Annotated zoogeography of non-marine Tardigrada. Part II: South America
Author
Kaczmarek, Łukasz
Author
Michalczyk, Łukasz
Author
Mcinnes, Sandra J.
text
Zootaxa
2015
3923
1
1
107
journal article
41991
10.11646/zootaxa.3923.1.1
2822ae22-53bb-45c1-b3a3-49f7b6c0fb53
1175-5326
241936
2305A96C-0A03-4524-93AA-90359893A4DD
210.
Paramacrobiotus richtersi
(
Murray, 1911
)
sensu
lato [T]
M. richtersii
J. Murr. (
Marcus 1939
)
Macrobiotus richtersii
J. Murr., 1908
(de
Barros 1942b
)
M. richtersi
J. Murray, 1911
(du
Bois-Reymond Marcus 1944
)
Macrobiotus richtersii
J. Murr. (
Iharos 1963
)
Macrobiotus richtersi
Murr. (
Iharos 1969
)
Macrobiotus richtersi
Murray, 1911
(
Rossi & Claps 1980
)
M. richtersi
Murray, 1911
(
Claps & Rossi 1981
, 1984, 1988)
Macrobiotus richtersi
J. Murray, 1911
(
Iharos 1982
)
Macrobiotus richtersi
Murray, 1911
(
Maucci 1988
,
Rossi & Claps 1991
,
Jerez & Narváez 2001
)
M. richtersi
J. Murr., 1911
(
Rossi & Claps 1989
)
M. richtersi
Murray (
Claps & Rossi 1997
)
Macrobiotus harmsworthi
/
richtersi
(
Montoya
et al.
2010
)
Terra typica
:
Ireland
(Europe)
Argentina
:
•
23°08′S
,
64°19′W
;
350 m
asl
: Salta Province,
Orán
[San Ramón de la Nueva
Orán
], mosses on rock.
Claps & Rossi (1984)
•
23°36′S
,
65°04′W
;
2,000 m
asl
: Jujuy Province, Departamento Santa Bárbara, lichens on vina lito.
Claps & Rossi (1984)
•
24°07′S
,
65°24′W
;
1,450 m
asl
: Jujuy Province, road to Yala, mosses from the edge of irrigation ditch and on tree (2 samples).
Claps & Rossi (1984)
•
24°18′S
,
64°54′W
;
700 m
asl
: Jujuy Province, road to Higueritas, mosses on soil.
Claps & Rossi (1984)
•
25°43′S
,
54°25′W
;
250 m
asl
: Misiones Province, Iguazú National Park, moss on rock and mosses on trees in the forest (3 samples).
Claps & Rossi (1988)
•
25°58′S
,
54°34′W
;
200 m
asl
: Misiones Province, Wanda, mosses on rocks.
Claps & Rossi (1988)
•
26°08′S
,
54°35′W
;
150 m
asl
: Misiones Province, Arroyo Torocuá,
12 km
S Puerto Esperanza, mosses on rocks (2 samples).
Claps & Rossi (1988)
•
26°47′S
,
65°20′W
;
750 m
asl
: Tucumán Province, Horco Molle, mosses on soil and tree (2 samples).
Claps & Rossi (1984)
•
26°51′S
,
65°26′W
;
700 m
asl
: Tucumán Province, road from San Miguel de Tucumán to Tafi del Valle, mosses and lichens on rocks (2 samples).
Claps & Rossi (1984)
•
26°56′S
,
55°04′W
;
200 m
asl
: Misiones Province, Capioví, mosses on trees in the forest (2 samples).
Claps & Rossi (1988)
•
27°27′S
,
58°39′W
;
50 m
asl
: Corrientes Province, old road to Santa Ana,
Selaginella
sp. in lagoon and moss on tree (2 samples).
Claps & Rossi (1988)
•
35°27′S
,
58°48′W
; 0 m asl
: Buenos Aires Province, Laguna San Miguel del Monte, on plant (
Ceratophyllum
sp.)
Rossi & Claps (1991)
.
•
36°22′S
,
56°43′W
; 0 m asl
: Buenos Aires Province, camping San Clemente del Tuy, lichens on tree (
Eucalyptus
) in shade.
Rossi & Claps (1980)
•
36°24′S
,
56°57′W
; 0 m asl
: Buenos Aires Province, Port General Lavalle, lichens on tree (
Tamarisk
) in full sun.
Rossi & Claps (1980)
•
40°50′S
,
71°38′W
;
900 m
asl
: Neuquén Province, Peninsula de Quetrihue, semi-wet, mosses on rocks on the sunny slope.
Claps & Rossi (1981)
•
41°00′S
,
71°30′W
;
800 m
asl
: Neuquén Province,
Isla
Victoria, mosses on tree.
Rossi & Claps (1989)
•
41°13′S
,
71°30′W
;
2,250 m
asl
: Rio Negro Province, Nahuel Huapi National Park, Cerro Catedral, semi-wet, lichens and mosses on trees and roots (
Nothofagus
sp.) in the forest, very wet, mosses on soil and dead wood and mosses on leaf litter in the forest (6 samples).
Claps & Rossi (1981)
•
41°58′S
,
71°29′W
;
1,200 m
asl
: Rio Negro Province, El Bolsón, Piltriquitron Mt., mosses on dry soil under
Mulinum spinosum
and between
Berberis
shrubs.
Iharos (1963)
•
41°58′S
,
71°28′W
;
1,460 m
asl
: Rio Negro Province, El Bolsón, Piltriquitron Mt., mosses on rocks and barks of trees near edge within
Nothofagus pumilio
forest.
Iharos (1963)
•
41°59′S
,
71°34′W
;
300 m
asl
: Rio Negro Province, El Bolsón, Valley of Rio Azul, mosses on tree (
Myrceugenia exupca
) in the
Myrceugenia exupca
-
Nothofagus dombeyi
marsh forest.
Iharos (1963)
•
41°59′S
,
71°31′W
;
360 m
asl
: Rio Negro Province, El Bolsón, foot of Piltriquitron Mt., mosses on rocks in shaded
Libocedrus chilensis
-Lomatia
obliqua
forest.
Iharos (1963)
•
41°59′S
,
71°31′W
;
370 m
asl
: Rio Negro Province, El Bolsón, foot of Piltriquitron Mt., slope above Pampa Azcona, mosses on soil of shrubby area.
Iharos (1963)
•
42°13′S
,
71°42′W
;
220 m
asl
: Chubut Province, El Turbio, Puelo Lake, mosses on wet soil in primeval
Nothofagus dombeyi
-
Libocedrus chilensis
forest near the lake.
Iharos (1963)
•
54°39′S
,
68°30′W
;
800 m
asl
: Tierra del Fuego Province, National Park Ushuaia [Tierra del Fuego National Park], soil and litter (2 samples).
Iharos (1982)
Bolivia
:
•
10°49′S
,
65°22′W
;
150 m
asl
: Beni Department, Guayaramerín, Estancia Esperanza, gallerian forest along Mamore river, leaf litter, leaf litter near water, decayed leaves on banana plantation, dry leaf litter, leaf litter from cacao plantation and soil from
Leguminosae
plantation (7 samples).
Iharos (1969)
•
10°56′S
,
65°28′W
;
150 m
asl
: Beni Department, Guayaramerín,
15 km
on W on the city, leaf litter in evergreen forest, near the road to Riberalta, leaf litter (2 samples).
Iharos (1969)
•
15°31′S
,
67°33′W
;
800 m
asl
: La Paz Department, between Alcoche and Puerto Linares,
20 km
on Alcoche, mosses in shaded place.
Iharos (1969)
•
16°08′S
,
67°43′W
;
1,600 m
asl
: La Paz Department, Coroico,
5 km
on N of the city, wet leaf litter from citrus and coffee plantation.
Iharos (1969)
•
16°11′S
,
69°05′W
;
4,100 m
asl
: La Paz Department, Copacabana [on the coast of Titicaca Lake], leaf litter from
Stipa
vegetation.
Iharos (1969)
•
16°16′S
,
67°53′W
;
3,100 m
asl
: La Paz Department,
5 km
on N of Unduavi, leaf litter and soil from
Vaccinum
plantation.
Iharos (1969)
•
16°19′S
,
67°54′W
;
4,000 m
asl
: La Paz Department, near Unduavi, near the river, very wet leaf litter under shrubs.
Iharos (1969)
Brazil
:
•
03°17′S
,
59°53′W
; 0 m asl
: Amazonas State, Manaus,
20 km
on S of the city, leaf litter from evergreen forest.
Iharos (1969)
•
21°49′S
,
49°05′W
: Undefined localities in São Paulo State, few localities.
Marcus (1939)
, de
Barros (1942b)
•
22°44′S
,
45°35′W
;
1,650 m
asl
: São Paulo State, Campos do Jordão, mosses or aquatic plants.
du
Bois-Reymond Marcus (1944)
•
22°45′S
,
47°24′W
;
550 m
asl
: São Paulo State, Município de Sta. Bárbara [Município Santa Bárbara d ′Oeste], mosses or aquatic plants.
du
Bois-Reymond Marcus (1944)
•
22°54′S
,
47°04′W
;
700 m
asl
: São Paulo State, Município Campinas, in bromeliads.
du
Bois-Reymond Marcus (1944)
•
25°26′S
,
49°16′W
;
900 m
asl
: Paraná State, Curitiba, mosses or aquatic plants.
du
Bois-Reymond Marcus (1944)
Chile
:
•
18°11′S
,
69°13′W
;
4,800 m
asl
[
4,550 m
asl]: Region XV Arica y Parinacota, Laguna La Cotacotani, leaf litter.
Iharos (1969)
•
18°34′S
,
70°05′W
;
500 m
asl
: Region XV Arica y Parinacota, Azapa Chico,
25 km
. on East of Arica, near the river Rio Lauca, wet soil on bamboo plantation.
Iharos (1969)
•
20°12′S
,
69°17′W
: Undefined locality in Region I Tarapac, Bofedal de Bajuco, soil under red cactus.
Iharos (1969)
•
30°17′S
,
71°16′W
;
400 m
asl
: Region IV Coquimbo, Las Cardas, leaf litter under shrubs.
Iharos (1969)
•
30°40′S
,
71°41′W
;
900 m
asl
[
650 m
asl]: Region IV Coquimbo, Fray Jorge [Bosque de Fray Jorge National Park], foggy forest, leaf litter.
Iharos (1969)
•
31°55′S
,
71°31′W
; 0 m asl
: Region IV Coquimbo, Los Vilos, jungle soil and leaf litter.
Iharos (1969)
•
32°58′S
,
71°01′W
;
1,900 m
asl
: Region V Valparaíso (Región de Valparaíso), Cordiliera de la Costa, Cerro El Roble,
84 km
NW of Santiago de
Chile
, leaf litter under snow, under the bushes.
Iharos (1969)
•
32°58′S
,
71°01′W
;
1,700 m
asl
: Region V Valparaíso (Región de Valparaíso), Cordiliera de la Costa, Cerro El Roble,
84 km
NW of Santiago de
Chile
, leaf litter between dry bushes.
Iharos (1969)
•
33°04′S
,
71°00′W
;
1,000 m
asl
: Region RM Metropolitana (Región Metropolitana de Santiago), Tiltil, Cuesta La Dormida, jungle leaf litter.
Iharos (1969)
•
33°21′S
,
70°19′W
;
2,300 m
asl
: Region RM Metropolitana (Región Metropolitana de Santiago), Farellones,
30 km
on East of Santiago de
Chile
, detritus and soil under the bushes, wet mosses from rocky cave and leaf litter (3 samples).
Iharos (1969)
•
33°26′S
,
70°38′W
;
800 m
asl
: Region RM Metropolitana (Región Metropolitana de Santiago), Santiago de
Chile
,
San Cristóbal
Mt., mosses on rocks.
Iharos (1969)
•
33°30′S
,
70°55′W
;
850 m
asl
: Region RM Metropolitana (Región Metropolitana de Santiago), Quebrada de La Plata, shrubs upland, moss and lichen on rocks (2 samples).
Iharos (1969)
•
37°13′S
,
72°23′W
;
100 m
asl
: Region VIII Biobío, Salto del Laja, near waterfalls, mosses on rocky wall.
Iharos (1969)
•
50°59′S
,
73°00′W
;
1,200 m
asl
: Region XII Magallanes (Región de Magallanes y de la
Antártica
Chilena), Torres del Paine National Park, mosses on trees and rocks, nearly all in half sunlight.
Maucci (1988)
•
51°34′S
,
72°36′W
;
200 m
asl
: Region XII Magallanes (Región de Magallanes y de la
Antártica
Chilena), Ultima Esperanza, near Cueva del Milodón, mosses on rocks, in sunlight.
Maucci (1988)
Colombia
:
•
07°07′N
,
73°02′W
;
1,795–1,970 m
asl
: Santander Department, El Diviso Natural Reserve, Pine forest (
Pinus patula
), mosses (
Hypopterygium tamariscinum
,
Fabronia ciliaris
and
Sematophyllum insularum
).
Jerez & Narváez (2001)
Uruguay
:
•
30°15′S
,
57°36′W
;
50 m
asl
: Artigas Department, Bella Unión, mosses or lichens on rocks, trees or posts or aquatic macrophytes.
Claps & Rossi (1997)
•
31°03′S
,
57°38′W
;
50 m
asl
: Salto Department, Arroyo Palomas, mosses or lichens on rocks, trees or posts or aquatic macrophytes.
Claps & Rossi (1997)
•
31°20′S
,
57°51′W
;
50 m
asl
: Salto Department, Arroyo San Antônio de Chico, mosses or lichens on rocks, trees or posts or aquatic macrophytes.
Claps & Rossi (1997)
Venezuela
:
•
08°46′N
,
70°49′W
;
4,300 m
asl
: Region Los Andes, Mérida, Mucuñuque Mt., moss.
Montoya
et al.
(2010)
•
08°45′N
,
70°49′W
;
4,450 m
asl
: Region Los Andes, Mérida, Mucuñuque Mt., surface samples of peat bog.
Montoya
et al.
(2010)
Record numbers:
Argentina
: 24,
Bolivia
: 7,
Brazil
: 6,
Chile
: 15,
Colombia
: 1,
Uruguay
: 3,
Venezuela
: 2;
total: 58.
Remarks:
Paramacrobiotus richtersi
was, until recently, believed to be cosmopolitan (
McInnes 1994a
). New taxonomic data has shown that this is the nominal species for a species complex – the ‘
richtersi
group’ – requiring careful examination of both adults and eggs. Several newly described species of this group have been added to the faunas of South and Central
America
(
e.g.
Michalczyk & Kaczmarek 2006b, Michalczyk
et al.
2006, Pilato
et al.
2006). In the light of these reports, and given
P. richtersi
was originally described from Europe, we suggest all species records from South
America
should be treated with caution until the original slides are re-examined or the species is otherwise verified from the region.