A taxonomic revision of Bromus (Poaceae: Pooideae: Bromeae) in México and Central America
Author
Saarela, Jeffery M.
Botany Section, Research & Collections Services, Canadian Museum of Nature, PO Box 3443, Station D, Ottawa, Ontario K 1 P 6 P 4, Canada; jsaarela @ mus-nature. ca & Author for correspondence
jsaarela@mus-nature.ca
Author
Peterson, Paul M.
Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington, D. C., 20013 - 7012, U. S. A.; peterson @ si. edu
peterson@si.edu
Author
Valdés-Reyna, Jesus
Botany Section, Research & Collections Services, Canadian Museum of Nature, PO Box 3443, Station D, Ottawa, Ontario K 1 P 6 P 4, Canada; jsaarela @ mus-nature. ca & Departamento de Botanica, Univerisidad Autonoma Agraria " Antonio Narro " Buenavista, Saltillo, Coahuila 25315, MÉXICO; jvaldés @ uaaan. mx & Botany Section, Research & Collections Services, Canadian Museum of Nature, PO Box 3443, Station D, Ottawa, Ontario K 1 P 6 P 4, Canada; jsaarela @ mus-nature. ca
jsaarela@mus-nature.ca
text
Phytotaxa
2014
2014-11-14
185
1
1
147
journal article
5187
10.11646/phytotaxa.185.1.1
4fc1902a-6bcc-45b9-9588-7cc7a150e063
1179-3163
10090001
5a.
Bromus carinatu
s
var. carinatus
.
Figs. 15A–D
,
16
.
Awns
7.1–10 mm
long.
Distribution:―
This variety is not as widespread as
var.
marginatus
. It is known from
Costa Rica
(
Cartago
,
Heredia
,
San José
),
Honduras
(
Ocotepeque
) and
Mexico
(
Baja California
,
Coahuila
,
Chiapas
,
Chihuahua
,
Distrito Federal
,
Durango
,
Guanajuato
,
Jalisco
, Estado de
México
,
Michoácan
,
Nuevo León
,
Oaxaca
,
Puebla
,
San Luis Potosí
,
Sinaloa
,
Tamaulipas
,
Tlaxcala
and
Veracruz
) (
Fig. 17
).
42
Phytotaxa
185 (1) © 2014 Magnolia Press
SAARELA
ET AL.
ATAXONOMIC REVISION OF
BROMUS
(POACEAE)
FIGURE 15.
Bromus carinatus
var.
carinatus
. A. Spikelet. B. Lemma. C. Inflorescence. D. Ligule.
Bromus carinatus
var.
marginatus
. E. Spikelet. F. Lemma. G. Inflorescence. Illustration by C.T. Roché, reproduced from Barkworth
et al.
(2007) with permission.
Phytotaxa
185 (1) © 2014 Magnolia Press
43
FIGURE 16.
Bromus carinatus
var.
carinatus
.
Gould 11621
(MICH-1119195).
44
Phytotaxa
185 (1) © 2014 Magnolia Press
SAARELA
ET AL.
Comments:—
Elsewhere in North America awns in this
variety range
to
15–17 mm
long (
Pavlick & Anderton 2007
,
Saarela & Peterson 2012
). In
México
and Central America we have not seen specimens with awns greater than
10 mm
long.
Specimens Examined:—
COSTA RICA
.
Cartago
:
6 km
SW of Pacayas
, [
9.8°N
,
84.08°W
],
1700 m
,
10 August 1968
,
R
.
W
.
Pohl
&
G
.
Davidse
10893
(CR-46403, F-1732776); km post 19 along road to the crater of
Volcán Irazú
, [
9.97°N
,
83.85°W
],
2600 m
,
1 August 1968
,
R
.
W
.
Pohl
&
G
.
Davidse
10807
(F-1733610);
Volcán Irazú
,
9.9769°N
,
83.8542°W
,
16 August 1964
,
W
.
E
.
Booth
166
(
US-3480655
)
.
Heredia
:
Volcán Irazu
, [
9.97°N
,
83.85°W
], 1912,
A
.
M
.
Brenes
s.n.
(
NY
)
.
San José
:
at km post 51 along the carretera
Interamericana
,
Nof
Empalme
,
1980 m
,
22 August 1968
,
R
.
W
.
Pohl
&
G
.
Davidse
11005
(CR-47205, F-1731856,
US-3578264
)
;
HONDURAS
.
Ocotepeque
:
Mt. Cocal of Cordillera Merendon
20 km
NW
of Ocotopeque
, [
14.49°N
,
89.04°W
],
1800 m
,
25 August 1968
,
A
.
Molina
R. 22140
(F-1675844, NY).
MÉXICO
.
Baja California
:
Ensenada
,
31.8606°N
,
116.6178°W
,
7 April 1882
,
Jones
s.n.
(
RSA-POM-112248
);
Ensenada
,
32.5°N
,
116.6°W
,
7 April 1886
,
C
.
R
.
Orcutt
s.n
. (
US-1009472
);
Sierra de Juárez
,
Parque Nacional Constitución
de 1857,
Laguna Hanson
,
26 May 1987
, [
32.05°N
,
115.9167°W
],
1610–1700 m
,
R
.
F
.
Thorne
,
A
.
Strid
,
K
.
Tan
,
F
.
Ehrendorfer
&
A
.
Liston
62392
(
NY
,
RSA-POM-394998
, F-2047437); just below head of
Prado del Corona
,
1 mi
above end of road from oak pasture and
Melling Ranch
, [
30.75°N
,
115.5167°W
],
7400 ft
,
6 September 1962
,
J
.
D. Olmsted
4737
(
RSA-POM-170872
);
Laguna Hanson
,
Constitución National Park
,
Sierra de Juárez
, [
31.97°N
,
115.83°W
],
1610–1625 m
,
28 May 1983
,
R
.
F
.
Thorne
,
W
.
Wisura
,
W
.
Steinmetz
et al. 55756
(
RSA-POM-310370
);
Northern Lower California
,
April 1886
,
C
.
R
.
Orcutt
s.n.
(
GH
,
MSC-4710
,
NY
);
Lower California
, 1886,
C
.
R
.
Orcutt
s.n.
(
NY
);
Sierra San Pedro Mártir
,
La Grulla
,
30.8917°N
,
115.4833°W
,
2050 m
,
8 June 1982
,
R
.
Moran
30920
(
SD-111378
);
Sierra de Juárez
,
Constitución National Park
,
Laguna Hanson
,
Nend
of lake, [
32.05°N
,
115°W
],
1610–1625 m
,
28 May 1983
,
R.F.
Thorne
,
W
.
Visura
,
W
.
Steinmetz
et al. 44803
(
SD-123836
); below
Rancho
Jatay,
32.025°N
,
116.8542°W
,
80 m
,
11 June 1980
,
R
.
Moran
28792
(
SD-105409
);
Wshore
of
Laguna Hanson
,
32.0542°N
,
115.075°W
,
1610 m
,
21 June 1980
,
R.
Moran
28864
(
SD-105489
);
Sierra San Pedro Mártir
,
La Joya
, on
Valladares Creek
, above creek,
30.925°N
,
115.6°W
,
1500 m
,
31 May 1976
,
R
.
Moran
2332
(
SD-96979
);
Sierra Juárez
,
1 km
SE of Rancho San Pedro
32.1667°N
,
115.4833°W
,
1675 m
,
22 June 1980
,
R
.
Moran
28908
(
SD-105488
);
Sierra San Pedro Mártir
,
Parque Nacional San Pedro Mártir
, at entrance to park in recently burned area, [
31.13°N
,
115.19°W
],
2040 m
,
27 August 1988
,
P
.
M
.
Peterson
,
C
.
R
.
Annable
,
R
.
F
.
Thorne
&
R
.
Noyes
5027
(
CAN
,
US
)
.
Coahuila
:
Sierra Zapalinamé
,
25.3468°N
,
100.908°W
,
2800 m
,
20 September 2003
,
P
.
M
.
Peterson
,
J
.
Valdés-Reyna
&
R
.
H
.
Cárdenas
17871
(
CAN
,
MO
,
US
);
Portrero Valley
, [
26.65 °N
,
112.05°W
],
June 1899
,
C
.
R
.
Orcutt
s.n.
(
US- 1009475
);
Todos Santos Bay
,
31.7667°N
,
116.6167°W
, 1882,
F
.
E
.
Fish
12
(
US-90886
)
.
Chiapas
:
Mpio. Tenejapa
, in the paraje of
Pahal Ton
,
16.85°N
,
92.45°W
,
2286 m
,
1 October 1965
,
D.
E
.
Breedlove
12608
(
DS
,
US-2999875
);
Mpio. Venustiano Carranza
, in the sitios of
Aguacatenango
,
16.45°N
,
92.4667°W
,
1768 m
,
22 July 1965
,
D.
E
.
Breedlove
11226
(
DS
,
MICH-1119255
,
US-2999853
);
Mt. Tacana
,
15.1333°N
,
92.1°W
,
1000–2000 m
,
August 1938
,
E
.
Matuda
2444
(F-1004956,
GH
,
MICH-1119182
,
US-1724501
,
NY
);
Mpio. Tenejapa
,
Paraje Shohleh
,
16.86°N
,
92.49°W
,
2499 m
,
4 April 1966
,
A
.
S
.
Ton
724
(
MICH-1119253
,
US-3005102
);
2 mi
SE of Pueblo Nuevo Solist
,
17.07°N
,
92.86°W
,
1585 m
,
26 July 1965
,
E
.
Lathrop
5971
(
RSA-POM-196173
,
US-2463686
)
.
Chihuahua:
Majalca (Pilares),
28.85°N
,
106.3333°W
,
2080 m
,
12 August 1939
,
L.H. Harvey 1485
(CAN, GH, mixed sheet with
B. frondosus
, MICH-1119267, MO, US-1762718); Cerro Mohinora,
10 mi
Sof
Guadalupe
y Calvo,
26.1°N
,
107.06°W
,
2300–2500 m
,
13 August 1960
,
R.M. Straw & M. Forman 2034
(RSA-POM-178217, mixed sheet with
B. carinatus
var.
marginatus
).
Distrito Federal:
10 mi
SW of México City, [
19.29°N
,
98.95°W
],
9000 ft
,
10 August 1947
,
F.A. Barkley, B.L. Westlund & J.B. Paxson 572
(F-1405758); Mixcoac, Barranca del Manicomio,
19.3833°N
,
99.1833°W
,
2280 m
,
11 August 1913
,
G. Arséne 8801
(F-484987, US-1003469);
10 mi
E of Amecameca on road towards Paso de Cortés, in Parque Nacional Izta-Popo,
19.0764°N
,
98.71°W
,
2840 m
,
12 October 2001
,
P.M. Peterson & Y. Herrera-Arrieta 16163
(CAN, MO,
US
); San Lorenzo, delgacíon de Milpa Alta,
2600 m
,
21 March 1976
,
A. Ventura A. 1191
(ANSM); Lomas de Mixcoac,
19.3833°N
,
99.1833°W
,
17 October 1937
,
E. Lyonnet & J. Elcoro 1731
(MO, US-1746365); Piedregal, Ajusco-Tlalpan,
19.2833°N
,
99.1667°W
,
2600 m
,
16 July 1950
,
E. Matuda 19091
(US-2040915);. Vallede México, Topilojo,
19.2008°N
,
99.1428°W
,
2500 m
,
3 August 1952
,
E. Matuda 26199
(US-2119848).
Durango:
Paraje de Potrillos, entre el Alto de Caballos y el Alto de Potrillos, [
25.4°N
,
106.1°W
],
2500 m
,
6 September 1989
,
A. Benítez P. 814
(ANSM); along road between El Salto and Durango, [
24.08°N
,
105.15°W
],
November 1968
,
Stephenson 68-556
(MSC-391573); Predio particular Las Cebollitas,
1 August 1990
,
A. Benítez 1715
(ANSM);
14.3 mi
Eof La Ventana and
2.7 mi
Wof Los Charcos,
ATAXONOMIC REVISION OF
BROMUS
(POACEAE)
Phytotaxa
185 (1) © 2014 Magnolia Press
45
23.0054°N
,
104.3234°W
,
2875 m
,
13 September 2003
,
P.M. Peterson & F. Sánchez-Alvarado 17766
(CAN, US).
Guanajuato
:
Mpio. Acambaro, Obrajuelos, Altura, [
20°N
,
100.71°W
],
1850 m
,
27 October 1992
,
J.J. Macias Cuellar 36
(MEXU).
Jalisco
:
1–2 mi
Eof Tapalpa,
19.94°N
,
103.76°W
,
2100–2200 m
,
30 October 1960
,
R. McVaugh 20490
(MICH-1119170); Mpio.
Valle
de Juáre, Cerro del Tigre,
97 km
al NE de Cd. Guzmán, [
24.76°N
,
105°W
],
2700 m
,
9 September 1988
,
M. Fuentes O. 698
(MICH-1119187); Mpio.
Vallede
Juáre,
19.5667°N
,
103.6°W
,
19 September 1980
,
A.A. Beetle & R. Guzmán M. M-5370
(SD-122031);
11.7 mi
SW of Ciudad Guzmán,
19.5924°N
,
103.5372°W
,
2165 m
,
5 October 2001
,
P.M. Peterson & O.
Rosales
16081
(CAN, MO, US);
23.5 mi
SW of Ciudad Guzmán, El Refugio, NE slopes of
Nevado
de
Colima
,
19.5871°N
,
103.5972°W
,
3345 m
,
6 October 2001
,
P.M. Peterson & O.
Rosales
16095
(CAN, MO, US);
6.3 mi
SW of Mazamitla along Hwy. 110 towards
Colima
,
19.894°N
,
103.084°W
,
1900 m
,
7 October 2001
,
P.M. Peterson & O.
Rosales
16111
(CAN, MO, US); Parque Nacional Volcán
Nevado
de
Colima
, near
2.5 km
mark from entrance up road to Observatorio Volcanologico,
19.6252°N
,
103.5474°W
,
2008 m
,
22 September 2005
,
P.M. Peterson & F. Sánchez Alvarado 19088
(CAN, MO, US).
Estado de México
:
Toluca,
19.3°N
,
99.65°W
,
9000 ft
,
28 August 1941
,
W. & M. Leavenworth 1926
(F-1248973, GH, MICH-1119176, MEXU-155026, MEXU-5884, MO, NY, US-1815927);
2 mi
Wof state line and ca.
20 mi
SE of
México City
on
Puebla
highway, [
19.33°N
,
98.62°W
],
9 September 1965
,
F.W. Gould 11621
(MICH-119195);
18 mi
Wof Toluca,
19.1083°N
,
99.7583°W
],
15 August 1961
,
F.W. Gould 9601
(US-2432890);
300 m
al Wde Tanango del Aire [
19.16°N
,
98.86°W
],
2350 m
,
14 August 1968
,
A.
Pineda
R. 342
(MICH-1119199);
1 mi
Wof Avila Camacho,
9200 ft
[
2804 m
],
31 July 1970
,
L.H. Harvey 8538
(MICH- 1119242);
27 km
SW of Toluca on road to Temaxcaltepic,
19.11°N
,
100°W
,
2800 m
,
9 July 1964
,
G. Mick & K. Roe 232
(MICH-1119247, US-2630326);
2 mi
above junction of MEX 130 and road to Volcán
Nevado
de Toluca, [
19.1083°N
,
99.7583°W
],
10400 ft
,
22 October 1972
,
L.H. Harvey & J.T. Witherspoon 9473
(MICH-1119121); Temascaltepec Dist., Nanchititla,
18.8667°N
,
100.4667°W
,
10 November 1933
or
11 October 1933
,
G.B. Hinton 4984
(GH, US-1867861);
400 m
al Wde Tanango del Aire, [
19.16°N
,
98.86°W
],
2350 m
,
14 July 1968
,
A.
Pineda
R. 368
(MSC-234697);
10.5 mi
Nof Aculco on Hwy. 55 towards
San Juan
del Río,
20.2247°N
,
99.9°W
,
2208 m
,
8 October 2007
,
P.M. Peterson, J.M. Saarela & M.J. Flores Villegas 21322
(CAN, MO, US);
1.7 mi
Eof Hwy. 10 along road to
Nevado
de Toluca,
19.1347°N
,
99.8°W
,
3688 m
,
9 October 2007
,
P.M. Peterson, J.M. Saarela & M.J. Flores Villegas 21349
(CAN, MO, US); Mpio. Donato Guerra, a
1.25 km
al Nde Las Mesas de
San Martín
, [
19.31°N
,
100.14°W
],
2672 m
,
29 October 1998
,
J.A.
García Ruíz
6
(MEXU); Campo Experimental de la Universidad Autonoma Chapingo,
2260 m
,
1 November 1990
,
O. Arana s.n.
(ANSM); Amecameca,
8600 ft
,
August 1904
,
O. Kuntze 23667
(NY); Popo Park,
19.0333°N
,
98.6333°W
,
4–8 August 1910
,
A.S. Hitchcock 5964
(US-1009500);
2.8 mi
Eof Ocuilán Arteaga near cornfield,
18.9808°N
,
99.38°W
,
2478 m
,
11 October 2007
,
P.M. Peterson, J.M. Saarela & M.J. Flores Villegas 21374
(CAN, MO, US).
Michoácan
:
14 mi
Nof Aguililla on road to Dos Aguas,
1570 m
,
8 August 1972
,
M.F. Denton 2036
(MICH-1119171); at km 56–57 above Quiterio on road from Pátzcuaro to Tacambaro,
19.4°N
,
101.5°W
,
2700 m
,
H.E. Moore, Jr., E. Hernández X. & H. Porras H. 5593
(GH, US-1983689); Estacíon Chincua, Reserva de la Biósfera Mariposa Monarca,
19.6792°N
,
100.2853°W
,
3030 m
,
5 August 2000
,
Ma. G. Cornejo Enotrio, S.R. Luna & G.I. Manríquez 65
(ANSM, MEXU);
10 km
al Sde Indaparapeo, sobre el camino a Las Peras, [
19.72°N
,
100.9181°W
],
2250 m
,
23 December 1986
,
J. Rzedowski 42310
(ANSM); Mpio. de Queréndaro, alrededores de
San José
Lagunillas,
2750 m
,
24 August 1986
,
J. Rzedowski 40468
(ANSM); Mpio. Huaniqueo, NE del pedregal pequeño,.
5 km
al Wde Tendeparacua,
2060 m
,
22 August 1992
,
P. Silva-Sáenz 75
(ANSM); ca.
18 mi
Sof Pátzcuaro,
19.31°N
,
101.66°W
,
2713–2743 m
,
20–25 November 1961
,
R.M. King & T.R. Soderstrom 5178
(NY, mixed sheet with
B. dolichocarpus
); ca.
2 km
al Nde AJuno, Cerro La Taza,
2280 m
,
2 September 2003
,
M.E.
Molina
327
(ANSM); Lieux inondes a' l'ouest, vicinity of Morelia,
19.7167°N
,
101.1833°W
,
1800 m
,
7 October 1909
,
G. Arséne 3166
(MO, US-1002651); near Zacapú,
19.8167°N
,
101.7833°W
,
28 September 1946
,
E. Hernández-Xolocotzi, J. Ruppert & J. Guevara
X-2825
(US-1962215); ca.
15 mi
Eof Mil Cumbres on road from Morelia to Toluca,
24 July 1950
,
J.R. Reeder, C.G. Reeder & L.N. Goodding 1477
(RSA-POM-253305, US-2473569).
Nuevo León
:
road to La Tinaja from roadside in disturbed area,
23.8873°N
,
99.7956°W
,
2500 m
,
21 September 2002
,
P.M. Peterson, J. Valdés-Reyna & M. Sosa Morales 16765a
(ANSM, CAN, US); Sierra Infiernillo, Cañón San Francisco, ca.
15 km
al NE de Pablillo,
24.55°N
,
99.9°W
,
1800–1900 m
,
4 September 1993
,
J. Valdés-Reyna 2321, M.A. Carranza & R. Banda S.
(ANSM); Sierra Las Cautivas,
8.7 mi
SW of Dulces Nombres and
18.1 mi
Eof Zaragoza,
23.9501°N
,
99.6473°W
,
2385 m
,
26 September 2007
,
P.M. Peterson & J.M. Saarela 21099
(CAN, MO, US).
Oaxaca
:
1.2 mi
Eof
San José
del Pacífico on road towards San Sebastián,
16.1698°N
,
96.4898°W
,
2641 m
,
22 September 2008
,
P.M. Peterson & J.M.
46
Phytotaxa
185 (1) © 2014 Magnolia Press
SAARELA
ET AL.
Saarela 22352
(CAN,
US
);
1.5 mi
Wof
San Juan Chicomezuchitl
,
17.3026°N
,
96.5016°W
,
1535 m
,
19 September 2008
,
P
.
M
.
Peterson
&
J
.
M
.
Saarela
22286
(
CAN
,
US
)
;
2.7 mi
SW of San Angustín Mixtepec
on road toward
Cieneguilla
,
16.2509°N
,
96.3507°W
,
2560 m
,
28 September 2006
,
P
.
M
.
Peterson
&
R
.
Garcia
20244
(
CAN
,
MO
,
P-03328784
,
US
).
Puebla
:
5 mi
Eof
Río Frio
,
19.34°N
,
98.59°W
,
2800 m
,
20 July 1953
,
J
.
R
.
Reeder
&
C
.
G
.
Reeder
1915
(
SD-101483
,
US
);
Parada Oligui
,
entre Teziutlán y Tlapacoyan
,
3 June 1968
, [
19.82°N
,
97.36°W
],
1850 m
,
D.
García Saucedo
95
(
MICH-1119193
,
MSC-234698
);
Mayorazgo
, sur
l'Atoyac
, vicinity of
Puebla
,
19.033°N
,
98.2°W
,
2120 m
,
18 July 1907
,
G
.
Arséne
1405
(
US-1002644
) &
1407
(
MO
,
US-1002658
); SW slopes of
Volcán Citlaltépetl
(
Pico de Orizaba
), below
Tesmalaquilla
,
18.95°N
,
97.29°W
,
2970 m
,
26 September 1962
,
D. Ugent
&
R
.
Flores-C. 2334
(
RSA-POM-264936
,
US-2543174
).
San Luis Potosí
:
±
39 km
carretera
S
.
L
.
P
.
to
Rioverde
S
.
L
.
P
.,
22 August 1961
,
A
.
Gómez
190
(
ANSM
);
Álvarez
, [
22.0333°N
,
100.6167°W
],
23–23 July 1904
,
E
.
Palmer
171
(
NY
)
;
3 mi
Nof
Los Hornos
, along road to
Ocurahui
, [
25.6833°N
,
108.4833°W
],
6500 ft
,
2 October 1970
,
D.
E
.
Breedlove
&
R
.
F
.
Thorne
18417
(
RSA-POM-222535
).
Sinaloa
:
Ocurahui
,
Sierra Surotato
,
6000–7000 ft
,
1–10 September 1941
,
H
.
S
.
Gentry
6380
(
GH
,
MICH-1119177
,
MO
,
NY
)
;
3 mi
Nof
Los Ornos
, along road to
Ocurahui
, [
25.6833°N
,
108.4833°W
],
6500 ft
,
2 October 1970
,
D.
E
.
Breedlove
&
R
.
F
.
Thorne
18417
(
RSA-POM- 222535
).
Tamaulipas
:
Sierra Las Cautivas
,
0.3 mi
Wof
Dulces Nombres
at edge of community,
24.0031°N
,
98.81°W
,
1984 m
,
26 September 2007
,
P
.
M
.
Peterson
&
J
.
M
.
Saarela
21074
(
CAN
,
MO
,
NY
,
US
);
Sierra Las Cautivas
,
23.1 mi
Wof
Ejido Guayabas
along road towards
Dulces Nombres
,
23.9756°N
,
99.49°W
,
1844 m
,
25 September 2007
,
P
.
M
.
Peterson
&
J
.
M
.
Saarela
21061
(
CAN
,
US
).
Tlaxcala
:
Mpio. Cuapiaxtla
,
2405 m
,
24 July 1974
,
E
.
Hernández X.
et al. R-79
(
ANSM
);
Contadero
,
19.4333°N
,
98.5167°W
,
3 August 1901
,
J
.
N
.
Rose
&
R
.
Hay
5966
(
NY
,
US-395757
).
Veracruz
:
Mpio. Altotonga
,
Nahualaco
,
19.76°N
,
97.24°W
,
1750 m
,
24 November 1969
,
F
.
Ventura
A
.
108
(
MICH-1119202
,
MSC-234831
);
Mpio. Calcahualco
,
9.5 km
by road
Wof
Escola
along Coscomatepec-Escola-Jacal road,
19.1167°N
,
97.08°W
,
2600 m
,
15 November 1981
,
M
.
Nee
23173
(F-1975254,
MO
,
NY
);
Orizaba
, [
18.51°N
,
97.06°W
],
August 1853
,
F
.
Müller
2114
(
NY
);
Jalapa
,
19.5333°N
,
96.9167°W
,
1402 m
,
2–4 September 1910
,
A
.
S
.
Hitchcock
6592
(
US-1009524
)
.