Revision of the subgenus Phortica (sensu stricto) (Diptera, Drosophilidae) from East Asia, with assessment of species delimitation using DNA barcodes Author Huang, Jia Author Gong, Lu Author Tsaur, Shun-Chern Author Zhu, Lin Author An, Keying Author Chen, Hongwei text Zootaxa 2019 2019-09-30 4678 1 1 75 journal article 25341 10.11646/zootaxa.4678.1.1 96c8a18a-e887-4345-b266-d76076e8416b 1175-5326 3468782 91B46647-B441-41F5-BE27-3A67D699788C Phortica ( Phortica ) setitabula Chen & Gao, 2005 Phortica ( Phortica ) setitabula Chen & Gao in Chen et al . 2005: 3970 . Diagnosis. Paramere with 1 acute process subapically; branch of aedeagal basal bridge basally with two slender processes bearing a few triangular projections laterally (modified from Chen et al . 2005 ). Specimens examined. 1♂ ( KIZ ), Wuliangshan , Jingdong , Yunnan , 24°17'27''N , 100°19'48''E , alt. 1700–2100m , 5.viii.2006 , J.J. Gao; 9♂ ( KIZ ), Menglun South 5km , Mengla , Yunnan , 21°59'57''N , 101°02'50''E , alt. 820–890m , 12.iv.2010 , J.J. Gao, M.J. Toda; 19♂ ( SCAU ), Hesong , Menghai , Yunnan , 21°50'17''N , 100°05'48''E , alt. 1700– 1857m , 3♂ on 16.iv.2010 , J.J. Gao, 8♂ on 8.iv.2011 , J.M. Lu, Z.F. Shao, Y.R. Su, S.J. Yan, 8♂ on 12.v.2012 , H.W. Chen; 21♂ ( SCAU ), Muyiji Park , Ximeng , Yunnan , 22°37'15''N , 99°35'42''E , alt. 1200m , 2,4. iv.2011 , J.M. Lu, Z.F. Shao, Y.R. Su, S.J. Yan; 2♂ ( SCAU ), Yixiang , Pu'er , Yunnan , 22°44'19''N , 101°07'01''E , alt. 1400m , 13.v.2012 , H.W. Chen, J.J. Gao; 4♂ ( SCAU ), Baihualing , Baoshan , Yunnan , 25°17'50''N , 98°48'12''E , alt. 1400m , 26.viii.2013 , Q.S. Gao, K. Liu; 4♂ ( SCAU ), Mengdong , Cangyuan , Yunnan , 23°10'08''N , 99°13'52''E , alt. 1260–1340m , 10.v.2016 , J. Huang , Y.Q. Liu , Y.L. Wang , L. Zhu ; 1♂ ( SCAU ), Wulaoshan , Lincang , Yunnan , 23°15'17''N , 100°11'38''E , alt. 2155m , 14.v.2016 , J. Huang; 2♂ ( SCAU ), Tongbiguan , Yingjiang , Yunnan , 24°37'04''N , 97°40'03''E , alt. 1313m , 19.v.2016 , J. Huang, Y.Q. Liu; 3♂ ( SCAU ), Langping , Tianlin , Guangxi , 24°27'43''N , 106°22'52''E , alt. 1490m , 30.vii.2016 , H.W. Chen, L. Gong, Y.Q. Liu; 1♂ ( SCAU ), Mengyuan , Mengla , Yunnan , 21°47'29''N , 101°22'59''E , alt. 990m , 7.viii.2016 , L. Gong; 2♂ ( SCAU ), Nanling ( Shangsongshan ), Lancang , Yunnan , 22°47'57''N , 99°55'11''E , alt. 1807m , 12.viii.2016 , H.W. Chen, L. Gong; 2♂ ( SCAU ), Banpo , Lüchun , Yunnan , 22°34'50''N , 102°34'50''E , alt. 560m , 28.x.2016 , H.W. Chen; 2♂ ( SCAU ), Shangnongkang , Gengma , Yunnan , 23°32'26''N , 99°21'21''E , alt. 1315m , 1.xi.2017 , L. Gong, B.X. Li; 3♂ ( SCAU ), Zhongshan , Mangshi , Yunnan , 24°07'49''N , 98°31'27''E , alt. 1486m , 3.xi.2017 , H.W. Chen, L. Gong, B.X. Li; 2♂ ( SCAU ), Bangdong , Ruili , Yunnan , 24°06'09''N , 97°50'55''E , alt. 1370m , 6.xi.2017 , H.W. Chen, L. Gong; 2♂ ( SCAU ), Bangda , Mangshi , Yunnan , 24°07'55''N , 98°13'42''E , alt. 1710m , 8.xi.2017 , L. Gong, B.X. Li; 3♂ ( SCAU ), Mucheng , Longling , Yunnan , 24°08'48''N , 98°44'48''E , alt. 1030m , 9.xi.2017 , H.W. Chen, L. Gong, B.X. Li. Distribution. China ( Guangxi * , Yunnan ). FIGURE 29. Phortica ( Phortica ) saeta ( Zhang & Gan, 1986 ) from Deqin, Yunnan, China, male. A. Epandrium, surstylus, and cercus (lateral view); B. Surstylus (frontal view); C. Hypandrium, paramere, aedeagus, gonopod, and aedeagal apodeme (lateral view). Scale bars = 0.1 mm. Phortica ( Phortica ) subradiata ( Okada, 1977 ) ( Figs. 6C, 6D , 30 ) Amiota ( Phortica ) subradiata Okada, 1977: 24 . Amiota ( Phortica ) orientalis : Okada, 1971: 91 (nec Hendel, 1914 , misidentified). Diagnosis. Vertical process of gonopod apically crispate, with 2 or 3 projections per side ( Fig. 30C ); two basal bridges of aedeagal median rod distinctly separated from each other ( Fig. 30C ). FIGURE 30. Phortica ( Phortica ) subradiata ( Okada, 1977 ) from Chiayi, Taiwan, China, male. A. Epandrium and cercus (lateral view); B. Surstylus (frontal view); C. Hypandrium, paramere, aedeagus, gonopod, and aedeagal apodeme (lateral view). Scale bars = 0.1 mm. Specimens examined. CHINA : 26♂ ( SCAU ), Jianfengling , Ledong , Hainan , 18°41'50''N , 108°50'04''E , alt. 700–890m , 3♂ on 20.v.2004 , H.W. Chen, 23♂ on 20.iv.2008 , X.P. Chen, J.J. Gao; 2♂ ( SCAU ), Bapen , Fush- ui, Guangxi , 22°32'36''N , 107°53'18''E , alt. 210m , 16.viii.2004 , H.W. Chen; 8♂ , Menglun , Mengla , Yunnan , 21°54'57''N , 101°16'23''E , alt. 580m , 5♂ ( KIZ ) on 15.iii.2005 , Y.F. Li , 3♂ ( SCAU ) on 27.ix.2011 , H.W. Chen ; 6♂ ( SCAU ), Conghua , Guangdong , 23°43'30''N , 113°48'20''E , alt. 350m , 12.x.2005 , H.W. Chen; 1♂ ( TMNS ), Beibeng , Motuo , Xizang , 29°19'00''N , 95°20'00''E , alt. 900m , 29.ix.2010 , J.J. Gao; 1♂ ( SCAU ), Bangda , Chiayi , Taiwan , 23°28'47''N , 120°39'11''E , alt. 820m , 15.x.2012 , H.W. Chen; 23♂ ( SCAU ), Jinshan , Menglian , Yunnan , 22°19'56''N , 99°34'51''E , alt. 940m , 3♂ on 17.iv.2016 , J. Huang, L. Zhu, 20♂ on 23–25.x.2017 , H.W. Chen, L. Gong, B.X. Li; 8♂ ( SCAU ), Guanlei , Mengla , Yunnan , 21°38'39''N , 101°09'52''E , alt. 560–740m , 22.iv.2016 , J. Huang, Y.Q. Liu; 8♂ ( SCAU ), Muyiji Park , Ximeng , Yunnan , 22°37'15''N , 99°35'42''E , alt. 1200m , 5♂ on 30.iv.2016 , J. Huang , Y.Q. Liu , Y.L. Wang , L. Zhu , 3♂ on 11.viii.2016 , H.W. Chen, L. Gong, Y.Q. Liu; 2♂ ( SCAU ), Likan , Xi- meng, Yunnan , 22°39'21''N , 99°36'28''E , alt. 840m , 1.v.2016 , J. Huang; 1♂ ( SCAU ), Wulaoshan , Lincang , Yunnan , 23°15'17''N , 100°11'38''E , alt. 2155m , 14.v.2016 , J. Huang; 5♂ ( SCAU ), Nabang , Yingjiang , Yunnan , 24°39'51''N , 97°35'31''E , alt. 700m , 17.v.2016 , 18,19. viii.2016 , H.W. Chen, J. Huang, L. Gong; 30♂ , Botanic Garden , Ruili , Yunnan , 24°01'12''N , 97°51'33''E , alt. 1170m , 2♂ ( SCAU ) on 23.v.2016 , J. Huang , 25♂ ( SCAU ) on 22.viii.2016 , H.W. Chen , L. Gong , Y.Q. Liu , 3♂ ( KIZ ) on 6.xi.2017 , H.W. Chen , L. Gong , B.X. Li ; 7♂ ( SCAU ), Qimaba , Lü- chun, Yunnan , 22°49'08''N , 102°17'25''E , alt. 1280m , 3.viii.2016 , H.W. Chen, L. Gong, Y.Q. Liu; 10♂ ( SCAU ), Mengyuan , Mengla , Yunnan , 21°47'29''N , 101°22'59''E , alt. 990m , 7.viii.2016 , 16.x.2017 , H.W. Chen, L. Gong, Y.Q. Liu; 3♂ ( SCAU ), Nanling ( Shangsongshan ), Lancang , Yunnan , 22°47'57''N , 99°55'11''E , alt. 1807m , 2♂ on 17.viii.2017 , H.W. Chen, 1♂ on 23.x.2017 , L. Gong; 12♂ ( SCAU ), Longtan Park , Ximeng , Yunnan , 22°38'44''N , 99°35'39''E , alt. 1170m , 26,27. x.2017 , H.W. Chen, L. Gong, B.X. Li; 6♂ ( SCAU ), Zhongshan , Mangshi , Yunnan , 24°07'49''N , 98°31'27''E , alt. 850m , 8.xi.2017 , H.W. Chen, L. Gong, B.X. Li; 3♂ ( KIZ ), Bangda , Mangshi , Yunnan , 24°07'55''N , 98°13'42''E , alt. 1710m , 8.xi.2017 , H.W. Chen, L. Gong, B.X. Li; 2♂ ( SCAU ), Mucheng , Longling , Yunnan , 24°08'48''N , 98°44'48''E , alt. 900m , 10.xi.2017 , L. Gong, B.X. Li; 9♂ ( SCAU ), Mengma , Menglian , Yun- nan, 22°13'44''N , 99°22'37''E , alt. 1140m , 15.iv.2018 , H.W. Chen, L. Gong, Y. Lin. Distribution . China ( Taiwan , Guangdong, Hainan, Guangxi, Yunnan, Xizang * ).