The Goblin Spider Genus Ischnothyreus (Araneae, Oonopidae) in the New World Author Platnick, Norman I. Author Dupérré, Nadine Author Ubick, Darrell Author Fannes, Wouter text American Museum Novitates 2012 2012-09-21 2012 3759 1 32 http://www.bioone.org/doi/abs/10.1206/3756.2 journal article 10.1206/3756.2 0003-0082 10109266 Ischnothyreus peltifer (Simon) Figures 1–99 Ischnaspis peltifer Simon, 1891: 562 ( four female syntypes from St. Vincent , in BMNH, one female syntype from St. Vincent , in MNHN; examined). Ischnothyreus peltifer: Simon, 1893a: 298 (not male, fig. 264, = Camptoscaphiella simoni Baehr ). – Petrunkevitch, 1929: 66 , figs. 51, 52. – Chickering, 1951: 219 , fig. 7. – Chickering, 1968: 80 , figs. 1–10. – Dumitresco and Georgesco, 1983: 96 , pl. 17, figs. 1–9, pl. 18, figs. 1–4. – Saaristo, 2001: 345 , figs. 146A, 147A–C, 148A–B, 149, 150, 152–154, 155A, 156, 157A–B. – Saaristo and van Harten, 2006: 135 , figs. 15a–b, 16a–c, 17. – Platnick et al., 2012a: 12 , figs. 16–20, 22, 23, 25 (not figs. 21, 24, = I. velox ). Dysderina antillana Bryant, 1942: 324 , figs. 1, 7 (male holotype from Christiansted, St. Croix, Virgin Islands , in MCZ; examined). First synonymized by Chickering, 1968: 80 . Ischnothyreus velox (misidentification): Bristowe, 1948: 890 (in part, male, fig. 1). – Locket and Millidge, 1951: 76 (in part, male, fig. 38E). Ischnothyreus omus Suman, 1965: 226 , figs. 1–8 (male holotype from Kailua, Oahu, Hawaii , in Bishop Museum; not examined). – Chen and Zhang, 1991: 63 , figs. 54.1–5 ( I. onus , lapsus). – Song, Zhu, and Chen, 1999: 69 , figs. 28I–K. – Ono, 2009: 103 , figs. 25– 31. First synonymized by Saaristo, 2001: 345 ( I. omosus , lapsus). Ischnothyreus formosus Brignoli, 1974: 80 , figs. 12–18 (male holotype from Akau, Taiwan , in Zoological Museum, Hamburg; not examined). – Song, 1987: 91 , fig. 55. – Chen and Zhang, 1991: 62 , figs. 53.1–7. – Song, Zhu, and Chen, 1999: 69 , figs. 28F–H. First synonymized by Saaristo, 2001: 345 . Ischnothyreus sechellorum Benoit, 1979: 208 , figs. 7A–E (female holotype from Mahe , Sechelles, in MRAC; examined). First synonymized by Saaristo, 1999: 3 . DIAGNOSIS: Males can easily be separated from those of I. velox by the large protuberance on the base of the cheliceral fang (figs. 5–8, 83), females by the narrow, posteriorly situated, sinuous ridge on the postepigastric scutum (fig. 72). MALE (PBI_OON 16071, figs. 1–45, 76–87): Total length 1.48. Cephalothorax: Carapace olive green, with dark brown egg-shaped patches behind eyes, ovoid in dorsal view (fig. 1), pars cephalica strongly elevated in lateral view (fig. 2), anteriorly narrowed to 0.49 times its maximum width or less, with rounded posterolateral corners, posterolateral edge without pits, posterior margin not bulging below posterior rim, anterolateral corners with slightly sclerotized triangular projections, posterolateral surface without spikes, surface of elevated portion of pars cephalica smooth, sides reticulate, thorax without depressions, fovea absent, without radiating rows of pits; lateral margin straight, smooth, without denticles; plumose setae near posterior margin of pars thoracica absent; nonmarginal pars cephalica setae light, needlelike, scattered; nonmarginal pars thoracica setae absent; marginal setae light, needlelike. Clypeus margin unmodified, straight in front view, vertical in lateral view, high, ALE separated from edge of carapace by their radius or more (fig. 3), median projection absent; setae light, needlelike. Chilum absent. Eyes six, well developed, ALE largest, ALE circular, PME oval, PLE oval; posterior eye row procurved from both above and front; ALE separated by less than their radius, ALE-PLE separated by less than ALE radius, PME touching throughout most of their length, PLE-PME touching (fig. 4). Sternum slightly longer than wide (fig. 12), pale orange, uniform, not fused to carapace, median concavity absent, without radial furrows between coxae I–II, II–III, III–IV, radial furrow opposite coxae III absent, surface smooth, without pits, microsculpture absent, sickle-shaped structures absent, anterior margin with continuous transverse groove, posterior margin not extending posteriorly of coxae IV, anterior corner unmodified, lateral margin with infracoxal grooves and anterior and posterior openings (figs. 13, 14), distance between coxae approximately equal, extensions of precoxal triangles present, lateral margins unmodified, without posterior hump; setae scattered, light, needlelike, more abundant on anterior half, originating from surface, without hair tufts. Chelicerae, endites and labium pale orange. Chelicerae straight, anterior face unmodified; with one tooth on both promargin and retromargin (figs. 5, 6); fangs without toothlike projections, directed posteriorly, shape normal, with prominent basal process (figs. 7, 8), tip unmodified; setae light, needlelike, evenly scattered; paturon inner margin with scattered setae, distal region unmodified, posterior surface unmodified, promargin unmodified, inner margin with many medial denticles, laminate groove absent. Labium rectangular, fused to sternum, anterior margin indented at middle, same as sternum in sclerotization; with six or more setae on anterior margin, subdistal portion with unmodified setae (fig. 9). Endites distally excavated, serrula absent (fig. 10), anteromedian tip with one strong, tooth-shaped projection, posteromedian part unmodified, same as sternum in sclerotization. Labrum with triangular projection (fig. 11); palps connected to complexly modified endosternite (figs. 43–45). Abdomen: ovoid, without long posterior extension, rounded posteriorly, interscutal membrane rows of small sclerotized platelets absent posteriorly; dorsum soft portions white, without color pattern. Book lung covers large, elliptical, without setae, anterolateral edge unmodified. Posterior spiracles not connected by groove. Pedicel tube short, unmodified, scutopedicel region unmodified, scutum extending far dorsal of pedicel, plumose setae, matted setae on anterior ventral abdomen in pedicel area, cuticular outgrowths near pedicel all absent. Dorsal scutum weakly sclerotized, olive green, without color pattern, covering 1/2 to 3/4 of abdomen, more than 1/2 to most of abdomen width, fused to epigastric scutum, middle surface smooth, sides smooth, anterior half without projecting denticles. Epigastric scutum weakly sclerotized, surrounding pedicel, not protruding, small lateral sclerites absent. Postepigastric scutum weakly sclerotized, pale orange, short, almost rectangular, covering about 2/3 of abdominal length, fused to epigastric scutum, anterior margin unmodified, without posteriorly directed lateral apodemes. Spinneret scutum present, incomplete ring. Dorsal, epigastric, postepigastric setae light, needlelike; epigastric setae not basally enlarged; spinneret scutum with fringe of needlelike setae; dense patch of setae anterior to spinnerets absent; interscutal membrane with setae. Colulus present, very small, bearing two hairs. Spinnerets (fig. 17): ALS with one major ampullate gland spigot and three piriform gland spigots (fig. 18); PMS with single spigot (fig. 19); PLS with three spigots (fig. 20). Legs: pale orange, without color pattern; femur IV not thickened, same size as femora I–III, patella plus tibia I shorter than carapace, tibia I unmodified, tibia IV specialized hairs on ventral apex absent, tibia IV ventral scopula absent, metatarsi I, II mesoapical comb absent, metatarsi III, IV weak ventral scopula absent, tarsi III, IV with “false claws” (figs. 23, 27, 28). Leg spination (only surfaces bearing spines listed, all spines longer than segment width): femora: I p0-1-1; II p0-0-1; tibiae I, II v4-2-2; metatarsi I, II v2-2-0. Tarsal proclaws and retroclaws inner face striate, with zero teeth on lateral surface, three teeth on median surface (figs. 21–28); inferior claw absent. Trichobothria: tibia each with three, metatarsi each with one; hood covered by numerous low, closely spaced ridges (fig. 33). Tarsal organ with three receptors on legs I, II, two receptors on legs III, IV, palps (figs. 29–32). Genitalia: Epigastric region with sperm pore large, triangular with rounded angles (figs. 15, 16), situated in front of anterior spiracles, unmodified; epigastric furrow apparently absent (fig. 15). Palp slightly reduced in size, strongly sclerotized, right and left palps symmetrical, proximal segments almost black; trochanter normal size, with ventral projection; femur normal size, one to two times as long as trochanter, without posteriorly rounded lateral dilation, attaching to patella basally; patella about as long as femur, not enlarged, without prolateral row of ridges, setae unmodified; tibia with three trichobothria; cymbium almost black, ovoid in dorsal view, fused with bulb but with clearly defined seam between, not extending beyond distal tip of bulb, plumose setae, stout setae, distal patch of setae all absent; bulb almost black, 1–1.5 times as long as cymbium, stout, gradually tapering apically, obtusely bent before apex (figs. 34, 35), middle part with two protuberances on ventral side (fig. 36); embolus light, prolateral excavation absent, set off from bulb by ventral collar, tip with multiple, complex processes (figs. 37–42). FIGURES 1–15. Ischnothyreus peltifer (Simon) , male. 1. Carapace, dorsal view. 2. Same, lateral view. 3. Same, anterior view. 4. Eyes, anterior view. 5. Chelicerae, anterior view. 6. Same, posterior view. 7. Fang, anterior view. 8. Same, posterior view. 9. Labium and endites, ventral view. 10. Labrum and endites, dorsal view. 11. Labral projection, dorsal view. 12. Sternum, ventral view. 13. Anterolateral corner of sternum, ventral view. 14. Side of sternum, oblique lateral view. 15. Epigastric scutum, ventral view (4, 13, 14 courtesy of Alexandre Bonaldo). FIGURES 16–30. Ischnothyreus peltifer (Simon) , male. 16. Sperm pore, ventral view. 17. Spinnerets, apical view. 18. Anterior lateral spinneret, same. 19. Posterior median spinneret, same. 20. Posterior lateral spinneret, same. 21. Claws of leg I, apical view. 22. Same, leg II. 23. Same, leg III. 24. Same, leg IV. 25. Claws of leg I, lateral view. 26. Same, leg II. 27. Same, leg III. 28. Same, leg IV. 29. Tarsal organ from leg I, dorsal view. 30. Same, leg II. FIGURES 31–45. Ischnothyreus peltifer (Simon) , male. 31. Tarsal organ from leg III, dorsal view. 32. Same, leg IV. 33. Trichobothrial base from metatarsus I, dorsal view. 34. Left palp, prolateral view. 35. Left palpal bulb, prolateral view. 36. Same, ventral view. 37. Left palp, retrolateral view. 38. Left palpal bulb, same. 39. Tip of palpal bulb, distal view. 40. Terminal sclerites of palpal bulb, distal view. 41. Same, oblique view, tip of leafshaped sclerite broken off. 42. Embolar opening, apical view. 43. Right palp and associated endoskeletal elements, ventral view. 44. Left half of endoskeletal elements, same. 45. Middle portion of endoskeletal elements, same (36, 38, 43–45 courtesy of Alexandre Bonaldo). FIGURES 46–60. Ischnothyreus peltifer (Simon) , female. 46. Carapace, dorsal view. 47. Same, lateral view. 48. Same, anterior view. 49. Chelicerae, anterior view. 50. Same, posterior view. 51. Labium and endites, ventral view. 52. Sternum, ventral view. 53. Palp, prolateral view. 54. Same, retrolateral view. 55. Spines on palpal femur, lateral view. 56. Palpal tibia, dorsal view. 57. Claws of leg II, apical view. 58. Same, leg III. 59. Same, leg IV. 60. Claws of leg I, lateral view. FIGURES 61–75. Ischnothyreus peltifer (Simon) , female. 61. Claws of leg II, lateral view. 62. Same, leg III. 63. Same, leg IV. 64. Tarsal organ from leg II, dorsal view. 65. Same, leg III. 66. Same, leg IV. 67. Same, palp. 68. Spinnerets, apical view. 69. Anterior lateral spinneret, same. 70. Posterior median spinneret, same. 71. Posterior lateral spinneret, same. 72. Postepigastric scutum, ventral view. 73. Anteromedian portion of postepigastric scutum, same. 74. Median portion of postepigastric scutum, same. 75. Genitalia, dorsal view. FIGURES 76–87. Ischnothyreus peltifer (Simon) , male. 76. Habitus, dorsal view. 77. Same, ventral view. 78. Abdomen, ventral view. 79. Carapace, dorsal view. 80. Same, lateral view. 81. Cephalothorax, ventral view. 82. Carapace, anterior view. 83. Chelicera, posterior view. 84. Habitus, lateral view. 85. Left palp, prolateral view. 86. Same, ventral view. 87. Same, retrolateral view. FIGURES 88–99. Ischnothyreus peltifer (Simon) , female. 88. Carapace, dorsal view. 89. Cephalothorax, ventral view. 90. Abdomen, dorsal view. 91. Carapace, anterior view. 92. Abdomen, ventral view. 93. Genitalic region, ventral view. 94. Habitus, lateral view. 95. Genitalia, ventral view. 96. Same, dorsal view. 97. Labrum and endites, dorsal view. 98. Labral projection, dorsal view. 99. Serrula, dorsal view. FIGURES 100–111. Ischnothyreus velox Jackson , male. 100. Carapace, dorsal view. 101. Sternum, ventral view. 102. Carapace, anterior view. 103. Same, lateral view. 104. Mouthparts, ventral view. 105. Habitus, lateral view. 106. Abdomen, dorsal view. 107. Same, ventral view. 108, 109. Left palp, prolateral view. 110. Same, ventral view. 111. Same, retrolateral view. FIGURES 112–120. Ischnothyreus velox Jackson , male. 112, 119. Left palp, prolateral view. 113. Same, ventral view. 114, 120. Same, retrolateral view. 115. Left embolus, prolateral view. 116. Same, ventral view. 117. Same, retrolateral view. 118. Same, distal view. FIGURES 121–132. Ischnothyreus velox Jackson , female. 121. Carapace, dorsal view. 122. Same, anterior view. 123. Habitus, lateral view. 124. Sternum, ventral view. 125. Habitus, dorsal view. 126. Abdomen, ventral view. 127, 128, 130. Genitalia, ventral view. 129. Same, dorsal view. 131. Postepigastric scutum, ventral view. 132. Copulatory opening, ventral view. FIGURES 133–144. Ischnothyreus browni Chickering , male. 133. Carapace, dorsal view. 134. Same, lateral view. 135. Same, anterior view. 136. Sternum, ventral view. 137. Abdomen, dorsal view. 138. Same, ventral view. 139. Same, lateral view. 140, 142. Left palp, prolateral view. 141, 144 . Same, retrolateral view. 143. Same, ventral view. FIGURES 145–161. Ischnothyreus browni Chickering , male (145–151) and female (152–161). 145. Left palp, prolateral view. 146. Same, ventral view. 147. Same, retrolateral view. 148. Left embolus, prolateral view. 149. Same, ventral view. 150. Same, retrolateral view. 151. Same, distal view. 152. Abdomen, dorsal view. 153. Same, ventral view. 154. Carapace, lateral view. 155. Abdomen, lateral view. 156. Carapace, dorsal view. 157. Same, anterior view. 158. Sternum, ventral view. 159, 160. Genitalia, ventral view. 161. Same, dorsal view. FEMALE (PBI_OON 16071, figs. 46–75, 88–99): Total length 1.70. As in male except as noted. Carapace without any pattern, broadly oval in dorsal view (fig. 46), pars cephalica slightly elevated in lateral view (figs. 47, 48). Sternum anterior margin unmodified (fig. 52). Fangs directed medially, without prominent basal process (figs. 49, 50). Labium anterior margin not indented at middle. Endites distally not excavated, serrula present in single row (figs. 51, 97, 99), anteromedian tip unmodified; labral projection rounded (fig. 98). Palpal claw absent (figs. 53, 54); femur with three spiniform setae ventrally (fig. 55); patella without prolateral row of ridges; tibia with three trichobothria (fig. 56); tarsus unmodified. Dorsal scutum between 1/4 and 1/2 abdomen width, not fused to epigastric scutum. Epigastric scutum slightly protruding, small lateral sclerites present (fig. 93), without lateral joints. Postepigastric scutum widely hexagonal, covering about 1/3 of abdominal length, not fused to epigastric scutum, with short, posteriorly directed lateral apodemes. Spinnerets (fig. 68): ALS as in male (fig. 69); PMS with four spigots (fig. 70); PLS with seven spigots (fig. 71). Leg spination: femora: I p0-1-1; II p0-0-1; tibiae I, II v4-2-2; metatarsi I, II v2-2-0. Tarsal organs (figs. 64–67) and claws (figs. 57–63) as in male except claws of leg IV with elongated distal tooth, tip therefore appearing bifid (fig. 59). Posterior margin of epigastric scutum with transverse row of large setae. Postepigastric scutum with narrow, sinuous, elevated ridge at about two-thirds its length (fig. 72); narrow, slit-shaped copulatory opening situated on midline near anterior edge of postepigastric scutum (fig. 73), followed posteriorly by series of tiny pores (fig. 74). Posterior genitalic tube originating at middle of anterior edge of postepigastric scutum, squiggled, ending at level of external ridge (fig. 75); anterior receptaculum small, T-shaped. MATERIAL EXAMINED: NORTH AMERICA: Canada : Ontario : African Pavilion, Toronto Metropolitan Zoo, Mar. 17, 1987 (J. Swann, CNC PBI_OON 38143), 3♂ , 2♀ . United States : Florida : Alachua Co.: San Felasco State Park, Gainesville, Sept. 24, 2007 , leaf litter (X. Wang, AMNH PBI_OON 31070), 1♂ ; Univ. of Florida campus, Gainesville, Sept. 24, 2007 , leaf litter (X. Wang, AMNH PBI_OON 31071), 3♂ , 1♀ . Collier Co.: Collier-Seminole State Park, Apr. 1, 1985 , leaf litter (W. Maddison, MCZ 70476, PBI_ OON 26768), 1♂ , 2♀ . Lee Co.: Fort Myers, 26°38′N , 81°50′W , Mar. 18, 1954 (W. Ivie, AMNH PBI_OON 1025), 2♀ . Martin Co.: Bluefield, Aug. 4, 1983 , on casuarina (K. Hibbard, FSCA PBI_OON 21194), 1♀ . Miami-Dade Co.: Deering Estate Park, SW 167 St., SW 72 Ave., Miami, Nov. 15, 1985 , hammock forest litter (S. Peck, AMNH PBI_OON 1971), 1♂ , Feb. 21–June 1, 1986 , young hammock forest, malaise flight intercept trap (S., J. Peck, AMNH PBI_OON 1040), 2♂ , 1♀ , June 1–Aug. 25, 1986 , same (S., J. Peck, AMNH PBI_OON 1038), 11♂ , 9♀ , July 26–Dec. 11, 1986 , old hammock, malaise flight intercept trap (S., J. Peck, AMNH PBI_OON 1032), 1♀ ; Grossman Hammock, Chekika State Recreation Area, 50 km SW Miami, Nov. 15, 1985Feb. 24, 1986 , malaise flight intercept traps (S., J. Peck, AMNH PBI_OON 1026), 2♀ ; Homestead, Mar. 12–25, 1968 , avocado grove (A. Chickering, MCZ 68268, PBI_OON 27430), 1♂ , 5♀ ; Long Pine Key, Everglades National Park, May 28–June 8, 1986 , pinelands, malaise flight intercept traps (S., J. Peck, AMNH PBI_OON 1034, 1041), 2♂ , 2♀ , June 8–Dec. 9, 1986 , same (S., J. Peck, AMNH PBI_OON 1972, 1973), 3♂ , 15♀ , Aug. 28–Sept. 5, 1986 , same (S., J. Peck, AMNH PBI_OON 1036), 1♂ , 1♀ ; Royal Palm Hammock, Everglades National Park, July 1981 , malaise trough (S. Peck, CNC PBI_OON 38144), 3♂ , 2♀ , Nov. 1, 1984Mar. 3, 1985 , hammock forest, malaise flight intercept traps (S., J. Peck, AMNH PBI_OON 1039), 1♂ , 6♀ , May 2–Aug. 2, 1985 , same (S., J. Peck, AMNH PBI_OON 1027), 1♂ , 2♀ , Dec. 9, 1986 , forest litter (S. Peck, Klimaszweski, AMNH PBI_OON 158), 1♀ . Palm Beach Co.: 5 mi S Lake Harbor on Miami Canal, Aug. 18, 1978 , pitfall, sugarcane field (C. Adams, CNC PBI_ OON 38140), 1♀ , Oct. 20, 1978 , same ( CNC PBI_OON 38139), 1♀ . Saint Lucie Co.: Bluefield, Aug. 24, 1983 , on casuarina (K. Hibbard, FSCA PBI_OON 21195), 2♀ ; Indrio, Aug. 17, 1983 , on cenchrus (K. Hibbard, FSCA PBI_OON 21193), 1♀ ; White City, Aug. 11, 1983 , on casuarina (K. Hibbard, FSCA PBI_OON 21196), 1♂ , 1♀ , Sept. 27, 1983 , on casuarina (K. Hibbard, FSCA PBI_OON 21192), 2♀ . Mexico : Quintana Roo : Cozumel, July 14, 1951 (L. Stannard, AMNH PBI_OON 1968), 2♀ . Tabasco : Parque La Venta, Villahermosa, 18°00′N , 92°53′W , Aug. 13, 1966 (J., W. Ivie, AMNH PBI_OON 160, 1385, 1967, 1970), 9♂ , 6♀ ; Peje Lagartero, 18°03′N , 93°10′W , Aug. 14, 1966 (J., W. Ivie, AMNH PBI_ OON 135), 2♂ , 2♀ ; Villahermosa, Apr. 3, 1969 ( AMNH PBI_OON 1969), 1♀ . Veracruz : 7 mi W Coatzacoalcos, 18°08′N , 94°30′W , Aug. 31, 1964 (J., W. Ivie, AMNH PBI_OON 168, 187), 4♂ ; 10 mi W Coatzacoalcos, 18°08′N , 94°32′W , Aug. 15, 1966 (J., W. Ivie, AMNH PBI_OON 132), 1♂ , 2♀ . Yucatán : Mérida. Feb. 2, 1939 ( AMNH PBI_OON 1012), 1♀ . CENTRAL AMERICA: Belize : Belize : Burrell Boom, July 28, 1972 , Berlese (C. Goodnight, AMNH PBI_OON 1016), 1♀ . Costa Rica : Alajuela : Monterrey de Upala, Jan. 10–12, 2009 , elev. 50 m (A. Solis, C. Víquez, R. Guries, INBIO PBI_OON 49460), 1♂ ; Muelle, San Carlos, Nov. 7, 2007 , in termite nest (C. Víquez, INBIO PBI_OON 49459), 1♀ ; Upala, Montecristo, 10°54′22″N , 84°58′09″W (INBIO PBI_OON 49451, 49453, 49455, 49456, 49461, 49462), 4♂ , 6♀ , Apr. 13–June 8, 2007 , humus, elev. 60 m (C. Víquez, INBIO PBI_OON 26369, 26371, 26372, 26374, 26375, 36979), 5♂ , 3♀ , 10.9668°N , 85.04363°W , Aug. 12–19, 2007 , cacao farm (C. Víquez, INBIO PBI_OON 49457), 3♀ , Aug. 15–17, 2009 , cacao (A. Solis, C. Víquez, R. Guries, INBIO PBI_OON 49458), 1♂ . Cartago : Turrialba, 9°58′N , 83°38′W , July 25–Aug. 15, 1965 (A. Chickering, MCZ 68267, PBI_OON 26769), 19♂ , 90♀ , Feb. 17, 1981 , banana stem (A. Young, AMNH PBI_OON 49442), 1♀ . Heredia : La Selva, 10.422159°N , 84.001525°W , Berlese, Aug. 9–15, 2010 (INBIO 99380, PBI_OON 49454), 1♀ ; Llorente de Flores, Feb. 4–Mar. 17, 2007 , coffee plantation, leaf litter (C. Víquez, INBIO PBI_OON 29680, 29682, 49463), 3♂ , 3♀ ; Parque Municipal Nacimiento, 9°58′57″N , 84°10′28″W , Jan. 27–28, 2010 , humus, elev. 950 m (C. Víquez, B. Hernandez, A. Solis, INBIO PBI_OON 49452), 1♂ , 2♀ ; San Joaquín, July 6, 1997 , humus (C. Víquez, INBIO PBI_OON 26368), 2♂ , 10.006214°N , 84.153652°W , Sept. 8, 2007 , garden humus, elev. 1050 m (C. Víquez, INBIO PBI_OON 26370), 2♂ , 1♀ ; San Lorenzo, road to San Pedro, Río Secundo bridge, 10°01′13″N , 84°08′46″W , Feb. 24, 2007 , Berlese, elev. 1090 m (C. Víquez, INBIO PBI_ OON 29681), 1♂ ; Santa Bárbara , San Juan, 10.02153°N , 84.16626°W , May 18, 2008 , Berlese, coffee plantation, elev. 1065 m (C. Víquez, INBIO PBI_OON 31119), 1♂ . Limón : Parque Nacional Tortuguero, Apr. 17–23, 1983 , wet second growth forest (D. Ubick, CDU PBI_OON 35943), 1♂ . Puntarenas : Sendero Río Madrigal, Estación La Leona, Península de Osa, Apr. 4, 2002 , humus, secondary forest (A. Azofeifa, INBIO PBI_OON 26373), 1♀ . Panama : Colón : Frijoles, Jan. 25, 1958 (A. Chickering, MCZ 71371, PBI_ OON 27254), 2♂ , 5♀ ; Gamboa, July 24, 1954 (A. Chickering, MCZ 71394, PBI_OON 27246), 1♀ , Jan. 1958 (A. Chickering, MCZ 71392, PBI_OON 27249), 18♂ , 47♀ , Feb.–Mar. 1958 (A. Chickering, MCZ 71393, PBI_OON 27241), 1♂ , 1♀ ; Gatún, Jan. 30, 1954 (A. Chickering, MCZ 71381, PBI_OON 27270), 2♀ , Feb. 15, 1958 (A. Chickering, MCZ 71383, PBI_OON 27271), 38♂ , 98♀ , Feb. 27, 1958 (A. Chickering, MCZ 71384, PBI_OON 27262), 17♂ , 56♀ , Mar. 6, 1958 (A. Chickering, MCZ 71382, PBI_OON 27272), 20♂ , 85♀ ; Pipeline Road, Gamboa, Aug. 7, 1983 , jungle, elev. 10 m (J., F. Murphy, AMNH PBI_OON 1548), 2♀ . Los Santos : Tonosí, Aug. 12, 2003 , fallen trunks (R. Mirando, A. Santos, MIUP PBI_OON 37745), 1♀ . Panamá : Arraijan, July 6, 1950 (A. Chickering, MCZ 71375, PBI_OON 27268), 1♀ ; Balboa, May 1964 (A. Chickering, MCZ 71391; PBI_OON 27244), 13♂ , 54♀ ; Barro Colorado Island, July 30, 1954 (A. Chickering, MCZ 71395, PBI_OON 27251), 1♀ , May 18, 1964 (A. Chickering, MCZ 71396, PBI_OON 27264), 1♀ ; Chilibre, July 27, 1939 (A. Chickering, MCZ 71374; PBI_OON 27247), 1♀ ; Chilibrillo Cave, near Chilibre, Feb. 7, 1959 , drift along stream in cave (H. Dybas, FMNH 33657, PBI_OON 10159), 1♀ ; Coccoli, Jan. 13, 1958 (A. Chickering, MCZ 71372, PBI_OON 27261), 4♀ ; Corozal, July 10, 1954 (A. Chickering, MCZ 71400, PBI_OON 27256), 1♂ , 9♀ , Dec. 23, 1957Jan. 4, 1958 (A. Chickering, MCZ 71398, PBI_OON 27260), 3♂ , 8♀ , May 25, 1964 (A. Chickering, MCZ 71399, PBI_ OON 27255), 5♂ , 9♀ ; Diablo, Dec. 19, 1957 (A. Chickering, MCZ 71373, PBI_OON 27267), 18♀ ; Forest Preserve, July 1939 (A. Chickering, MCZ 71390, PBI_OON 27257), 1♀ , July 23, 1950 (A. Chickering, MCZ 71389, PBI_OON 27258), 1♀ , July 28, 1954 (A. Chickering, MCZ 71388, PBI_OON 27243) 5♀ , Dec. 24, 1957 (A. Chickering, MCZ 71386, PBI_OON 27252), 3♀ , Feb. 14, 1958 (A. Chickering, MCZ 71387, PBI_OON 27240), 2♂ ; Madden Dam, Feb. 12, 1958 (A. Chickering, MCZ 71385, PBI_OON 27250), 1♀ ; 2 mi N Paraíso, Jan. 21, 1958 (A. Chickering, MCZ 68264, PBI_OON 27259), 1♂ , 1♀ ; Parque Metropolitano, near canopy crane, 8.99483°N , 79.54366°W , Dec. 30, 2007 , sifting rainforest litter (M. Draney et al., FMNH 34885, PBI_OON 10598), 1♀ ; Pedro Miguel, Aug. 1954 (A. Chickering, MCZ 71401, PBI_OON 27248), 11♂ , 36♀ , Dec. 26, 1957 (A. Chickering, MCZ 71397, PBI_OON 27263), 10♂ , 17♀ , Jan.–Mar. 1958 (A. Chickering, MCZ 71402, PBI_OON 27245), 3♂ , 17♀ ; Summit, July–Aug. 1950 (A. Chickering, MCZ 71380, PBI_OON 27253), 3♂ , 9♀ , Aug. 17, 1954 (A. Chickering, MCZ 71379, PBI_OON 27265), 2♂ , 13♀ ; Summit Gardens, July–Aug. 1954 (A. Chickering, MCZ 71377, PBI_OON 27242), 6♂ , 36♀ , Dec. 12, 1957 (A. Chickering, MCZ 71378, PBI_OON 27266), 6♂ , 20♀ , May 13, 1964 (A. Chickering, MCZ 71376, PBI_OON 27269), 3♂ . WEST INDIES: Cuba : Camagüey : Hoyo de Bonet, Sierra de Cubitas, Feb. 2009 (Y. Martínez, MNH PBI_OON 49444), 1♂ , 5♀ , Sept. 2009 (Y. Martínez, MNH PBI_OON 49445), 1♀ . Cienfuegos : Soledad, Aug. 1–11, 1934 (P. Darlington, MCZ 71320, PBI_ OON 27427), 1♀ . Guantánamo : Valle del Río Jojo, 20°07′35″N , 74°28′42″W , May 11, 2010 , litter (N. Platnick, A. Pérez, A. Sánchez, G. Alayón, AMNH PBI_OON 49448), 1♀ . Holguín : Bahía de Taco, Moa, Nov. 9, 2004 , under rock (A. Sánchez, BSC PBI_OON 49446), 1♀ . La Habana : Havana , June 12 (Baker, MCZ 66705, PBI_OON 27421), 1♀ . Santiago de Cuba : Arroyo Grovert, 10 km NE Caney, 1995, leaf and log litter, elev. 300 m (S. Peck, AMNH PBI_OON 49443), 1♀ ; Reserva Ecológica Siboney-Juticí, 19°57′39″N , 75°42′52″W , May 5, 2010 , dry litter, elev. 30 m (N. Platnick, A. Sánchez, A. Pérez, G. Alayón, AMNH PBI_OON 49447), 1♂ ; Río Bacanao tributary, Mata Yegua, 2 km NE El Escandel, 20°05′34″N , 75°41′49″W , May 10, 2010 (N. Platnick, A. Sánchez, A. Pérez, G. Alayón, AMNH PBI_OON 49449), 3♂ , 4♀ . Jamaica : Clarendon : 2.5 mi N May Pen, Nov. 10, 1963 (A. Chickering, MCZ 65630, PBI_OON 27296), 1♀ ; Salt River, Nov. 24, 1963 (A. Chickering, MCZ 68262, PBI_OON 27297), 3♀ . Kingston : Kingston , Aug. 27–29, 1934 (P. Darlington, MCZ 42624, PBI_OON 27282), 2♀ . Portland : Port Maria, Dec. 31, 1951 , under litter (C. Perraton, MCZ 66523, PBI_OON 27278), 1♀ . St. Andrew: no specific locality, May 5, 1956 (C. Hoff, MCZ 65094, PBI_OON 27289), 3♂ ; Hermitage Rd., Nov. 28, 1963 (A. Chickering, MCZ 71328, PBI_OON 27284), 1♂ , 3♀ ; Hope Gardens, Nov. 1963 (A. Chickering, MCZ 71330, PBI_OON 27279), 9♂ , 38♀ , Dec. 1963 (A. Chickering, MCZ 71331, 27293), 10♂ , 76♀ ; Liguanea, Oct.15, 1957 (A. Chickering, MCZ 71329, PBI_OON 27294), 1♀ , Dec. 1963 (A. Chickering, MCZ 71327, PBI_OON 27276), 2♂ , 4♀ ; Mona, Nov. 11, 1957 , pasture (A. Chickering, MCZ 71332, PBI_OON 27288), 1♀ , Nov. 1963 (A. Chickering, MCZ 71336, PBI_OON 27287), 4♂ , 6♀ , Dec. 1963 (A. Chickering, MCZ 71326, PBI_OON 27290), 1♀ ; Red Hills Road, Oct. 28, 1957 (A. Chickering, MCZ 71340, PBI_OON 27280), 1♀ ; Richards Reservoir, Nov. 1957 (A. Chickering, MCZ 71333, PBI_OON 27292), 1♀ , Dec. 18, 1963 (A. Chickering, MCZ 71335, PBI_OON 27283), 1♂ , 2♀ ; Stony Hill, Oct. 18, 1957 (A. Chickering, MCZ 71334, 71337, PBI_OON 27274, 27281), 2♂ , 1♀ . St. Ann: Ferry, Claremont, June 19, 1954 (A. Chickering, MCZ 65909, PBI_OON 27273), 1♀ . Saint Catherine : 3 mi. E Old Harbour, Oct. 21, 1957 (A. Chickering, MCZ 71138, PBI_OON 27295), 3♀ ; School of Agriculture, Nov. 23, 1957 (A. Chickering, MCZ 32461, PBI_OON 27291), 1♀ . St. Thomas: 6 mi NE Bath, Dec. 10, 1957 (A. Chickering, MCZ 66522, PBI_OON 27286), 1♀ ; Roselle Falls, Oct.–Nov. 1957 (A. Chickering, MCZ 71339, PBI_OON 27275), 1♂ , 6♀ . Puerto Rico : Fajardo : Isleta Marina, Isla Obispo, Nov. 7, 1964 (H. Heatwole, F. McKenzie, AMNH PBI_OON 1954), 1♂ . Guayama : Punta Pozuela, Dec. 25, 1985 (V., B. Roth, CAS 26297, PBI_OON 2587), 1♀ . Lajas : Laguna Cartagena, Valle de Lajas , Jan. 8, 1964 (A. Chickering, MCZ 71370, PBI_OON 27391), 2♂ , 3♀ . Loíza : 15 km E San Juan, Jan. 26, 1964 (A. Chickering, MCZ 71368, PBI_OON 27380), 1♀ . Maricao : El Vivero, Jan. 9, 1964 (A. Chickering, MCZ 71365, PBI_OON 27383), 1♂ ; Montañas de Uroyan, Rt. 108, km 12–22, Jan. 12, 1964 (A. Chickering, MCZ 71369, PBI_OON 27392), 3♂ , 2♀ . Mayagüez : 3 mi S Mayagüez on road to Hormigueros , Jan. 8, 1964 (A. Chickering, MCZ 71361, PBI_OON 27379), 1♂ , 14♀ ; 5 km E Mayagüez on Route 106, Jan. 1964 (A. Chickering, MCZ 71363, PBI_OON 27381), 5♂ , 22♀ ; woods near Nuclear Center, Jan. 1964 (A. Chickering, MCZ 71366, PBI_OON 27386), 4♂ , 19♀ ; University campus, Jan. 1964 (A. Chickering, MCZ 71359, PBI_OON 27384), 16♂ , 42♀ ; 5 km N University campus, Jan. 5, 1964 (A. Chickering, MCZ 71362, PBI_OON 27387), 1♂ , 2♀ ; University farm, E campus, Jan. 1964 (A. Chickering, MCZ 71360, PBI_OON 27385), 3♂ , 3♀ ; University farm, N campus, Jan. 1964 (A. Chickering, MCZ 71357, PBI_OON 27393), 3♂ , 11♀ , Feb. 1964 (A. Chickering, MCZ 71358, PBI_OON 27382), 7♂ , 27♀ , Feb. 2, 1964 (A. Chickering, MCZ 68265, PBI_OON 27389), 1♂ , 1♀ . Río Grande : El Yunque Biological Station, Jan. 25, 1964 , elev. 2100 ft (A. Chickering, MCZ 71364, PBI_ OON 27390), 1♂ , 2♀ . San Germán : Port Grilo, Feb. 23–Apr. 4, 1955 (A. Nadler, AMNH PBI_OON 1975), 1♀ . San Juan: Rio Piedras, Dos Pinos, Sept. 21, 1963 , under boards (E. Nelson, MCZ 71367, PBI_OON 27388), 1♀ ; between Toa Alta and Toa Baha, ca. 17.5 km from San Juan, forest ( AMNH PBI_OON 1955), 1♀ (details from Petrunkevitch, 1929 ). Virgin Islands : St. Croix: no specific locality, Sept. 1966 (A. Chickering, MCZ 71351, PBI_OON 27410), 13♂ , 16♀ ; Christiansted (Beatty, MCZ PBI_OON 49466), 1♂ ( holotype ); Frederiksted, Mar. 1964 (A. Chickering, MCZ 71341, PBI_OON 27425), 7♂ , 10♀ ; vicinity of King’s Hill, Mar. 1964 (A. Chickering, MCZ 71347, PBI_OON 27405), 1♂ , 9♀ ; Lavaetz Gardens, Mar. 24, 1964 (A. Chickering, MCZ 71349, PBI_OON 27416), 1♂ , 8♀ . St. John: no specific locality, Feb. 1964 (A. Chickering, MCZ 71348, PBI_OON 27407), 1♂ , 2♀ , Mar. 1964 (A. Chickering, MCZ 71350, PBI_ OON 27406), 1♀ , July 1966 (A. Chickering, MCZ 71346, PBI_OON 27413), 4♂ , 10♀ . St. Thomas: no specific locality, Feb. 1964 (A. Chickering, MCZ 71352, PBI_OON 27420), 4♂ , 2♀ , July 27, 1966 (A. Chickering, MCZ 71353, PBI_OON 27403), 1♂ , 18♀ , Aug. 1966 (A. Chickering, MCZ 71319, PBI_OON 27424), 26♂ , 52♀ ; High School grounds, Feb. 22, 1964 (A. Chickering, MCZ 71189, PBI_OON 27404), 4♀ , Mar. 10–11, 1964 (A. Chickering, MCZ 71354, PBI_OON 27402), 25♂ , 43♀ . Tortola: no specific locality, July 30–Aug. 5, 1966 (A. Chickering, MCZ 71190, PBI_OON 27414), 20♀ . Leeward Islands: St. Kitts: no specific locality, Sept.14–22, 1966 (A. Chickering, MCZ 68261, PBI_OON 27417), 1♀ . Windward Islands: Barbados: Turners Hall Woods, Feb. 23, 1979 , Berlese, termite nest (S. Peck, AMNH PBI_ OON 1236), 1♂ , same ( FMNH PBI_OON 49468), 1♂ . St. Lucia: no specific locality, Oct.5–13, 1966 (A. Chickering, MCZ 71316, PBI_OON 27412), 2♂ , 26♀ . St. Vincent: no specific locality ( MNHN 5728, PBI_OON 32322), 1♀ ( syntype ), Oct. 15–24, 1966 (A. Chickering, MCZ 71355, 71356, PBI_OON 27418, 27419), 4♂ , 15♀ . Richmond Estate, leeward open valley near sea level, webs under rotting banana flowers ( BMNH PBI_OON 32324), 4♀ ( syntypes ). Trinidad and Tobago : Trinidad: no specific locality (N. Weber, MCZ 71342, PBI_OON 27409), 2♂ . Arima : Simla, Arima Valley, Apr. 10–22, 1964 (A. Chickering, MCZ 71343, PBI_OON 27426), 10♂ , 27♀ , Apr. 23–27, 1964 (A. Chickering, MCZ 67071, PBI_OON 27422), 11♂ , 20♀ . St. Andrew: Salybia, Feb. 1972 (J. Cooke, AMNH PBI_OON 49450), 7♀ . St. George: Curepe, Nov. 21–Dec. 11, 1977 , pan trap (W. Mason, CNC PBI_OON 38142), 1♀ ; Curepe, Santa Margareta, Sept. 26–Oct. 26, 1974, yellow pan trap (N. Beg, AMNH PBI_OON 37072), 2♀ ; Saint Augustine, Nov. 16, 1943Feb. 23, 1944 (Strickland, AMNH PBI_OON 1956), 1♂ ; St. Augustine, University Campus, Apr. 1–6, 1964 (A. Chickering, MCZ 71344, PBI_OON 27408), 10♂ , 20♀ , Apr. 7–9, 1964 (A. Chickering, MCZ 71345, PBI_OON 27411), 14♂ , 39♀ . SOUTH AMERICA: Colombia : Amazonas: Leticia, June 18, 1965 , grass at night (P. Craig, J. Robb, CDU PBI_OON 35946), 2♀ . Cauca : Poblado, Parque Nacional Gorgona, July 5, 2003 , elev. 5 m (A. Rico et al., ICN 2510, PBI_OON 49469), 1♀ . Tolima : Espinal, Granja los Marañones, 4°09′10″N , 74°53′19″W , July 2004 , cotton field, elev. 320 m (L. Gomez, C. Perafan, ICN 4098, PBI_OON 49470), 1♀ . Venezuela : Portuguesa : Guanare, Sept. 10–13, 1957 (B. Malkin, CAS 26296, PBI_OON 2586), 1♀ . Sucre : 6 km SE Río Caribe, July 24, 1987 , soil and litter, gallery forest (S., J. Peck, AMNH PBI_OON 1957), 1♂ . Brazil : Pará: Bosque Rodrigues Alves , 1°25′49.0″S , 48°27′22.3″W (J. Barreiros, MPEG 10478, 10479, PBI_OON 40726, 40727), 2♀ ; Ilha de Cotijuba, Belém, 1°14′S , 48°35′W , Nov. 2004 ( MPEG 10271, PBI_OON 40724), 1♀ ; Mata, Várzea, Belém, Oct. 2005 (L. Macambira, MPEG 10709, PBI_OON 40692), 1♂ , 2♀ . Galapagos Islands : Pinta : no specific locality, July 18, 1977 , sweeping, beating, dry perennial grass, elev. 400 m (W. Reeder, TMM PBI_OON 49471), 2♀ . Santa Cruz : no specific locality, May 7, 1980 , turning rocks and downed wood, fern zone, elev. 150 m (W. Reeder, TMM 57847, PBI_OON 36930), 1♀ ; Bellavista, Feb. 3–8, 1989 , banana traps (S. Peck, Sinclair, KBIN PBI_OON 49504), 1♀ , Apr. 10–30, 1996 , pitfall trap, guava forest. elev. 350 m (S. Peck, KBIN PBI_OON 49497), 2♀ ; near Bellavista, Apr. 9, 1981 (Y. Lubin, MCZ 71323, PBI_OON 27429), 1♀ ; 2 km N Bellavista, May 14–July 13, 1985 , flight intercept trap, guava thicket, elev. 360 m (S., J. Peck, KBIN PBI_OON 49501, 49502, 49514), 5♂ , 12♀ ; Caseta Occidente, Feb. 11–Mar. 18, 1982 , elev. 170 m (L. Baert, J. Maelfait, KBIN PBI_OON 49484, 49488), 1♂ , 8♀ ; Caseta Tortuga, Feb. 19, 1988 , elev. 180 m (L. Baert, J. Maelfait, K. Desender, KBIN PBI_OON 49483), 1♂ ; El Carmen farm, Mar. 10–21, 1998 , elev. 350 m (L. Baert, J. Maelfait, K. Desender, KBIN PBI_OON 49503), 2♂ , 4♀ ; Hornemann farm, Bellavista, Apr. 3, 1975 , screening litter at base of bamboo, elev. 200 m (W. Reeder, TMM 59909, PBI_OON 36931), 2♂ , 3♀ ; Los Gemelos, Feb. 12–Apr. 2, 1986 –1988, scalesia, pampa, elev. 580–600 m (L. Baert, J. Maelfait, K. Desender, KBIN PBI_OON 49474. 49477, 49479–48481, 49490, 49498, 49512, 49515, 49519), 82♂ , 128♀ ; Media Luna , June 21–July 21, 1992 , elev. 600 m (J. Palacios, KBIN PBI_OON 49506), 1♂ , 3♀ , Apr. 10–30, 1996 , pitfall, miconia, elev. 575 m (S. Peck, KBIN 49499), 1♂ , 2♀ ; Media Luna trail, Mar. 10, 1982 , elev. 200–250 m (L. Baert, J. Maelfait, KBIN PBI_OON 49489), 2♀ ; Feb. 15–Apr. 4, 1986 –2000, transition zone, elev. 350 m (L. Baert, J. Maelfait, K. Desender, KBIN PBI_OON 49486, 49487, 49491, 49505, 49513), 7♂ , 8♀ , Feb. 15–Apr. 2, 1986 –1989, miconia, elev. 500–550 m (L. Baert, J. Maelfait, K. Desender, KBIN PBI_OON 49473, 49493, 49507, 49517), 15♂ , 30♀ ; northern transect, Feb. 12–Apr. 2, 1988 , elev. 400–560 m (L. Baert, J. Maelfait, K. Desender, KBIN PBI_OON 49478, 49510, 49516), 4♂ , 7♀ ; 5 km N Puerto Ayora, May 1–30, 1991 , malaise-flight intercept trap, elev. 110 m (S., J. Peck, KBIN PBI_OON 49496), 1♀ ; 4 km E Santa Rosa, Apr. 10–May 4, 1996, pitfall trap, elev. 350 m (S. Peck, KBIN PBI_OON 49500), 1♂ ; 10 km N Santa Rosa, Mar. 1–30, 1992, elev. 500 m (S. Peck, KBIN PBI_OON 49508), 1♀ ; southern transect, Feb. 15–Mar. 25, 1986 , elev. 140–500 m (L. Baert, J. Maelfait, K. Desender, KBIN PBI_OON 49475, 49476, 49482, 49485, 49492, 49494, 49495), 7♂ , 12♀ ; southern transect, E Santa Rosa, Mar. 30–Apr. 10, 1996, elev. 350 m (L. Baert, K. Desender, P. Verdijck, KBIN PBI_OON 49509), 1♀ ; southern transect, N Santa Rosa, Apr. 3, 2000, elev. 500 m (L. Baert, J. Maelfait, K. Desender KBIN PBI_OON 49511), 1♀ . OLD WORLD: Equatorial Guinea : Micomeseng , July 18–27, 1989 , pitfall , swamp with ferns near river ( M. Alderweireldt , MRAC 169997 , PBI_OON 29241), 1♂ . Gabon : Estuaire : Ntoum , July 1985 , forest litter ( A. Pauly , MRAC 220031 , PBI_OON 28807), 1♀ . Comoros : Anjouan : Domoni , road to Tatinga bridge, May 19, 2003 , litter, forest remnants ( R. Jocqué , D. Van den Spiegel , MRAC 213420 , PBI_OON 29284), 1♂ ; Mutsamudu , May 20, 2003 , mango litter, hospital garden ( R. Jocqué , D. Van den Spiegel , MRAC 213152 , PBI_OON 29281), 1♀ . Mayotte : Coconi , campus of DAF, July 18–Aug. 2, 1998 , Winkler , sifting litter ( R. Jocqué , MRAC 208204 , 208268 , 208316 , PBI_OON 28806, 29234, 29333), 4♀ . Mohéli : Hoani , May 23, 2003 , sifting litter on road ( R. Jocqué , D. Van den Spiegel , MRAC 213155 , PBI_OON 29285), 1♀ . Madagascar : Antsiranana: Lokobe Forest, Nosy Be , 13°24′58″S , 48°18′26″E , Aug. 11–14, 1992 (V., B. Roth , CAS 30882, PBI_OON 3371), 1♀ . Fianarantsoa : Ranomafana town , 21°14.9′S , 47°27.7′E , Apr. 30, 1998 , on tree bark ( C. Griswold et al., CAS 29564, PBI_OON 3369), 1♀ . Toamasina : Foulpointe, Sept. – Dec. 1993 ( A. Pauly , MRAC 177947 , 200227 , 200344 , 200380 , 220052 , 220054 , 220086 , PBI_OON 9768, 9779–9781, 9787, 9789, 9790), 5♂ , 5♀ , July–Sept. 1994 (A. Pauly , MRAC 220055–220057 , 220085 , PBI_ OON 9766, 9769. 9770, 9774), 1♂ , 3♀ ; Île Sainte-Marie , east beach ( R. Legendre , MNHN PBI_OON 36330), 1♀ . Toliara: Forêt Classee Tsikongambarika, Ivolo forest, Andily , ca. Fort Dauphin , 24°56′13.5″S , 46°55′58.4″E , Mar. 13, 2003 , Ludd tree trunks, litter, elev. 20 m ( D. Andriamalala et al., CAS 15863, PBI_OON 3370), 1♀ . Seychelles : no specific locality, Jan. 23–Feb. 5, 1977 (A. Rundle , AMNH PBI_OON 1556, 1558, 1559), 3♀ . Mahé : Baie Lazare , June 26, 1972 , border of beach ( P. Benoit , J. Van Mol , MRAC 143253 , 146758 , PBI_OON 49522, 49523), 2♂ , 2♀ ( holotype , allotype , paratypes ) ; Dugand , June 25, 1972 ( P. Benoit , J. Van Mol , MRAC 143182 , PBI_OON 49524), 1♂ . Praslin : Fond Ferdinand , July 24, 1972 , lodoicea forest ( P. Benoit , J. Van Mol , MRAC 143397 , PBI_OON 49525), 1♂ . Silhouette : Anse Lacars-Anse Paptes , Jan. 12, 1999 ( M. Saaristo , ZMUT 1233 , PBI_OON 16071), 2♂ , 2♀ . Mascarene Islands : La Réunion : St. Philippe , Mare-Longue Forest , May 27, 2009 ( M. Dierkens , CMD PBI_OON 49472), 1♂ . China : Hong Kong : New Territories , Mai Po , Feb. 27, 1988 , mangroves, elev. 5 m ( J., F. Murphy , AMNH PBI_OON 1549), 1♀ . Singapore : Kranji , July 7, 1992 , mangroves, elev. 1 m ( J., F. Murphy , AMNH PBI_ OON 31308), 2♀ ; University , Jan. 22, 1986 , water tanks, elev. 40 m ( J., F. Murphy , AMNH PBI_OON 31306), 2♀ . Caroline Islands : Palau Islands : Koror Island , Mar. 15, 1973 , compost near biology lab ( AMNH PBI_OON 38558), 1♀ , Apr. 3, 1973 , litter, taro patch ( AMNH PBI_OON 38601), 1♀ , Mar. 26–May 9, 1973 , litter, taro patch ( J. Beatty , J. Berry , AMNH PBI_OON 37430, 38614), 3♂ , 3♀ ; Malakal Island , Apr. 17, 1973 , around grass roots in field ( AMNH PBI_OON 38552), 2♀ . Hawaii : Hawaii: Iliahi St., Hilo, Feb. 1, 1995, litter ( J. Berry , AMNH PBI_OON 38603), 1♀ ; Isaac Hale Beach Park , Puna District , Feb. 24, 1995 , pandanus litter ( J., E. Berry , AMNH PBI_OON 38374), 1♀ ; 1 mi W Mackenzie State Park , Route 137, Puna District , Jan. 31, 1997 , casuarina litter ( J., E. Berry , AMNH PBI_OON 37438), 1♂ , 3♀ ; Naalehu , Route 11, mile marker 61, Kau District , Feb. 7, 1997 , litter along driveway ( J. Berry , AMNH PBI_OON 38542), 1♂ , 2♀ , same, mile marker 66, Feb. 8, 1997 , forest litter on hill, elev. 1000–1100 ft (J., E. Berry , AMNH PBI_OON 38373, 38378), 1♂ , 5♀ . Kauai : Haena State Park , Jan. 15, 1998 , litter, almond/ ficus forest ( J., E. Berry , AMNH PBI_OON 37439), 4♂ , 3♀ ; Kahili Mountain , 5 mi N Koloa , Jan. 11, 1998 , mixed forest, elev. 920 ft ( J., E. Berry , AMNH PBI_OON 38375), 1♂ ; Makaha Ridge Road , Jan. 23, 1998 , pine forest litter, elev. 2000 ft ( J., E. Berry , AMNH PBI_OON 38376), 2♀ ; Moloaa Beach , Kuaehu Point , Jan. 11, 1988 ( J. Berry , AMNH PBI_OON 38543), 1♀ ; National Tropical Botanical Garden , Lawai , Jan. 21, 1998 , litter ( J. Berry , AMNH PBI_OON 38616), 3♀ , Feb. 4, 1998 , litter on bank beside stream at waterfall and pool (J. Berry , AMNH PBI_OON 37763), 1♀ . Oahu : Wahiawa ( H. Hagan , ex MCZ 71324, PBI_OON 96), 1♀ . DISTRIBUTION: Pantropical, and also introduced in north temperate buildings; sometimes sympatric with I. velox . However, not all the published records of the species are accurate. For example, the specimens from St. Helena described by Benoit (1977: 41 , figs. 15a–c), including his invalid “ allotype ,” belong neither to I. peltifer nor I. velox .