The spider genus Patrera Simon (Araneae: Dionycha, Anyphaeninae) in the Atlantic Forest biome, with the description of one new species from Brazil Author Oliveira, Luiz Fernando M. Programa de Pós-Graduação em Zoologia, Instituto de Biociências de São Paulo, Rua do Matão, 14, 05508 - 090, Universidade de São Paulo, São Paulo, SP, Brazil. & Laboratório de Coleções Zoológicas, Instituto Butantan, Av. Vital Brasil, 1500, 05503 - 900, São Paulo, SP, Brazil. Author Brescovit, Antonio D. 0000-0002-1511-5324 Laboratório de Coleções Zoológicas, Instituto Butantan, Av. Vital Brasil, 1500, 05503 - 900, São Paulo, SP, Brazil. & antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 0002 - 1511 - 5324 antonio.brescovit@butantan.gov.br text Zootaxa 2021 2021-06-24 4991 2 201 246 journal article 5537 10.11646/zootaxa.4991.2.1 f9dde51c-a2b9-4f3e-abe5-a7a3d7b0002d 1175-5326 5028185 62BE9E59-1DAB-425F-A322-4E48F12391C6 Patrera procera ( Keyserling, 1891 ) Figs 1A–G ; 2A–D ; 3A–F ; 27C Anyphaena procera Keyserling, 1891: 86 , figs 49–49a ( Syntypes male and female from Rio de Janeiro ( 22°54’10”S ; 43°12’28”W ), Rio de Janeiro , Brazil , 7.I.1890 , Keyserling leg., deposited in BMNH 1890.7.1.613-614, examined. Lectotype male and paralectotype female here designated). Teudis procerus : Petrunkevitch, 1911: 518 . Aysha cinereovittata Mello-Leitão, 1945: 242 ( partim , only the females MLP; males = Iguarima censoria (Keyserling) , see Brescovit, 1997: 50 . Patrera procera : Brescovit, 1997: 33 , 50. Other material examined. BRAZIL . No date, 1 ♂ (MNHN 18666) ; Paraíba : João Pessoa , Mata do Buraquinho ( 7°8′17″S ; 34°50′42″W ), 1♀ , 2003–2004, S.C. Dias leg. (IBSP 99968) ; Areia , Reserva da Mata do Pau Ferro ( 06º57’S ; 35º44’W ), 1♂ , 23–29.IX.1999 , A.D. Brescovit et al. leg. (IBSP 138929); 2 ♂ (IBSP 138939); 1♂ (IBSP 138949); 1♂ (IBSP 138950); 1♂ (IBSP 138958); 1♂ (IBSP 138960); all collected in 23–29.IX.1999 by A.D. Bresco- vit et al . ; Bahia : 2♂ (MNHN 7135); Una , Reserva Biológica do Una ( 15°10′23″S ; 39°7′56″W ), 1♀ , 15–28.XI.2000 , A.D. Brescovit et al. leg. (IBSP 45700); Jussari ( 15°11’27”S ; 39°29’42”W ), 3♀ , 1970, Cepec / Ceplac leg. (MNRJ 3143) ; 1♂ , 1♀ (MNRJ 3236); 1♂ (MNRJ 3194); 2♂ (MNRJ 3256); 1♂ , 2♀ (MNRJ 13313); 2♂ (MNRJ 3183); Coaraci ( 14°38′27″S ; 39°33′3″W ), Fazenda Boa Esperança, 1♂ , 01.X.1970 , Cepec/Ceplac leg. (MNRJ 3231); Mato Grosso do Sul : Bonito , Baia Bonita ( 21º09’S ; 56º26’W ), 1♂ (IBSP 138887); 1♀ (IBSP 138890); 3♂ , 2♀ (IBSP 138902); all collected in 14–23.X.2002 , Equipe Biota ; Minas Gerais : Belo Horizonte , Estação Ecológica UFMG, Malaise trap ( 19°52′19″S 43°57′58″W ), 2♂ , 09–16.XI.1999 , J.C.R. Fontenelle leg. (IBSP 41119) ; Espírito Santo : São José do Calçado , Rio Itabapoana , Usina Hidrelétrica de Rosal ( 20°57’7”S ; 41°43’5”W ), 1♂ , 1♀ , XI.1999 , I. Knysak leg. (IBSP 26383) ; Apiacá ( 21°9′14″S ; 41°34′4″W ), 1♂ , XI.1986 , R.L.C. Baptista leg. (MNRJ 3188) ; Ca- riacica, Alto do Alegre , Reserva Estadual de Duas Caras ( 20°17’29.7”S ; 40°31’10.2”W ), 1♂ , 13.X.2005 , T . Souza et al. leg. (IBSP 133182); Linhares , Flona de Goytacazes ( 19°23’48”S ; 40°3’42”W ), 42m alt., 2♂ , 21–23.X.2005 , T . Souza et al. leg. (IBSP 134825); Conceição da Barra , Floresta Nacional do Rio Preto , Trilha do Mata-Pau , 60m alt. ( 18°24’00”S ; 39°50’00”W ), 1♂ , 19.X.2005 , T . Souza et al. leg. (IBSP 135120); Rio de Janeiro : Volta Redonda , Floresta da Cicuta ( 22°32′53″S ; 44°5′34″W ), 3♀ , VIII.2002 , E.L.C. Silva et al. leg. (IBSP 48942; IBSP 48950; IBSP 48954); Cachoeiras de Macacu , Reserva Ecológica de Guapiaçu ( 22°35′14.61″S ; 42°56′24.69″W ), 11♂ (IBSP 138968; IBSP 138970; IBSP 138972; IBSP 138974; IBSP 138984; IBSP 138991; IBSP 138993; IBSP 138994; IBSP 138997); all collected in 08–12.X.2001 by Equipe Biota ; Pinheiral , Fazenda Santa Helena ( 22º34’00”S ; 44º21’00”W ), 21♂ , 3♀ (IBSP 139401; IBSP 139406-IBSP 139408; IBSP 139410; IBSP 139411; IBSP 139414; IBSP 139415; IBSP 139418-IBSP 139420; IBSP 139423-IBSP 139427; IBSP 139430; IBSP 139432- IBSP 139434); all collected in 05–11.X.1999 by A.D. Brescovit et al. ; Pinheiro (currently Piraí , Pinheiral ) ( 22°30′46″S ; 44°00′3″W ), 2♂ , I.1988 , R.L.C. Baptista leg. (MNRJ 3189) ; 1♂ , 1♀ , 4#imm. [without date], C.F. de Mello-Leitão leg. (MNRJ 616); São Paulo : Teodoro Sampaio, Porto Cabral , Rio Paraná ( 24°31’00.1”S ; 47°42’00”W ), 1♀ , 1941, L.P. Travassos Filho leg. (MZSP 9788) ; Ribeirão Preto ( 21°10′40″S ; 47°48′36″W ), 1♂ , 1992, E. Camillo leg. (IBSP 210237) ; Itapetin- inga, Estação Ecológica de Angatuba ( 23º24’00”S ; 48º21’00”W ), 2♂ , 7♀ (IBSP 138710; IBSP 138711; IBSP 138714; IBSP 138717; IBSP 138720; IBSP 138722; IBSP 138723; IBSP 138725), all collected in 11–16.XI.2002 by Equipe Biota ; Iporanga ( 24°35’09”S ; 48°35’34”W ), Lajeado , 1♂ , XII.1944 , J. Damico leg. (MZSP 12217); Itaberá ( 23°51′43″S ; 49°8′13″W ), 2♂ , 1♀ , XI.1943 , F.S. Pereira leg. (MZSP 12207) ; Botucatu , Fazenda Butignoli , Rubião Júnior ( 23°59’21”S ; 48°26’21”W ), 1♀ , 07.I.1987 , I.M.P. Rinaldi & L.C. Forti leg. (MCN 22486) ; 1♂ , 07.X.1987 , I.M.P. Rinaldi & L.C. Forti leg. (MCN 22485) ; 1♀ , 12.II.1987 , I.M.P. Rinaldi leg. (MZSP 60386); 2♀ , 18.XII.1978 , N. Carneiro leg. (MZSP 12129); 1♀ , 02.XII.1987 , I.M.P. Rinaldi & L.C. Forti leg. (MCN 22483) ; 1♂ , 02.XII.1987 , I.M.P. Rinaldi & L.C. Forti leg. (MCN 22484) ; 3♂ , 1♀ , 14–16.XII.1966 , (MZSP 12123; MZSP 12141; MZSP 12317; MZSP 14137); all collected by V.C. de Jesus; 1♂ , 16.XII.1986 , I. Rinaldi leg. (MZSP 60360); Botucatu ( 22°53’9”S ; 48°26’42”W ), Parque Municipal , 1♀ , 11.V.1986 (MZSP 60383); 1♂ , 12.II.1987 (MZSP 60385); 1♂ , 13.I.1988 (MZSP 60361); 3♀ , 02.VIII.1988 (MZSP 60362); 3♀ , 12.II.1987 (MZSP 60363), all collected by I. Rin- aldi; Fazenda Edgárdia , campus UNESP ( 22°48’56”S ; 48°23’44”W ), 1♀ , 07.VI.1988 , I.M.P; Rinaldi leg. (MZSP 60384) ; Rio Claro ( 22°24′39″S ; 47°33′39″W ), 1♀ , 17–19.I.1962 , H.R. Filho leg. (MZSP 10216) ; Assis , Estação Ecológica de Assis ( 22º34’00”S ; 50º24’00”W ), 1♂ , 3♀ , 25–30.XI.2002 , Equipe Biota leg. (IBSP 138827; IBSP 138842; IBSP 138860) ; Atibaia ( 23º07’01’’S ; 46º33’01”W ), 1♂ , 13.X.2002 , G. Machado et al. leg. (IBSP 35177); 1♂ , X.2002 , M.B. Rosseto & G. Machado leg. (IBSP 35845) ; Biritiba Mirim ( 23°34′22″S ; 46°2′20″W ), 1♂ , 2003, I. Knysak leg. (IBSP 72718) ; Jundiaí ( 23º11’11”S ; 46º53’03”W ), 1♂ , 09–12.X.1999 , C.A. Rheims leg. (IBSP 26252) ; Reserva Biológica Municipal da Serra do Japi ( 23°12’00”S ; 46º30’00”W ), 2♂ , 2004, Alunos Curso Pós USP leg. (IBSP 138871; IBSP 138872); 1♀ , III.2007 , A.D. Brescovit et al. leg. (IBSP 215510); 1♂ , 2♀ , 15– 17.XII.2014 , Exc. Alunos USP leg. (IBSP 231027); Cabreúva ( 23º18’27”S ; 47º07’58’’W ), 27.X.2001 , D.F. Candi- ani & P.E.D. Francesconi leg. (IBSP 31976) ; Itapecerica da Serra ( 23°43′1″S ; 46°50′56″W ), 1♂ , 10.I.2006 , L.V. de Camões leg. (IBSP 75579) ; Mogi das Cruzes , Rio Tietê ( 23°31′22″S ; 46°11′16″W ), 1♀ , VII–VIII.1996 , R. Martins leg. (IBSP 13831) ; Barueri ( 23°30′38″S ; 46°54′34″W ), 1♀ , 07.I.1961 (MZSP 12250); 2♀ , 31.XII.1965 (MZSP 14108); 2♀ , 21.XII.1965 (MZSP 14139), 1♂ , 08.I.1966 (MZSP 5958); 1♂ , 08.I.1966 (MZSP 12314); 3♂ , 1♀ , 31.XI.1965 (MZSP 14136); 1♂ , 1♀ , 05.II.1965 (MZSP 14138); all collected by K. Lenko ; Embu das Artes ( 23°38′56″S ; 46°51′7″W ), 1♀ , II.1946 , F. Lane leg. (MZSP 12242) ; São Paulo ( 23º32’51”S ; 46º38’10”W ), 1♀ , X.1990 , Equipe IBSP leg. (IBSP 4367); Vila Mascote ( 23°38’44”S ; 46°40’2”W ), 1♀ , 04.XI.2002 , D.M.B. Battesti leg. (IBSP 36392) ; Perdizes ( 19°21’19’’S ; 47°16’58’’W ), 1♂ , 01.XII.1951 , H. Urban leg. (MZSP 12276) ; Parque da Previdência ( 23º34’00”S ; 45º43’00”W ), 1♂ , 22.X.1999 , F.S. Cunha leg. (IBSP 29400) ; 1♂ , 29.X.1999 , J. Im- pério leg. (IBSP 29401); Campus USP, Reserva Florestal da CUASO ( 23º33’00”S ; 46º43’00”W ), 3♀ , 14.XII.1999 (IBSP 29393); 1♀ , 29.X.1999 (IBSP 29394); 1♂ , 29.X.1999 (IBSP 29395); 1♀ , 31.X.2000 (IBSP 29396); 1♀ , 14.XII.1999 (IBSP 29398); 1♂ , 27/X/1999 (IBSP 32984); 2♀ , all collected by F.S. Cunha ; 29.X.1999 , D.F. Candi- ani leg. (IBSP 29399); 1♂ , 05.IX.2000 , J. Império leg. (IBSP 29397); Campus Instituto Butantan ( 23º33’00”S ; 46º43’00”W ), 4♂ , 2♀ , 11.XI.2004 , no coll. (IBSP 48427); 2♀ , 2010, G.S. Ruiz leg. (IBSP 168580) ; Campus Insti- tuto Butantan , Mata IBSP, 5♂ , 6♀ , 05.X.2005 , Alunos USP curso 2006 leg. (IBSP 57444); Represa de Guarapir- anga, Ilha dos Eucaliptos ( 23°44’26”S ; 46°44’09”W ), 1♂ , 09.IX.2001 , R.P. Indicatti leg. (IBSP 32580) ; Ilha dos Eucaliptos , 4♂ 8♀ (IBSP 61264; IBSP 61265; IBSP 61266; IBSP 61267; IBSP 61268; IBSP 61269; IBSP 61270; IBSP 61271; IBSP 61272); all collected in 07–13.X.2004 by I. Cizauskas & C.R.M Garcia ; Parque Estadual do Jaraguá ( 23°27′34.3″S ; 46°46′2.8″W ), 2♂ , 2♀ , 27–28.X.2005 , Curso UFRRJ leg. (IBSP 71185; IBSP 71237; IBSP 71239); 10♂ , 4♀ , 14–19.X.2002 , Equipe Biota leg. (IBSP 139596; IBSP 139598; IBSP 139599; IBSP 139602; IBSP 139603; IBSP 139605; IBSP 139615; IBSP 139616; IBSP 139618; IBSP 139624; IBSP 139636; IBSP 139647; IBSP 139648) ; Parque Estadual da Cantareira , Sede ( 23°24′5″S ; 46°35′24″W ), 1♀ , 20.XII.2000 (IBSP 85701); 2♀ , 21.XII.2000 (IBSP 85702); 1♀ , 28.II.2001 (IBSP 85703); 2♂ , 15.XII.2000 (MCN 41880; MCN 41882); 2♂ , 3♀ (MZSP 24966–MZSP 24970), all collected in 22.XII.2000 by R. Pinto da Rocha et al. ; Núcleo Pedra Grande , 1♀ , 15.XII.2002 , R. Pinto da Rocha et al. leg. (IBSP 210221); 3♂ , 3♀ (MZSP 24959–MZSP 24964), all collected in 21.XII.2000 - 6.I.2001 by R. Pinto da Rocha et al. ; Parque Estadual Fontes do Ipiranga ( 23°38′24.6″S ; 46°37′3.1″W ), 1♀ , 28.X.2017 (IBSP 222758); 1♂ , 27.VII.2018 (IBSP 222766); 1♀ , 28.X.2017 (IBSP 222768); 1♀ , 25.XI.2017 (IBSP 222769); 1♂ , 30.IX.2017 (IBSP 222777); 1♂ , 27.VIII.2017 (IBSP 222778); 1♂ , 22.VII.2017 (IBSP 222780); 1♀ , 25.XI.2017 (IBSP 222783); 1♂ , 30.IX.2017 (IBSP 222787); 1♂ , 27.VIII.2017 (IBSP 222792); 1♀ , 27.VIII.2017 (IBSP 222793); 1♀ , 27.VIII.2017 (IBSP 222794), all collected by Z. R. Mendes ; Horto Florestal (currently Parque Estadual Alberto Löfgren ) ( 23°27′33″S ; 46°38′2″W ), 1♂ , 1imm. , 1.XI.1994 , R. Pinto da Rocha leg. (MZSP 14310) ; Bosque da Saúde ( 23°34’21”S ; 46°38’70”W ), 2♀ , 09.IV.1944 , F. Lane leg. (MZSP 5165) ; 1♂ , 22.III.1942 , F. Lane leg. (MZSP 14134) ; Cocaia , Represa Billings ( 23°47’13”S ; 46°35’2”W ), 1♀ , 11.III.1951 , H. Urban leg. (MZSP 12212) ; Parque Estadual Serra da Cantareira , Sede ( 23°24′5″S ; 46°35′24″W ), 1♂ , 22.XI.2000 , R. Pinto da Rocha et al. leg. (MZSP 25011); São José do Barreiro , Parque Nacional da Serra da Bocaina ( 23°2′30″S ; 44°39′42″W ), 1♂ , 22–25.VII.2006 , F.U. Yamamoto leg. (IBSP 66747) ; Ribeirão Grande , Parque Estadual de Intervales ( 24°16′6.3″S ; 48°24′49.7″W ), in Bamboo , 1♀ , 2002–2003, M.O. Gonzaga leg. (IBSP 71452) ; São Bernardo do Campo ( 23°41′38″S ; 46°33′54″W ), Clube de Campo São Camilo , 1♂ , 1♀ , 20.X.2008 , V. Passanha & C.V. Janini leg. (IBSP 119790) ; Mairiporã ( 23°22′44″S ; 46°31′38″W ), Serra da Cantareira , 1♂ , VII.2000 , C.R. Macagnan leg. (IBSP 27347) ; Mogi Guaçu , Reserva Biológica Mogi Guaçu ( 22º10’00”S ; 47º08’00”W ), 2♀ , 16.XII.1998 , E. Camillo leg. (IBSP 28113; IBSP 28155) ; Reserva Biológica Mogi Guaçu ( 22º10’00”S ; 47º08’00”W ), 33 ♂ , 2 ♀ (IBSP 54646; IBSP 139460; IBSP 139466; IBSP 139468; IBSP 139469; IBSP 139473; IBSP 139482; IBSP 139484; IBSP 139486; IBSP 139489; IBSP 139491; IBSP 139493; IBSP 139497; IBSP 139500; IBSP 139501; IBSP 139502; IBSP 139504; IBSP 139505; IBSP 139509; IBSP 139510; IBSP139511; IBSP 139512; IBSP 139514; IBSP 139516; IBSP 139517; IBSP 139518), all collected in 13–19/X/2003 by Equipe Biota ; Pariquera-Açu (24°42’54”S; 47°52’51”W), Vale do Ribeira , cam- pus do Instituto Agronômico de Campinas , 1♂ , [without date], E. Wienskoski leg. (MZSP 60391) ; Ubatuba ( 23º26’02”S ; 45º04’16”W ), Fazenda Experimental , 1♂ , 1♀ , 14.XI.1943 , A. Zoppei leg. (MZSP 12228) ; Paraná : Cruzeiro do Iguaçu , Usina Foz do Chopim ( 25º33’45”S ; 53º10’37”W ), 1♂ , 08–15.X.1998 , Equipe IBSP leg. (IBSP 21241); Adrianópolis , Vale do Ribeira (24°39′25″S; 48°59′27″W), 1♀ , 18.XI.2004 , D.M.B. Battesti leg. (IBSP 47949) ; Bela Vista do Paraíso ( 22º59’49”S ; 51º11’27”W ), 1♂ , 1♀ , 11–16.XI.1991 , A.B. Bonaldo leg. (IBSP 221089) ; Fênix ( 23º54’57”S ; 51º58’44”W ), 1♂ , 24.XI.1986 , Equipe CIIF leg. (IBSP 221097); 1♂ , 01.XI.1986 , Equipe Profaupar leg. (IBSP 221098) ; Parque Estadual de Vila Rica do Espírito Santo ( 23°55’00”S ; 51°57’00”W ), 3♀ , 12.XI.1987 , A.D. Brescovit leg. (IBSP 221096) ; Curitiba , campus UFPR, Centro Politécnico ( 25°25′44″S ; 49°16′3″W ), 2♂ , 2♀ , 26.X.1995 , A.B. Bonaldo leg. (MCN 26698) ; Almirante Tamandaré ( 25°19′30″S ; 49°18′36″W ), 1♀ , 10.VII.1984 (MCN 12389); 1♂ , 25.IV.1984 (MCN 12414); 1♂ , 08.VI.1984 (MCN 12454); 1♂ , 25.IV.1984 (MCN 12467); 1♂ , 22.VI.1984 (MCN 12488); 1♂ , 28.IX.1984 (MCN 12517); 1♂ , 28.IX.1984 (MCN 12518); all collected by E. C. Costa ; Capanema ( 25°40′19″S ; 53°48′32″W ), 3♂ , 11.III.1981 , S. Scherer leg. (MCN 9777) ; Santa Catarina : Chapecó , Floresta Nacional de Chapecó ( 27º06’10,5”S ; 52º46’48,8”W ), 1♀ , I.2003 XII.2004 , M. Scartezini leg. (IBSP 91070) ; Nova Teutônia ( 27°11’00”S ; 52°23’00”W ), 2♂ , 1♀ , [without date], F. Plaumann leg. (FMS 13579; FMS 13583; FMS 13578) ; Florianópolis , Itacorubí , UCAD ( 27°33′21.44″S ; 48°29′37.46″W ), 1♂ , 2004, J. Steiner leg. (IBSP 91097) ; Criciúma ( 28°40′40″S ; 49°22′12″W ), 1♂ , 08.XI.2007 , R.A. Teixeira leg. (IBSP 132769) ; Rancho Queimado ( 27°40′22″S ; 49°1′19″W ), 1♂ , 15–18.XI.1995 , L.A. Moura leg. (MCN 27039) ; 2♀ , 13–15.I.1995 , A.B. Bonaldo leg. (MCN 26356) ; Rio Grande do Sul : Iraí ( 27°11′38″S ; 53°15′3″W ), 1♂ , 2♀ , 20.XI.1975 , A.A. Lise leg. (MCN 3106) ; Derrubadas , Parque Estadual do Turvo , Salto do Yucumã ( 27°08’32.5″S ; 53°53’09.0″W ), 1♀ , 16.I.1985 (MCN 13002); 1♂ , 16.I.1985 (MCN 13005); all collected by A.A. Lise ; São Fran- cisco de Paula ( 29°26′52″S ; 50°35′2″W ), 1♀ , 22.IX.1974 (MCN 2304); 1♂ , 1♀ , 22.I.1981 (MCN 9531); 1♂ , 2♀ , 23.I.1981 (MCN 9540); all collected by A.A. Lise ; Barragem dos Bugres ( 29°20’35.87”S ; 50°41’44.15”W ), 2♂ , 23–25.XI.1998 , A.F. Franceschini leg. (MCN 30656) ; 2♂ , 1♀ , 01–04.II.1999 , L.A. Moura leg. (MCN 30669) ; Rincão dos Kroeff ( 29°28’3”S ; 50°20’47”W ), 1♀ , 05.I.1985 , A.A. Lise leg. (MCN 12739) ; Cambará do Sul , Itaim- bézinho ( 29º02’52”S ; 50º08’41”W ), 2♂ , 1♀ , 06.I.1985 , A. Lise leg. (MCN 12777; MCN 12779; MCN 12780) ; Garabi ( 28°12’50”S ; 55°41’17”W ), 1♀ , XI.1988 , A.A. Lise leg. (IBSP 258334) ; Marcelino Ramos , Estreito Au- gusto Cesar ( 27°24’51”S , 51°27’2”W ), 1♀ , 03.II.1990 , C.A. Martinazzo leg. (MCN 19530) ; Maquiné ( 29°40’30”S ; 50°12’25”W ), Fepagro , 1♀ , 18–27.I.2002 , Equipe Biota leg. (IBSP 139226) ; Derrubadas , Parque Estadual do Turvo ( 27°8’44”S ; 53°53’10”W ), 6♀ (IBSP 139987; IBSP 139989; IBSP 139990; IBSP 140007; IBSP 140018), all col- lected in 11–18.I.2002 by Equipe Biota ; Parque Estadual do Turvo (27°8’44”S, 53°53’10”W), 12♂ , 3♀ , 27– 31.X.2002 (MCN 37768); 1♂ , 1♀ , 19–22.X.2004 (MCN 38812); 1♂ , 27–31.X.2003 (MCN 38813); 6♂ , 27–31. X.2003 (MCN 38814); 1♂ , 19–22.X.2004 (MCN 38836); 10♂ , 6♀ , 19–22.X.2004 (MCN 38841), all collected by R. Ott et al. ; Canela ( 29°21’57”S ; 50°48’57”W ), 1♀ , 31.XII.1973 (MCN 2070); 1♂ , 20.III.1976 (MCN 3854); 1♀ , 05.III.1977 (MCN 5209); 1♀ , X.1976 (MCN 6300), all collected by A.A. Lise ; 1♂ , 1♀ , 03–04.IX.1988 , A.B. Bon- aldo leg. (MCN 17841); 1♀ , 05.II.1973 (MCN 852); 2♂ , 31.XII.1973 (MCN 2044); 1♀ , 11.V.1974 (MCN 2163); 1♂ , 26.XII.1974 (MCN 2475); 1♂ , 1♀ , 26.II.1975 (MCN 3748); all collected by A.A. Lise ; Carazinho ( 28°17’2”S ; 52°47’9”W ), 2♂ , 10.XI.1979 , H. Bischoff leg. (MCN 8655) ; 1♀ , 10.XI.1979 , A.A. Lise leg. (MCN 8686) ; So- bradinho ( 29°25’15”S ; 53°1’44”W ), 2♀ , 10.I.1985 , A.A. Lise leg. (MCN 18717) ; Marcelino Ramos , Estreito Au- gusto Cesar ( 27°24’51”S ; 51°27’2”W ), 1♀ , 03.II.1990 , C.A. Martinazzo leg. (MCN 19529) ; Nova Petrópolis , Boê- mios, Linha Imperial ( 29°21’36”S ; 51°2’54.8”W ), 1♂ , 20.XII.1973 , A.A. Lise leg. (MCN 2206) ; Caxias do Sul , Vila Oliva ( 29°12’56”S ; 50°53’22”W ), 1♂ , 10.I.1976 , P.A. Buckup leg. (MCN 3959) ; Caxias do Sul ( 29°10’04”S ; 51°10’44”W ), Água Azul , 1♀ , 15.IX.1976 , H. Bischoff leg. (MCN 4532) ; 1♂ , 04.IV.1975 , A.A. Lise leg. (MCN 18708) ; General Câmara ( 29°54’18”S ; 51°45’36”W ), 1♂ , 1♀ , 14.X.1982 , C.J. Becker leg. (MCN 10810) ; Far- roupilha ( 29°13’30”S ; 51°20’52”W ), 2♂ , 1♀ , 29.IX.1978 , H. Bischoff leg. (MCN 8283; MCN 8306; MCN 8329); Agudo , Usina Hidrelétrica Dona Francisca ( 29°27’7”S ; 53°16’55”W ), 1♀ , 02.XI.2002 , A.B.B. Morais leg. (IBSP 122191) ; 1♀ , Passo Fundo ( 28°15’46”S 52°24’25”W ), 1♀ , 13.X.1985 , A.A. Lise leg. (MCN 14321) ; 1♂ , 01.VIII.1986 , A.A. Lise leg. (MCN 15573) ; Estrela Velha , Barragem de Itaúba ( 29°15’40”S ; 53°14’8”W ), 2♂ , 1♀ , 21.X.1998 , A.B. Bonaldo leg. (MCN 30606; MCN 30607) ; Garruchos ( 28°11’2”S ; 55°38’20”W ), 4♀ , 10.XII.1975 (MCN 3152); 3♀ , 10.XII.1975 (MCN 3172); 1♀ , 06.XII.1975 (MCN 18703); all collected by A.A. Lise ; Triunfo ( 29°56’34”S ; 51°43’4”W ), 2♂ , 1♀ , 15.IX.1977 , H. Bischoff leg. (MCN 6516; MCN 6969) ; 1♂ , 27.X.1977 , M.H. Galileo leg. (MCN 7031) ; 1♂ , 2♀ , 28.XI.1977 , A.A. Lise leg. (MCN 7393); Polo Petroquímico do Sul ( 29°54’4”S ; 51°22’27”W ), 1♂ , 1♀ , 20.X.1977 , E.H. Buckup leg. (MCN 18706; MCN 18707) ; Parobé ( 29°37’44”S ; 50°50’6”W ), Rio dos Sinos , 1♀ , 10.XI.2008 , E.N.L. & P.E.S. Rodrigues leg. (MCN 47891) ; 1♀ , 18.XII.2007 , E.N.L. & P.E.S. Rodrigues leg. (MCN 48029) ; Parque Braskem de Proteção Ambiental ( 29°51’59.44”S ; 51°21’54.00”W ), 2♂ , 24.X.1989 , A.D. Brescovit leg. (MCN 18903) ; 1♂ , 28.XI.1989 , E.H. Buckup leg. (MCN 19042) ; 1♀ , 24.XI.1999 , M.A.L. Marques leg. (MCN 31739) ; 1♂ , 21.X.2003 , A. Barcellos & L. Schmidt leg. (MCN 36458) ; 1♀ , 2.XI.2007 , A. Barcellos & L. Schmidt leg. (MCN 43644) ; 1♀ , 04.XII.2009 , M.R.M. Poeta leg. (MCN 46917) ; 2♂ , 04.XII.2009 , M. Pairet Jr , leg. (MCN 46925); 1♀ , 04.XII.2009 , P.E.S. Rodrigues leg. (MCN 46933) ; Arroio Grande , Mata Ciliar do Rio Piratini ( 31°54’06.47”S ; 52°39’08.29”W ), 1♀ , 13.VI.2008 , E.N.L. & P.E.S. Rodrigues leg. (MCN 54732) ; Campo Bom ( 29°40’44″S ; 51°3’10”W ), 1♀ , 16.XI.1979 , C.J. Becker leg. (MCN 8789) ; Novo Hamburgo ( 29º40’40”S ; 51º07’51”W ), 2♂ , 22.IX.1986 , C.J. Becker leg. (IBSP 221094) ; 1♂ , 21.X.1983 (MCN 11814); 1♀ , 25.XI.1983 (MCN 11847); 2♂ , 11.XII.1984 (MCN 12645); 2♂ , 07.X.1985 (MCN 14262; MCN 12646); 2♂ , 14.XII.1984 (MCN 12717); 1♀ , 31.III.1986 , C.J. Becker leg. (MCN 14557) , all collected by C.J. Becker; Morro Reuter ( 29°32’16”S ; 51°4’51”W ), 1♀ , 28.III.1976 , C.J. Becker leg. (MCN 3576) ; Viamão , Estação Experimental Fitotécnica Águas Belas ( 30°05’00”S ; 51°02’00”W ), 1♂ , 30.XI.1984 , M.A.L. Marques leg. (MCN 12611) ; 1♂ , 06.I.1977 , A.A. Lise leg. (MCN 5899); São Leopoldo ( 29º45’36”S ; 51º08’49”W ), 1♂ , 28.IX.1987 , C.J. Becker leg. (IBSP 221095) ; 2♂ , 17.IX.1982 (MCN 10965); 3♂ , 24.IX.1982 (MCN 11049); 2♂ , 15.X.1982 (MCN 11146); 1♀ , 15.X.1982 (MCN 11147); 1♀ , 20.XII.1982 (MCN 11339); 2♂ , 12.XII.1983 (MCN 11865); 1♂ , 26.XII.1983 (MCN 11911); all collected by C.J. Becker ; Montenegro ( 29°41’20”S ; 51°27’39”W ), 2♂ , 2♀ , 06.X.1977 (MCN 6806; MCN 7277); 3♂ , 17.XI.1977 (MCN 7278); 1♀ , 20.XII.1985 (MCN 14077); 4♂ , 29.IX.1977 (MCN 6718); 1♂ , 17.XI.1977 , (MCN 7280); 8♂ , 01.XII.1977 (MCN 7482); 4♂ , 20.XII.1977 (MCN 7667); all collected by E.H. Buckup & A.A. Lise ; Tapes ( 30°40’22”S ; 51°23’45”W ), Fazenda Guará , 1♀ , 19.XII.2003 , Equipe Probio leg. (MCN 36821) ; Fazenda São Miguel , 2♂ , 1♀ , 17.XII.2003 , Equipe Probio leg. (MCN 36847) ; Barra do Ribeiro ( 30°17’27”S ; 51°18’3”W ), Fazenda Boa Vista , 2♂ , 18.XII.2003 , Equipe Probio leg. (MCN 36850) ; Capão do Leão , Campus UFPEL ( 31°45’46”S ; 52°29’2”W ), 1♀ , 24.II.2000 (IBSP 36534); 2♂ , 1♀ , 30.IX.2001 (IBSP 36536); 2♂ , 30.IX.2001 (IBSP 36537); 1♀ , 30.IX.2001 (IBSP 36538); all collected by E.N.L. Rodrigues. PARAGUAY . Departamento Paraguari : Parque Nacional de Ibycuí ( 26º05’49.08”S ; 56º50’20.16”W ), 1♀ , 28.I.1982 (MNHNP); 1♀ , 20.X.1981 (MNHNP) ; Departamento Amambay : Parque Nacional Cerro Corá , Base de Cerro Tuya Hog ( 22°38’28”S ; 56°0’47”W ), 1♀ , X.1983 , (MNHNP); 1♀ , 8.VI.1982 (IBNP) ; Departamento Itapúa : Pirapó ( 28°2’42”S ; 55°11’56”W ), 1♂ , 4.VIII.1982 (MNHNP) ; Departamento Caaguazú : Caaguazú ( 25°28’11”S ; 56°0’55”W ), 1♂ , 23.VII.1982 (MNHNP); all collected by J.A. Kochalka. ARGENTINA . Misiones : Loreto ( 27°19’55.12”S ; 55°31’23.47”W ), 1♂ , 27.III.1932 8/III/1933 , A.A. Oglobin leg. (MLP) ; Puerto Iguazú ( 25°35’50”S ; 54°34’41”W ), 8♀ , XI.1948 , M. Birabén leg. (MLP) ; Puerto Victoria ( 26°20’04”S ; 54°39’44”W ), 1♀ , [without date], A. Zenzes leg. (MLP) ; Parque Nacional del Iguazú , Cataratas del Iguazú ( 25°41’43”S ; 54°26’12”W ), 2♂ , XI.1989 , (MACN- Ar ); 1♀ , XI.1991 , (MACN-Ar); 1♂ , 1♀ , XI.1992 , (MACN-Ar); all collected by M.J. Ramírez ; Cantera , 200 m , 1♀ , 8.XII.1990 06.I.1991 , S. & J. Peck leg. (AMNH) ; Porto Canoas , 180m , 1♀ , 8.XII.1990 06.I.1992 , S. & J. Peck leg. (AMNH) ; Eldorado ( 26°28’00”S ; 54°43’00”W ), 2♂ , 01.X–15.XI.1964 , A. Kovacs leg. (AMNH) ; 2♂ , 1♀ , 01.X– 15.XI.1965 , A. Kovacs leg. (AMNH). Diagnosis. Males of Patrera procera can be distinguished from the remaining species by the characteristic tibial retrolateral apophysis wide, subrectangular, with a transverse fold at the tip, forming a conspicuous groove, and by the conical dorsal tibial apophysis ( Figs 1D , 2B , 3E ) in the male palp. Females are diagnosed by their short, narrow epigynal lateral borders with lateral excavation at the internal borders and wide deep atrium in the epigynum ( Figs 2C , 3F ). FIGURE 1A–G. Patrera procera (Keyserling, 1981) . Male (IBSP 138902): A habitus, dorsal view; C left palp, ventral view; D same, retrolateral view; E same, dorsal view. Female (IBSP 138902): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view. Abbreviations: A, atrium; dTA, dorsal tibial apophysis; E, embolus; LB, lateral borders; MA, median apophysis; RTA, retrolateral tibial apophysis; T, tegulum. Scale bars: A, B 3.5mm; C 0.6mm; D 0.73mm; E 0.66mm; F 0.4mm; G 0.25mm. FIGURE 2A–D. Patrera procera ( Keyserling, 1891 ) . Male (MCN 10965): A left palp, ventral view; B same, retrolateral view. Female (MCN 3172): C epigynum, ventral view (arrow indicating the excavation at the lateral borders); D same, dorsal view. Abbreviations: A, atrium; CD, copulatory ducts; dTA, dorsal tibial apophysis; E, embolus; FD, fertilization ducts; H, hood; LB, lateral borders; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SR, seminal receptacles; T, tegulum. Scale bars: A–D 0.25mm. FIGURE 3A–F. Patrera procera ( Keyserling, 1891 ) . Male (IBSP 139486): A chelicerae, ventral view; B left palp, ventral view; C same, distal view; D same, ventral view; E same, retrolateral view. Female (IBSP 138902): F epigynum, ventral view (arrow indicating the excavation at the lateral borders). Abbreviations: A, atrium; CO, copulatory opening; dTA, dorsal tibial apophysis; E, embolus; LB, lateral borders; MA, median tibial apophysis; RTA, retrolateral tibial apophysis; T, tegulum. Scale bars: A 0.3mm; B 0.4mm; C, F 0.2mm; D, E 0.1mm. Description. Male ( Lectotype , BMNH). Carapace orange brown, with orange paramedian stripes. Chelicerae brown yellowish, labium and endites orange. Sternum light orange, with dark borders. Legs orange. Abdomen dorsally brownish, ventrally cream ( Fig. 1A ). Total length 8.6, carapace length 3.5, width 2.65. Clypeus height 0.15. Eye diameters and interdistances: AME 0.11, ALE 0.20, PME 0.17, PLE 0.20; AME–AME 0.06, AME–ALE 0.08, PME–PME 0.21, PME–PLE 0.17, ALE–PLE 0.07. Chelicerae 2.3 long, four promarginal teeth, nine small retromarginal denticles ( Fig. 3A ). Leg measurements: leg I—femur 5.5/ patella 1.5/ tibia 6.1/ metatarsus 5.2/ tarsus 2.05/ total 20.35; II—4.85/ 1.3/ 5.25/ 4.6/ 1.9/ 17.9; III—3.1/ 1.05/ 2.5/ 3.4/ 1.0/ 11.05; IV—4.8/ 1.3/ 4.35/ 5.75/ 1.2/ 17.75. Leg spination: I—II—tibia v2-2-0, p1-1-1, r1-1-1, metatarsus v2-0-0, p1-1-0, r1-1-0; III—IV—tibia v2- 2-2, p1-1-1, r1-1-1, metatarsus v2-2-2, p1-1-1, r1-1-1. Abdomen: length 5.35, epigastric furrow 1.4 from tracheal spiracle, spiracle 2.3 from base of spinnerets. Palp: tibia as long as cymbium; tegulum oval; spermatic duct “Ushaped” ventrally; short distal tegular projection, close to the embolus apex; subdistal median apophysis, inverted “S-shaped”; embolus with median prolateral origin, thin, with broad base, slightly curved ( Figs 1C , 2A , 3B, C ). Female ( Paralectotype BMNH). Coloration as in male, except dorsal area of the abdomen lighter ( Fig. 1B ). Total length 9.2, carapace length 3.8, width 2.8. Clypeus height 0.16. Eye diameters and interdistances: AME 0.11, ALE 0.20, PME 0.16, PLE 0.18; AME–AME 0.07, AME–ALE 0.06, PME–PME 0.22, PME–PLE 0.18, ALE–PLE 0.06. Chelicerae 1.67 long, five promarginal teeth, eight retromarginal denticles. Leg measurements: leg I—femur 4.8/ patella 1.55/ tibia 5.1/ metatarsus 4.1/ tarsus 2.1/ total 17.65; II—4.3/ 1.4/ 4.5/ 3.85/ 1.7/ 15.75; III—3.1/ 1.2/ 2.4/ 3.1/ 1.0/ 10.8; IV—4.6/ 1.4/ 4.1/ 5.0/ 5.0/ 1.3/ 16.4. Leg spination: I—tibia v2-2-0, p1-0-0, r1-0-0, metatarsus v2-0-0, p0, r0; II—tibia v2-2-0, p1-1-1, r0, metatarsus v2-0-0, p1-0-0, r0; III—IV—tibia v2-2-2, p1-1-1-, r1-1-1, metatarsos v2-2-2, p1-1-1, r1-1-1. Abdomen: length 5.7, epigastric furrow 1.55 from tracheal spiracle, spiracle 2.25 from base of spinnerets. Epigynum: lateral borders narrow, not distally joined; internally with long, coiled copulatory ducts; seminal receptacles in the proximal area of the copulatory ducts; oval spermathecae; fertilization ducts as long as the spermathecae, curved and basally inserted ( Figs 1F, G , 2C, D , 3F ). Variation. Males (n=10): total length 7.4–9.2; carapace length 2.8–3.8; femur I length 3.8–5.2. Females (n=10): total length: 8.5–9.7; carapace length 3.3–3.6; femur I length 4.2–4.8. Distribution . Northeast to southern Brazil , West Paraguay and Northeastern Argentina ( Fig. 27C ).