margin (
Figs 14A & B
); sternites light with dark fasciae of varying thickness (
Fig. 18D
).
Male Genitalia: Surstylus more or less triangular in shape in lateral view, flattened posteriorly, long pile on dorsal, laterodorsal and posterodorsal sides, row of spines on posteromedial edge; cercus oval with long pile over entire surface; gonostylus boot-shaped but more narrow apically than basally, with projection on posterodorsal surface, projecting dorsally, pile on dorsal surface; basiphallus elongate, basally bent at a slightly more than 90 degree angle towards dorsum, apical end curved ventrally, but does not greatly extend ventrally and is partly membranous dorsally, with spines on posterodorsal side; distiphallus elongate, apical end flared out into horn, which is mostly sclerotized but membranous at apex, with no setulae dorsally, basally bent towards dorsum and usually greatly enlarged at bend with a triangular projection, projecting ventrally (
Fig. 14D
). Extremely similar to
D. laticaudus
(
Fig. 8C
).
FIGURE 14.
Dasysyrphus pauxillus
: A: male (JSS 24616) dorsal habitus; B: male (JSS 24616) lateral habitus; C: wing base (CNCD 13830); D: lateral view of male (CNCD 13746) genitalia. Dorsal and lateral habitus scale bar = 1mm, genitalia and wing scale bar = 500µm.
Intraspecific variation: Wings are typically densely microtrichose with cell r
1
having bare areas at base ranging from entire base bare to very small areas at base being bare, cells br, bm and c may all be entirely microtrichose or they may have bare areas; female usually have a very small, anterolateral projection on maculae of tergite 2, males do not have this projection.
Etymology: From the Latin
pauxillus
, meaning small.
Distribution: Rocky Mountains; Western
Canada
(AB, BC, MB, YT) and western
USA
(AK, CO, ID, NM, UT, WY) (
Fig. 21D
).
Ecology: Collected from March–July, but more commonly collected in June. Collected at elevations of
14– 3505m
. Habitats collected from are listed as “moraine lake”, “grassy lakeshore”, “over stream in fir forest”, “on tundra”, “spruce forest” and “
Pinus-Salix
forest”. It has been collected from the following plant species: common cow parsnip (
Heracleum maximum
W. Bartram
,
Apiaceae
), blackberry (
Rubus
Linnaeus
,
Rosaceae
), willow (
Salix
Linnaeus
,
Salicaceae
),
Canada
gooseberry (
Ribes oxyacanthoides
Linnaeus
,
Grossulariaceae
) and spruce (
Picea
A. Dietrich
,
Pinaceae
).
Discussion: Many authors have used the name
pauxillus
in the Palaearctic. During the course of this revision few specimens from that region were examined and no DNA evidence could be studied. Therefore, we cannot comment on whether the use of this name is appropriate in the Palaearctic. However, based on the geographic breakdown of
pauxillus
and its allies (
laticaudus
,
pacificus
and
nigricornis
) in the Nearctic it seems likely that only
nigricornis
is Holarctic. European and Asian specimens should be studied in greater depth, using both morphology and DNA, to confirm this suspicion.
Due to the widespread use of the name
pauxillus
much confusion exists over the concept of this species. Many Holarctic species have been named which have an extremely similar habitus to
pauxillus
. The main reason for such confusion is that the
holotype
specimen is lost (
Fluke 1933
). In
Williston’s (1887)
original designation he describes
one female
from
New Mexico
. He does not indicate where this specimen is deposited. The only other information that is provided is that the specimen was collected by G.F. Gaumer. In
Fluke’s (1933)
revision of
Metasyrphus
he indicates that he was unable to locate the type for examination. No record exists in the literature of any author examining Williston’s specimen; either Williston’s species description was used or specimens determined by other researchers were used for comparison.
Dasysyrphus pauxillus
was described in ‘Synopsis of the North American Syrphidae’, Williston’s PhD dissertation (
Beckemeyer 2009
). The specimens from his dissertation were donated to the USNM (
Flint
et al.
1976
, F.C. Thompson, pers. comm.). According to Thompson (pers. comm.) the type specimen is not in the USNM collection today and it is not known what happened to it. Williston worked in
Kansas
and Chicago until his death, after which his collection was donated to the American Museum of Natural History (AMNH). Byers
et al
. (
Byers
et al.
1962
) do not list
pauxillus
as a type in the
Kansas
collection and the specimen is not listed in the type database of the AMNH (The
American Museum of Natural History 2005
). A
neotype
is designated here to bring stability to the name
pauxillus
. This will create a solid concept for this species, allowing for better reconciliation of closely related species. The specimen used in the original designation provided a locality of ‘New Mexico’. The
neotype
specimen was chosen from a series of males from
Utah
, as the
two specimens
from
New Mexico
were a female and a male in poor condition. This is a locality that is not overlapped by the other species with a similar habitus to
pauxillus
(
pacificus
,
laticaudus
and
nigricornis
). The
neotype
varies from Williston’s description in a few characters that are variable in many
Dasysyrphus
species.
Also, Williston’s description was of a female and the
neotype
is male, which may account for some of the variation. The dorsum of the scutum in the
neotype
is covered by mainly dark pile, Williston’s description states the pile is light yellowish. This character is variable within most
Dasysyrphus
species
and often males have more dark pile and females have more light pile on the scutum. The face is also not projecting to the extent described by Williston. The face seems to project slightly more in females than males, but in most specimens the distance between the antennae and the tip of the tubercle is more or less equal to the distance from the antennae to the ocelli. The specimen matches the description in all other characters. The
Neotype
is a male from Tony Grove Lake, Wasatch-Cache National Forest, UT,
USA
, collected on
1.vii.2010
by M.M. Locke. It is deposited in the CNC and databased with the number Jeff Skevington Specimen 24616.
Type Material:
“
USA
:
UT
:Wasatch-CacheNF:/ Tony Grove Lake;/
41.893136°N
,
111.641841°W
;/
1.vii.2010
;
M.M.Locke
”, “
J. Skevington
/ Specimen #/ 24616”, “
NEOTYPE
/
Syrphus
/ pauxillus/ Williston”, “designated by/ Locke &/ Skevington/ 2013”;
♂
, deposited in the
Canadian National Collection of Insects
, Arachnids and Nematodes, Ottawa,
Ontario
,
Canada
.
Material Examined:
Canada
:
Alberta
:
WATERTON LAKES NATIONAL PARK
:
Waterton
, [
49.051122
,
-113.911807
],
6.vi.1962
,
2♂
,
CNCD 13764
,
13765
,
4♀
,
CNCD 13836–13839
, W.
R
.
M.
Mason
(
CNC
),
2♀
,
CNCD 13834
,
13835
,
K.C.
Herrmann
(
CNC
). KANANASKIS:
25 km
WNW Bragg Creek Powderface
trail
Rd.
, [
51.02166667
,
-114.9016667
],
8.vi.2001
,
♂
,
RBCM
ENT 6006051
,
C.J.
Borkent
(
RBCM
);
Forest Experimental Station Seebe
, [
51.100879
,
- 115.087692
],
2.vi.1968
,
♀
,
CNCD 13840
,
13.vi.1968
,
♀
,
CNCD 13841
,
15.vi.1968
,
♀
,
CNCD 13842
,
H.J.
Teskey
(
CNC
).
30 miles
West of Calgary
, [
51.04575
,
-113.631333
],
3.vi.1973
,
CNCD 60100
,
G.G.E.
Scudder
(
CNC
).
BANFF NATIONAL PARK
:
Banff
, [
51.180275
,
-115.568433
],
27.v.1925
,
♂
,
CNCD 13762
,
E.
Hearle
(
CNC
);
2.vi.1922
,
♂
,
CNCD 13763
,
3♀
,
CNCD 13845–13847
,
9.vi.1922
,
♀
,
CNCD 13849
,
3.vii.1922
,
♀
,
CNCD 13848
,
C.B.D.
Garrett
(
CNC
);
2.vi.1914
,
♀
, JSS 20605, N.B.
Sanson
(
EMEC
);
30.v.1948
,
♀
,
UASM# 149024
,
2.vi.1948
,
♀
,
UASM# 149025
,
E.H.
Strickland
(
UASM
);
Mount Eisenhower
, [
51.298694
,
-115.925724
],
3.vi.1968
,
2♂
,
CNCD 13770
,
13771
,
5.vi.1968
,
♂
,
CNCD 13769
,
7.vi.1968
,
♂
,
CNCD 13768
,
Mosquin
&
Seaborn
(
CNC
);
Moraine Lake
, [
51.324073
,
-116.184418
],
2256m
,
20.vii.1955
,
♀
,
CNCD 13850
, J.
R
.
McGillis
(
CNC
);
8 km
West of Lake Louise
, [
51.413684
,
-116.306046
],
31.v.1992
,
♀
, JSS 20801, A.
Borkent
(
CNC
);
Lake Agnes
, [
51.415101
,
-116.247314
],
2073m
,
10.vii.1925
,
♂
, JSS 19508,
O. Bryant
(
CNC
);
Snow Creek Pass
, [
51.60524
,
-115.808617
],
2256m
,
20.vii.1962
,
♀
,
CNCD 13844
,
K.C.
Herrmann
(
CNC
);
Peyto Lookout
, [
51.717631
,
-116.508919
],
2316m
,
23.vii.1962
,
♀
,
CNCD 13843
, W.
R
.
M. Mason
(
CNC
).
Edmonton
,
12.v.1924
,
♂
, JSS 23724,
Owen Bryant
(
CAS
);
Laggen
,
Lake Agnes
,
10.vii.1925
,
♀
, JSS 23714,
O. Bryant
(
CAS
).
Edmonton
;
W. Hawrelak Park
,
640m
,
3.v.2005
,
3♂
, JSS 25540–25542,
W. van Steenis
(
WVSC
).
Pincher Creek
;
Beauvais Lake
,
19.v.1989
23.v.1989
,
♀
, JSS 25550,
C. v. Nidek
(
WVSC
)
.
British Columbia
:
Victoria
,
Mt. Work
, [48.53, -123.47],
26.v.1982
,
♂
,
RBCM
ENT 99188111
,
R
.
A.
Cannings
(
RBCM
).
Galiano
I.
North End
, [48.92, -123.42],
29.iii.1987
,
♂
, JSS 20184, G.G.E.
Scudder
(
UBCZ
).
WATERTON LAKES NATIONAL PARK
:
Cameron Lake
, [
49.011457
,
-114.049238
],
17.vi.1956
,
9♂
,
CNCD 13754–13761
,
13767
,
9♀
,
CNCD 13822–13830
,
18.vi.1956
,
♀
,
CNCD 13831
,
19.vi.1956
,
2♀
,
CNCD 13832
,
13833
,
E.E.
Sterns
(
CNC
). CENTRAL
KOOTENAY
L.D.:
Robson
, [
49.333015
,
-117.692871
],
21.iv.1947
,
♂
,
CNCD 13746
,
8.v.1950
,
♂
,
CNCD 13750
, H.
R
.
Foxlee
(
CNC
);
25.iv.1958
,
♂
, JSS 20123, H.
R
.
Foxlee
(
UBCZ
);
Boswell
, [
49.414407
,
-116.746741
],
9.v.1958
,
♂
,
CNCD 13742
,
H. & A.
Howden
(
CNC
). SQUAMISH- LILLOOET
R
.
D.:
Meager Creek Hot Springs
, [
50.561617
,
-123.486344
],
14.v.1987
,
♂
, JSS 18370,
C.S. Guppy
(
CNC
).
KOOTENAY NATIONAL PARK
:
Daer-Pitts Aspen Control
1, [50.95, -116.0333333],
3.vi.2000
17.vi.2000
,
CNCD 7969
,
G.
Gareau
(
CNC
);
Daer-Pitts Aspen Control
2,
17.vi.2000
2.vii.2000
,
♀
,
CNCD 30906
,
G.
Gareau
(
CNC
).
32 miles
South West of Terrace
, [
54.194076
,
-129.140296
],
30.5m
,
6.vi.1960
,
♀
,
CNCD 13819
,
G.E.
Shewell
(
CNC
);
6.vi.1960
,
♀
,
CNCD 13820
,
11.vi.1960
,
♂
,
CNCD 13753
,
B.
Heming
(
CNC
).
Shames
, 18
Miles South West of Terrace
, [
54.409648
,
-128.935301
],
23.vi.1960
,
♀
,
CNCD 13818
,
C. H.
Mann
(
CNC
).
Kleanza Creek
,
Near Terrace
, [
54.597897
,
-128.386241
],
76.2m
,
29.vi.1960
,
♀
,
CNCD 13817
,
B.
Heming
(
CNC
);
30.vi.1960
,
♀
,
CNCD 13821
,
J.G.
Chillcott
(
CNC
).
Atlin
, [
59.5775
,
-133.69236
],
670.56m
,
2.vi.1955
,
♂
,
CNCD 13751
,.
vii.1955
,
♀
,
CNCD 13809
,
H.J.
Huckel
(
CNC
);
20.vi.1955
,
♂
,
CNCD 14358
,
♀
,
CNCD 13808
,
B.A.
Gibbard
(
CNC
)
.
Manitoba
:
26km
SE Churchill
;
Twin Lakes
, [58.63, -93.819],
14m
,
19.vii.2009
,
♀
, 09
PROBE 197
,
22.vii.2009
,
♂
, 09
PROBE 255
,
Arctic Ecology
2009 (
CNC
)
.
Yukon Territory
:
Snafu Creekl
, [60.13333333, -138.8],
22.vi.1982
,
CNCD 60079
,
S.G.
Cannings
,
R
.
D.
Wilkie
, L.
Vasington
(
CNC
).
Whitehorse
, [
60.71965
,
-135.052213
],
6.vii.1988
,
♀
,
CNCD 5812
,
F.
Brodo
(
CNC
).
Simpson L.
81 km
n
Watson
L., [60.75, -129.25],
15.vi.1984
1.ix.1984
,
♀
, JSS 20770,
S. & J. Peck
(
CNC
).
Nines Creek
,
3 km
West of Alaska Highway
, [61.18333333, -139.7],
4.vi.1981
,
CNCD 60072
,
C.
Guppy
(
CNC
).
La Force Lake
, [
62.683333
,
-132.333333
],
914m
,
26.vi.1960
,
♀
,
CNCD 13585
,
10.vii.1960
,
♀
,
CNCD 13728
,
E.W.
Rockburne
(
CNC
)
.
United States of America
:
Alaska
:
MATANUSKA- SUSITNA
CO
.:
Matanuska
, [
61.541945
,
-149.229674
],
12.vi.1944
,
♀
, JSS 19502,
J. Chamberlin
(
CNC
)
.
Colorado
:
Rocky Gulch
, [
37.817035
,
-107.579606
],
3444m
, 6.xxii.2002,
♂
, JSS 20810, C.
Slater
(
CSUC
).
Monarch Pass
,
2mi
W Garfield
, [
38.501433
,
-106.327747
],
3048m
,
19.vi.1985
,
♂
,
USNM
ENT 258074
,
David
R
.
Smith
(
USNM
).
Rock Cr.
,
May Mus.
, [
38.705655
,
-104.837421
],
8.v.1996
,
♀
, JSS 20808,
D. Leatherman
(
CSUC
).
Tenmile Range
, unamed
Cr.
avalanche chute W side, [
39.425
,
-106.111
],
3200m
,
15.vii.1998
17.vii.1998
,
♂
, JSS 20694,
S. Fitzgerald
&
A. Foley
(
CSUC
).
CLEAR CREEK
CO
.:
Mt. Evans
,
Echo Lake Campground
, [
39.65622
,
105.595042
],
329m
,
29.vi.2010
,
CNCD 60035
,
M.M.
Locke
(
CNC
);
Mt. Evans
,
Echo Lake
, [
39.658085
,
- 105.6035
],
3231m
,
13.vii.1961
,
♀
,
CNCD 26416
,
C.H.
Mann
(
CNC
);
29.vi.2010
,
♀
, JSS 24618, M.M.
Locke
(
CNC
);
29.vi.2010
,
♀
,
CNCD 35312
,
A.
Young
(
CNC
);
Mt. Evans
,
Doolittle Ranch
, [
39.675739
,
-105.601348
],
2987m
,
13.vii.1961
,
♀
,
CNCD 26417
, W.
R
.
M.
Mason
(
CNC
). BOULDER
CO
.:
Nederland
,
Science Lodge
, [
40.031645
,
-105.533886
],
2896m
,
4.vii.1961
,
♀
,
CNCD 26418
,
B.H.
Poole
(
CNC
);
Niwot Ridge
,
Near Ward
, [
40.069933
,
-105.607397
],
3444m
,
4.vi.1961
,
♂
,
CNCD 26440
,
28.vi.1961
,
♂
,
CNCD 26439
,
C.H. Mann
(
CNC
).
North Park
, [40.73, -106.18],
1.vii.1932
,
♀
, JSS 20746, (
CSUC
).
Rawah Wilderness
,
Blue
Lake
Trail
near
Fall Creek
, [
40.76944444
,
-105.9894444
],
2926m
,
20.vi.1998
,
♂
, JSS 20693,
S. Fitzgerald
&
A. Foley
(
CSUC
). Cherokee Park, [
40.90972
,
-105.47917
],
30.vii.1913
,
♀
, JSS 20713,
Chas L. Fluke
(
CSUC
). MINERAL
CO
.:
Continental Divide
,
3292m
,
20.vi.1919
,
♂
, JSS 19512, (
CNC
).
Camera Point
,
Pike’s Peak
,
16.vi.1956
,
♀
, JSS 21157,
R
. &
K.
Dreisbach
(
MSUC
)
.
Idaho
:
Island Park
, [
44.424364
,
-111.371064
], JSS 25099,
Bernard Knapp
(
EMUS
).
NEZ
PERCE
CO
.:
Lake Waha
, [
46.205942
,
-116.837015
],
9.vi.1918
,
♀
,
CNCD 26402
,
A.L.
Melander
(
CNC
);
18.vi.1930
,
♂
, JSS 19514, W.M.
Davidson
(
CNC
);
Waha
, [
46.211439
,
-116.844439
],
18.vi.1909
,
♀
, JSS 19501, (
CNC
);
30.v.1924
,
♀
, JSS 23778,
A.L. Melander
(
USNM
).
LATAH
CO
.:
Moscow Mt.
, [
46.800972
,
- 116.866093
],
8.vi.1921
,
♀
, JSS 23776,
26.vi.1922
,
♀
, JSS 23777,
A.L. Melander
(
USNM
);
Mountains Near Moscow
,
Latah County
, [
46.807258
,
-116.868969
],
1.vi.1907
,
♀
,
CNCD 26403
, (
CNC
);
17.vi.1918
,
♂
, JSS 19509,
26.vi.1920
,
♂
, JSS 19506,
9.vii.1920
,
♀
, JSS 19505,
A.L. Melander
(
CNC
);
vii.1910
,
♀
, JSS 19510,
25.vi.1920
,
♂
, JSS 19516,
R
.
C.
Shannon
(
CNC
).
Savary
,
29.iv.1916
,
♀
, JSS 21329, (
MCZ
)
.
New Mexico
:
Las Cruces
,
Dona Ana
, [32.3, -106.7666667],
♀
,
CNCD 26423
,
F.M.
Hull
(
CNC
). LINCOLN
CO
.,
Sierra Blanca
, [
33.478065
,
- 105.552907
],
2957m
,
10–26.vi.1979
,
♂
, JSS 18307,
S. & J. Peck
(
CNC
). SAN
MIGUEL
CO
.,
Santa Fe National Forest
, [
35.705996
,
-105.545731
],
27.v.1981
,
♀
, JSS 18346,
B.J. Sinclair
(
CNC
). SANDOVAL
CO
.:
Jemez Springs
, [
35.771336
,
-106.690555
], iv,
♀
,
CNCD 26422
, (
CNC
)
.
Utah
:
WASATCH
CO
.:
Daniels Pass
,
2 miles
South
, [
40.280034
,
-111.223675
],
2591m
,
9.vii.1961
,
♀
,
CNCD 26421
,
J.G.
Chillcott
(
CNC
). DUSCHENE
CO
.:
2 miles
West of Mirror Lake
, [
40.70363
,
-110.928511
],
23.vi.1968
,
♂
,
CNCD 26438
, B.
V
.
Peterson
(
CNC
).
CACHE
CO
.:
Card Canyon
, [
41.76236
,
-111.663482
],
13.vi.1937
,
♀
,
CNCD 26420
,
A.E.
Nye
(
CNC
);
Cowley Canyon
, [
41.772669
,
-111.60407
],
13–27.v.1989
,
♂
, JSS 24612,
W.J. Hanson
(
EMUS
);
Tony Grove Lake
, [
41.893136
,
- 111.641841
],
1.vii.2010
,
CNCD 60037–60040
,
♂
, JSS 24615,
M.M. Locke
(
CNC
);
West Hodges Canyon
, [
41.910305
,
-111.42277
],
13–20.vi.1980
, JSS 25096, (
EMUS
);
Left Hand Fork
,
Blacksmith Fork Canyon
,
24.v.2004
,
♀
, JSS 24617,
Brammer
(
EMUS
).
RICH
CO
.:
Logan Canyon Summit
, [
41.862735
,
-111.494404
],
26.vi.–2.vii.1982
, JSS 25098, (
EMUS
)
.
Wyoming
:
YELLOWSTONE NATIONAL PARK
:
Lewis Lake
, [
44.306126
,
- 110.537673
],
17.vii.1923
,
♀
, JSS 19500,
A.L. Melander
(
CNC
);
17.vii.1923
,
♀
, JSS 23774,
A.L. Melander
(
USNM
)
.