Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae) Author Locke, Michelle M. michellemlocke@gmail.com Author Skevington, Jeffrey H. michellemlocke@gmail.com text Zootaxa 2013 2013-05-22 3660 1 1 80 http://dx.doi.org/10.11646/zootaxa.3660.1.1 journal article 10.11646/zootaxa.3660.1.1 1175-5326 5265855 95ADD39C-98BE-4879-B070-34A5D86BD67B Dasysyrphus creper (Snow) Figures: 6A–D, 20A Synonyms and References: ? Syrphus lotus Williston 1887: 75 var. (short description of a variety of lotus ) Syrphus pauxillus Snow 1892: 37 (nec. Williston; short description) Syrphus creper Snow 1895: 234 (original description) Syrphus lotus Curran 1925: 92 Pl. VI Fig. 51 (nec. Williston; redescription) Syrphus lotus var. creper Curran 1925: 180 (discussion) Metasyrphus creper (Snow) : Fluke 1933: 107 (key, redescription and transfer to Metasyrphus ) Dasysyrphus creper (Snow) : Stone et al. 1965: 563 (catalogue and transfer to Dasysyrphus ) Vockeroth 1986: 202 ( lectotype designation) Vockeroth 1992: 65 (key and redescription) Diagnosis: Abdominal maculae do not meet abdominal margin; maculae on tergites 3&4 arcuate with medial edge flattened and usually broadly contacting anterior edge; markings unlike any other species ( Figs 6A & C ). Redescription: Body Length: 6.7–10.5mm ; Wing Length: 6.3–9.2mm Head: Frons dark with light pollenose fascia (in male it runs along ventral edge of where eyes meet, in females it is ½ the length between the antennae and ocelli) and dark pile; face light with dark vitta ¼–⅓ the width of the face, usually not reaching antennal socket, face with light pile, dark pile lateral to the eye; gena light ( Fig. 6C ) with light pile and usually some dark pile below eye; occiput dark, covered in light pollen, pile light; scape and pedicel light to dark, flagellomere dark. Thorax: Scutum dark, may appear shiny or metallic, pile light; scutellum light to dark with darker lateral edges, light pile along anterior edge and dark pile elsewhere; wing cells br and bm entirely bare, cell c bare from base to ¾, r 1 bare at base and cup bare at base and along vein CuP ( Fig. 6D ); haltere light; pro- and mesofemora ⅓–½ dark basially, ½–⅔ light apically, metafemur ⅔–¾ dark basially, ¼–⅓ light apically, tibiae light, metatibia with dark band ¼ of the way from the apex, tarsii light, sometime dark anteriorly. Abdomen: Maculae on tergite 2 large and oval, maculae on tergites 3&4 are arcuate, with medial edge widened, flattened and usually broadly contacting the anterior edge of tergite, sometimes meeting medially, maculae do not reach abdominal margin ( Figs 6A & C ); sternites light with dark fasciae. Male Genitalia: Surstylus more or less triangular in shape in lateral view, flattened posteriorly, long pile on dorsal, laterodorsal and posterodorsal sides, row of spines on posteromedial edge; cercus oval with long pile over entire surface; gonostylus boot-shaped with projection on posterodorsal surface that projects dorsally, pile on dorsal surface; basiphallus elongate, basally bent at 90 degree angle towards dorsum, apical end curved ventrally and is partly membranous with pile on dorsal side; distiphallus elongate, apical end flared out into horn, which is mostly sclerotized but membranous at apex with minute setulae dorsally, basally bent towards dorsum and slightly enlarged at bend ( Fig. 6B ). FIGURE 6. Dasysyrphus creper : A: male (CNCD 26355) dorsal habitus; B: lateral view of male (JSS 20682) genitalia; C: male (CNCD 26355) lateral habitus, genitalia removed; D: wing base (CNCD 26363). Dorsal and lateral habitus scale bar = 1mm, genitalia and wing scale bar = 500µm. Intraspecific variation: Scutellum colour varies from light to dark; maculae on tergites 3&4 sometimes meet medially and sometimes are separated, medial edge usually broadly contacts anterior edge of tergite but if not then it comes close to anterior edge and edge of maculae are rounded; dark fasciae on sternites vary in width. Etymology: The word creper is Latin, meaning dark or obscure. Distribution: Southwestern Canada (AB, BC) and western USA (AZ, CA, CO, ID, MT, NM, OR, SD, UT, WA, WY). ( Fig. 20A ) Ecology: Collected on ponderosa pine ( Pinus ponderosa Douglas ex Lawson & C. Lawson , Pinaceae ), sierra willow ( Salix orestera C.K. Schneid. , Salicaceae ), willow ( Salix Linnaeus , Salicaceae ), chokecherry ( Prunus virginiana , Rosaceae ), Ross’ avens ( Geum rossii (R. Brown) Seringe , Rosaceae ), avens ( Geum Linnaeus , Rosaceae ), tall ragwort ( Senecio serra Hooker , Asteraceae ), threenerve goldenrod ( Solidago velutina DC. , Asteraceae ), rabbitbush ( Chrysothamnus Nuttall , Asteraceae ). Habitats listed on labels include “along timberline”, “creek margin” and “on tundra”. The specimen collected on the “tundra” was collected in Colorado at 3505m , so this is in reference to an alpine tundra habitat. Collected April to October, although much more frequently collected in July and August. Collected from elevations of 305m – 3688m , although most frequently collected from elevations above 2000m . Discussion: Presumably Williston (1887) was referring to a specimen of D. creper when he described the female from New Mexico as a second specimen in his description of lotus . He noted there were differences but that it was “perhaps allied” with the female from Arizona that he described as lotus . He describes the abdominal maculae of this second specimen as “more oblique, and strongly concave in front” and “more convex behind” and the gena as having a “black stripe”, which are characters more befitting of Snow’s description of creper , which came some seven years later. Three years before Snow formally described creper in 1895 he described specimens as pauxillus ( Snow 1892 ) . In his description of creper he synonymizes his 1892 pauxillus description with creper . Snow is also the one who suggests Williston’s (1887) variety of lotus is probably creper . In Curran’s monograph ( Curran 1925 ) he demotes S. creper from the rank of species to a variety of S. lotus . He claimed that there were no differences between the two except for the amount of interruption in the second and third abdominal bands. He believed it was just an extreme in coloration and did not warrant the rank of species. However, in the same paper he provides a redescription of lotus and based on the characters discussed and illustrated (Plate VI: 51) it is clear that his concept of lotus is not the same as Williston’s and he is actually describing creper . In 1933, Fluke re-elevated creper to the rank of species. Fluke states that at that time lotus was only known from the type and that published records of lotus probably refer to creper . Fluke also notes that Curran was probably looking at specimens of creper , not lotus . Fluke (1933) clearly distinguishes between these two species based on the abdominal markings, with lotus having more straight bands and those of creper being more arcuate. Fluke also places both species into Metasyrphus and into the amalopis group with other species currently recognized as Dasysyrphus . Type Material: “Magdalena,/ New Mexico ”, “Type”, blank red label, red label “ Syrphus / creper Snow / LECTOTYPE / Des. Vockeroth/ 1984”, “Jeff Skevington/ Specimen #/ 23782”; ♂ ; deposited in the Snow Entomological Museum, University of Kansas . (examined) Material Examined: Paralectotypes : United States of America : Colorado : Estes Park , [ 40.376877 , -105.52137 ], viii.1892 , ♂ , JSS 20203, F.H. Snow ( FMNH ), ♂ , OSUC 319691 , F.H. Snow ( OSU ) . Other Material Examined: Canada : Alberta : Elkwater , Hwy 41, [49.65, -110.2666667], 7.vii.1982 , ♀ , JSS 18293, B. V . Peterson ( CNC ). Medicine Hat , Division No. 1, [ 50.040483 , -110.673523 ] , 15.v.1930 , ♂ , JSS 19477, F.S. Carr ( CNC ). Kananaskis , Forest Experimental Station Seebe, [ 51.100879 , -115.087692 ] , 15.vi.1968 , ♂ , CNCD 13634 , H.J. Teskey, 20.vi.1950 , ♀ , CNCD 13643 ( CNC ). BANFF NATIONAL PARK : Sulphur Mountain , [ 51.149438 , -115.583484 ], 2195m , 28.vii.1967 , 2♂ , CNCD 13635 , 13636 , J. R . Vockeroth ( CNC ); Banff , [ 51.180275 , -115.568433 ] , 27.vi.1925 , ♀ , JSS 19442, O. Bryant ( CNC ); Snow Creek Pass , [ 51.60524 , -115.808617 ], 2256m , 28.vii.1962 , ♀ , CNCD 13642 , 13.viii.1962 , ♀ , CNCD 13644 , K.C. Herrmann ( CNC ); Peyto Lookout , [ 51.717631 , -116.508919 ], 2438m , 23.vii.1962 , ♂ , CNCD 13630 , ♀ , CNCD 13641 , K.C. Herrmann ( CNC ) . British Columbia : CATHEDRAL PROVINCIAL PARK : Glacier Lake , [ 49.055785 , -120.207299 ] , 6.vii.1986 , ♂ , CNCD 13631 , R . A. Cannings ( CNC ) ; 7.vii.1986 , ♂ , RBCM ENT 9881939 , ♀ , RBCM ENT 9881942 , R . A. Cannings ( RBCM ); Quiniscoe Mtn. summit, [49.08, -120.18] , 26.vii.1988 , ♂ , JSS 20170, S.G. Cannings ( UBCZ ). OKANAGAN- SIMILKAMEEN R . D.: Manning Provincial Park , Valleyview , [ 49.07995 , -120.766842 ], 1830m , 14.vii.1986 , ♂ , JSS 18368, S.G. Cannings ( CNC ); Blackwall Mountain , [ 49.098666 , -120.766925 ], 1828m , 7.viii.1953 , ♂ , CNCD 13632 , D.F. Hardwick ( CNC ); Osoyoos , Mount Kobau , [ 49.111594 , -119.664702 ] , 9.vii.1968 , ♂ , CNCD 13627 , ♀ , CNCD 13640 , D. Allen , 13.viii.1967 , ♂ , CNCD 13628 , J. R . Vockeroth ( CNC ); Oliver , [ 49.182338 , -119.550442 ], 305m , 23.viii.1953 , ♀ , CNCD 13637 , J.E.H. Martin , 14.ix.1923 , ♀ , CNCD 13638 , C.B. Garrett ( CNC ); Hedley , [ 49.357784 , -120.075962 ] , 7.viii.1953 , ♂ , CNCD 13633 , D.F. Hardwick ( CNC ); Apex Mountain , [ 49.482486 , -119.934458 ], 1900m , 15.vii.1988 , ♂ , RBCM ENT 9885672 , R . A. Cannings ( RBCM ). Robson , [ 49.342029 , -117.697831 ] , 22.vi.1959 , ♀ , CNCD 13639 , H. R . Foxlee ( CNC ). Cranbrook , Elizabeth Lake , [ 49.497195 , -115.791152 ] , 6.vii.1980 , CNCD 60067 , R . A. Cannings ( CNC ). MOUNT REVELSTOKE NATIONAL PARK : Mount Harry , First Summit , [ 51.035979 , -118.142509 ], 2316m , 20.viii.1952 , ♂ , CNCD 13629 , G.P. Holland ( CNC ) . United States of America : Arizona : Rustler Park : w. Portal , [ 31.90556 , -109.27917 ], 2560m , 2.viii.1977 , ♀ , USNM ENT 258093 . COCHISE CO.: [ 31.906851 , - 109.278543 ], 2560m , 13.ix.1965 , ♀ , JSS 19475, C.W. Sabrosky ( CNC ). Bradshaw Mountains , [ 34.41472222 , - 112.4038889 ], 1951m , 7.v.1999 , ♂ , JSS 20683, R . S. Beal ( CSUC ). COCONINO CO .: Oak Creek Canyon , [ 34.961452 , -111.753046 ], ♀ , CNCD 26378 , F.H. Snow ( CNC ). Weimer Spring , [ 34.973632 , -111.528763 ] , 4.vi.1963 , ♂ , JSS 19445, C.H. Spitzer ( CNC ). Flagstaff , [ 35.340306 , -111.670881 ], 3505m , 14.viii.1934 , ♂ , JSS 19435, ♀ , JSS 19436, E.L. Bell ( CNC ). GRAND CANYON : North Rim [ 36.17849 , -113.226775 ] , 17.viii.1966 , JSS 25081, W.J. Hanson ( EMUS ). South Arizona , viii.1902 , 2♀ , CNCD 26379 , JSS 19474, C.W. Johnson ( CNC ) ; viii.1902 , ♂ , JSS 21342, ♀ , JSS 21340, F.H. Snow ( MCZ ). California: RIVERSIDE CO. : Quaking Aspen , [ 33.69554 , -117.163778 ] , 5.vii.1984 , ♂ , JSS 18296, D.J. Burdick ( CNC ); San Jacinto Mountains , [ 33.814491 , - 116.679201 ], ♀ , JSS 19459, J.M. Aldrich ( CNC ). LOS ANGELES CO .: Westwood , [ 34.056121 , -118.430635 ], 17.vi.1959 , ♀ , CNCD 26374 , Kelton & Madge ( CNC ). SAN BERNARDINO CO .: Upper Santa Ana River , [ 34.1003 , - 117.020931 ] , 7.vii.1957 , 3♂ , JSS 19466–19468, 2♀ , JSS 19438, 19465, A.L. Melander ( CNC ). YOSEMITE NATIONAL PARK : Glacier Point Road , [ 37.673917 , -119.637698 ] , 1.vii.1947 , ♀ , JSS 19464, A.L. Melander ( CNC ); Yosemite , [ 37.741951 , -119.577767 ], 1183m , 26.v.1931 , ♂ , JSS 20584, ( EMEC ). TUOLUMNE CO .: Mather Site , Buck Meadows , [ 37.880173 , -119.856977 ], 1970, ♂ , CNCD 26352 , ♀ , CNCD 26375 , A. R . Moldenke ( CNC ). MONO CO .: Tioga Pass , Hall Area site, [ 37.919391 , -119.255072 ], 26.vii.1969 , 2♀ , CNCD 26376 , 26377 , A. R . Moldenke ( CNC ). CALAVERAS BIG TREES SP: Beaver Creek , [ 38.26166667 , -120.2577778 ], 1375m , 22.v.– 11.vi.2007 , 3♂ , JSS 20954–20956, ♀ , JSS 20951, A. R . Cline & P.H. Kerr ( MHPC ). Wright’s Lake , [ 38.84944444 , -120.2269444 ], 2120m , 20.vi.2006 , ♂ , JSS 20962, M. Hauser , S. Winterton , & E. Fisher ( MHPC ). Yuba Pass , [ 39.323059 , -120.599169 ], 2042m , 9.vii.1954 , ♀ , JSS 20589, R . H. Goodwin ( EMEC ). WHITE MOUNTAINS : Blanco’s Corral , 3048m , 30.vi.1953 , ♀ , JSS 20590, J.W. MacSwain ( EMEC ). Truckee , 25.vi.1943 , 3♂ , JSS 23769, 23771, 23772, ♀ , JSS 23770, P.H. Arnaud ( USNM ). S. CA ., v.1908 , 4♂ , JSS 20585–20588, R . Woglum ( EMEC ) . Colorado : SAN JUAN CO .: 10 Miles East of Silverton , [ 37.809565 , -107.485032 ], 2743m , 13.viii, ♂ , CNCD 26351 , F.M. Carpenter ( CNC ). Bennet Crk. , [ 37.941103 , -108.309527 ] , 7.ix.1933 , ♀ , JSS 20707 ( CSUC ). Garfield , [ 38.55185 , -106.292119 ], 3658m , 5.viii.1971 , ♀ , USNM ENT 258077 , W.E. Steiner ( USNM ). Paonia , [ 38.86832 , - 107.592002 ] , 16.ix.1912 , JSS 20711, Chas. L. Fluke ( CSUC ). LAKE CO .: Independence Pass , [ 39.104722 , - 106.557321 ], 3688m , 31.vii.1961 , ♂ , CNCD 26354 , B.H. Poole ( CNC ); Tennessee Pass , [ 39.357246 , - 106.313889 ], 3131m , 7.vii, ♀ , JSS 19458, J.M. Aldrich ( CNC ). Tenmile Range , unamed Cr. avalanche chute W side, [ 39.425 , -106.111 ], 3200m , 15.vii.–17.vii.1998 , ♂ , JSS 20682, S. Fitzgerald & A. Foley ( CSUC ). Rogers Pk. , [ 39.615821 , -105.61778 ], 3660m , 12.vii.1997 , ♂ , JSS 20805 , 10.viii.2002 , ♀ , JSS 20806, C. Slater ( CSUC ). ARAPAHO NATIONAL FOREST , MT . EVANS : Mount Evans , [39.25, -106.1666667], 3658m , 3.viii.1961 , ♂ , CNCD 26355 , ♀ , CNCD 26368 , S.M. Clark ( CNC ); Timberline , [ 39.645575 , -105.594897 ], 3566m , 22.vii.1961 , ♀ , CNCD 26367 , B.H. Poole ( CNC ); Echo Lake , [ 39.659767 , -105.604767 ], 3176m , 29.vi.2010 , 2♂ , CNCD 60033– 60034 , ♀ , JSS 24626, M.M. Locke ( CNC ); Doolittle Ranch , [ 39.675739 , -105.601348 ], 2987m , 10.vii.1961 , ♀ , CNCD 26364 , 3.viii.1961 , ♀ , CNCD 26365 , W. R . M. Mason ( CNC ). BOULDER CO .: Nederland , Science Lodge , [ 40.031645 , -105.533886 ], 2896m , 1.vii.1961 , ♂ , CNCD 26357 , A.A. Bucknell , 28.vi.1961 , ♀ , CNCD 26361 , W. R . M. Mason , 6.vii.1961 , ♀ , CNCD 26362 , 1.vii.1961 , ♀ , CNCD 26363 , B.H. Poole ( CNC ); Niwot Ridge , Near Ward , [ 40.069933 , -105.607397 ], 3505m , 4.vii.1961 , ♂ , CNCD 26356 , ♀ , CNCD 26366 , C.H. Mann, 7.vii.1965 , ♀ , JSS 19446, B. Petersen ( CNC ); Ward , [ 40.072208 , -105.508332 ] , 9.viii.1913 , ♂ , JSS 20712, ( CSUC ). Estes Park , [ 40.376877 , -105.52137 ] , 8.viii.1931 , ♀ , JSS 20669 , 20.v.1933 , ♀ , JSS 20671, ( CSUC ) , viii.1892 , ♂ , JSS 21335, F.H. Snow ( MCZ ). ROCKY MOUNTAIN NATIONAL PARK , Fall R . entrance, [ 40.40344 , -105.589203 ], 4.vi.1999 , ♀ , JSS 20684, D. Leatherman , 1.viii.1997 , ♀ , JSS 20685, D. Leatherman , Manhattan, W . Rustic ( CSUC ). Masonville , [ 40.487171 , -105.210056 ] , 5.ix.1933 , ♀ , JSS 20674, ( CSUC ). Cameron Pass , [ 40.520816 , - 118.227668 ] , 7.vii.1931 , ♀ , JSS 20709, ( CSUC ); 3048m , vii, ♂ , JSS 19981, ( FMNH ). Pingree Park , [ 40.561094 , - 105.597778 ] , 20.viii.1924 , ♀ , JSS 20710, C.L. Fluke , 19.viii.1932 , ♂ , JSS 20668, C.M. Drage , ( CSUC ) , 14.viii.1934 , ♂ , JSS 23764, C.W. Sabrosky ( USNM ). LORY STATE PARK : Well Gulch Trail , [ 40.572996 , - 105.191669 ] , 18.vi.1995 , ♀ , JSS 20809, D. Leatherman ( CSUC ). Fort Collins , [ 40.58333333 , -105.0833333 ] , 19.ix.1942 , ♀ , JSS 20743, R .F. duChanois, 5.x.1921 , ♀ , JSS 20676 , 19.ix.1932 , ♀ , JSS 20680 , 21.ix.1932 , ♀ , JSS 20697 , 20.ix.1932 , ♀ , JSS 20698 , 8.ix.1913 , ♀ , JSS 20742 , 30.ix.1942 , ♀ , JSS 20703, ( CSUC ). Chambers Lake , [ 40.603663 , -105.853645 ] , 25.vi.1932 , ♂ , JSS 20672, 2♀ , JSS 20675, 20701, C. R . Jones ( CSUC ). North Park , [40.73, -106.18] , 1.vii.1932 , 3♂ , JSS 20695, 20696, 20705, 6♀ , JSS 20670, 20673, 20677, 20678, 20699, 20700, ( CSUC ). Glendevey , [ 40.8078 , -105.935501 ] , 7.vii.1932 , ♀ , JSS 20679 , 3.vii.1932 , ♂ , JSS 20704, C. R . Jones, iv.1931 , ♀ , JSS 20708, ( CSUC ). HOMER , 7.ix.1934 , ♀ , JSS 20706, ( CSUC ). Boulder Canyon , 2377m , 8.viii.1960 , ♀ , JSS 21113, R .& K. Dreisbach ( MSUC ). Colorado State , 4♂ , JSS 19450–19453, 2♀ , JSS 19454, 19455 ( CNC ). Idaho : CUSTER CO .: 4th of July Creek , [ 44.39658 , -114.672022 ], 27.vii.1975 , JSS 25139, T . Griswold ( EMUS ). LATAH CO : Mountains Near Moscow , [ 46.807258 , -116.868969 ], 9.vii.1911 , ♂ , CNCD 26358 , ♀ , CNCD 26373 , F.M. Hull, 8.vii.1911 , ♀ , JSS 19470 , 5.vii.1912 , ♂ , JSS 19472 , 6.vii.1912 , ♀ , JSS 19471, ( CNC ). Craig’s Mountain , 29.iv.1916 , ♀ , JSS 21330, ( MCZ ). Holbrook , 4 mi. N.W ., 16.v.1969 , JSS 25080, G.F. Knowlton & G.E. Bohart ( EMUS ). Montana : MISSOULA CO .: Missoula , [ 46.871403 , -113.994267 ] , 22.viii.1904 , ♀ , JSS 19473, ( CNC ). New Mexico : DONA ANA CO.: Las Cruces , [32.3, -106.7666667], 2♂ , CNCD 26359 , 26360 , 2♀ , CNCD 26380 , 26381 , F.M. Hull ( CNC ). LINCOLN CO .: White Mountains , North Fork Ruidoso , [ 33.331649 , -105.672958 ], 2499m , viii.1920 , ♂ , JSS 19440, Townsend ( CNC ). SAN MIGUEL CO .: Trout Spring Canyon , [ 35.659688 , - 105.34231 ] , iv.1927 , ♀ , JSS 19449, ( CNC ). Truchos Peak , [ 35.974028 , -105.636663 ], 4.viii, ♀ , JSS 19456, W.P. Cockerell ( CNC ). Embudo , Rio Arriba , [ 36.207523 , -105.961132 ] , 25.ix.1897 , ♀ , JSS 19441, ( CNC ). Las Valles , Gallinas River 3505m , 20.iv, ♀ , JSS 19437, E.L. Bell ( CNC ). Oregon : KALMOTH CO .: Crater Lake National Park , [ 42.925321 , -122.172722 ], 10.vii.1968 , ♀ , JSS 18297, B. V . Peterson ( CNC ). Sisters : 15 mi S, [ 44.060385 , - 121.542268 ] , 20.vii.1976 , ♂ , USNM ENT 258064 , George Steyskal ( USNM ). WHITMAN NATIONL FOREST : Anthony Lake , [ 44.958805 , -118.23057 ], 2164m , 28.vii.1954 , ♀ , OSAC 133387 , J.H. Baker ( OSAC ), ♂ , JSS 23767, 3♀ , JSS 23765, 23766, 23768, J.H. Baker ( USNM ) , 4.viii.1929 , ♀ , OSAC 133388 , H.A. Scullen ( OSAC ). South Dakota : LAWRENCE CO .: Spearfish Creek , [ 44.589399 , -103.886595 ] , 14.vi.1969 , ♂ , JSS 19476, W.W. Wirth ( CNC ). HARDY W.C. : T3 N, R1 E, S30, 3.vii.1965 , ♂ , CNCD 5445 , R . W. Hodges ( USNM ). Utah: GARFIELD CO .: Henry Mountains , [ 38.028415 , -110.712679 ], 2286m , 29.vii.1968 , ♂ , JSS 18295, J.E.H. Martin ( CNC ). WAYNE CO .: Capitol Reef National Park , [ 38.177815 , -111.176958 ] , 3.x.1984 , JSS 25137, W.P. Nye ( EMUS ). DUCHESNE CO .: Mirror Lake , [ 38.31073 , -112.367492 ], 2743m , 21.vii.–9.viii.1977 , JSS 25135 , 2.viii.1971 , JSS 25138, G.F. Knowlton ( EMUS ). BOX ELDER CO .: Willard Basin , [ 41.394101 , -111.973766 ], 2743m , 26.viii.1964 , JSS 25134, W.J Hanson ( EMUS ). CACHE CO .: Tony Grove Creek , [ 41.886599 , -111.564102 ] , 10.viii.1976 , JSS 25136, ( EMUS ); Green Canyon M. T ., 31.viii.2003 – 25.viii.2003 , ♀ , JSS 24611, W.J. Hanson & Brammer ( EMUS ). DAGGETT CO .: Rt. 44, 2438m, 10.vii.1962 , ♀ , JSS 21110, R .& K. Dreisbach ( MSUC ). SUMMIT CO .: Henry’s Fork Pk. , 2926m , 1.viii.1979 – 10.viii.1979 , ♀ , CNCD 5807 , S. & J. Peck ( CNC ). Washington : Mount Adams , [ 46.204174 , -121.493902 ] , 3.viii.1930 , ♀ , JSS 19460, F.P. Dean ( CNC ). Mt. Ranier , White River , [ 46.782709 , - 121.727952 ] , 20.vii.1924 , ♂ , JSS 23763, A.L. Melander ( USNM ). SPOKANE CO .: Buckeye , [ 47.842393 , - 117.374385 ] , 21.vi.1930 , ♂ , JSS 19444, J.M. Aldrich ( CNC ); Mount Spokane , [ 47.923711 , -117.111646 ] , 22.vi.1930 , ♀ , JSS 19457 , 27.vi.1930 , ♀ , JSS 19448 , 2.vii.1930 , ♀ , JSS 19447, J.M. Aldrich ( CNC ). Wawawai , 28.v.1922 , ♂ , JSS 23762, A.L. Melander ( USNM ). Wyoming : ALBANY CO ., [ 41.727896 , -105.684712 ], 2.viii.1927 , ♂ , CNCD 26353 , C.L. Corkins ( CNC ). Camp Guernsey , Ryan Springs , [ 42.260113 , -104.728624 ] , 31.v.2005 , ♂ , JSS 20686, B. Kondratieff , J. Schmidt , J. Owens ( CSUC ). YELLOWSTONE NATIONAL PARK : Jet Lake , [ 44.550878 , -110.808744 ] , 16.viii.1927 , ♀ , JSS 19439, J.M. Aldrich ( CNC ); Canyon Village , [ 44.587719 , - 110.537673 ] , 21.vii.1971 , ♂ , CNCD 5444 , G.C. Steyskal ( USNM ); Canyon Campground , [ 44.733262 , - 110.486225 ] , 12.viii.1918 , ♂ , JSS 19461, ♀ , JSS 19462, A.L. Melander ( CNC ); Mount Washburn [ 44.797716 , - 110.434368 ], ♂ , JSS 21343, C. T . Brues ( MCZ ). Centennial : 23.viii.1951 , 2♀ , JSS 21109, 21111, R . R . Dreisbach ( MSUC ); Snowy Range , 3048m , 8.ix.1950 , 2♀ , JSS 21105, 21114, R . R . D. & R .K. Schwab , 23.viii.1951 , ♂ , JSS 21108, 2♀ , JSS 21106-21107, R . R . Dreisbach ( MSUC ). Wyoming , 1952, 4♀ , CNCD 26369–26372 , F.M. Hull ( CNC ) .