<document id="8F6CAC705E04234CB5FA6ED9E4698E8C" ID-DOI="10.3767/000651914X684376" ID-ISSN="2212-1676" ID-Zenodo-Dep="16999789" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1756503404943" checkinUser="felipe" docAuthor="Veldkamp, J. F." docDate="2014" docId="03B70E2F8F69FFFFFCBBA257FD07FDC8" docLanguage="en" docName="Blumea.59.1.59-75.pdf" docOrigin="Blumea 59 (1)" docSource="https://doi.org/10.3767/000651914x684376" docStyle="DocumentStyle:00E2B8BC255D682B3CB921028E4A34F4.2:Blumea.2011-.journal_article.type1" docStyleId="00E2B8BC255D682B3CB921028E4A34F4" docStyleName="Blumea.2011-.journal_article.type1" docStyleVersion="2" docTitle="Cenchrus echinatus" docType="treatment" docVersion="2" lastPageNumber="68" masterDocId="FF8E76578F61FFF6FF90A428FFECFFF0" masterDocTitle="A revision of Cenchrus incl. Pennisetum (Gramineae) in Malesia with some general nomenclatural notes" masterLastPageNumber="75" masterPageNumber="59" pageNumber="67" updateTime="1756513612755" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="AD86A9F83A3D08F08C3622685E63F1B2">A revision of Cenchrus incl. Pennisetum (Gramineae) in Malesia with some general nomenclatural notes</mods:title>
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<mods:namePart id="6D2D52F7BD3F075BFA7189B2A84C5238">Veldkamp, J. F.</mods:namePart>
<mods:affiliation id="BD64477A5260C6DB2F6C9316A7ACDDE3">Naturalis Biodiversity Center, section Botany, P. O. Box 9517, 2300 RA Leiden, The Netherlands; e-mail: jef. veldkamp @ naturalis. nl</mods:affiliation>
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<mods:title id="F9B9C5E4307AF08353B9E1D4AC8F0885">Blumea</mods:title>
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<mods:date id="AB194E1802E1F6262AF9358EDA0DD739">2014</mods:date>
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<subSubSection id="C304ECB28F69FFFEFCBBA257FBAAF966" box="[811,1094,1663,1686]" pageId="8" pageNumber="67" type="nomenclature">
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<heading id="D0E908558F69FFFEFCBBA257FBAAF966" box="[811,1094,1663,1686]" fontSize="9" level="2" pageId="8" pageNumber="67" reason="4">
<emphasis id="B96A632B8F69FFFEFCBBA257FBC7F966" bold="true" box="[811,1067,1663,1686]" pageId="8" pageNumber="67">
7. 
<taxonomicName id="4C1EC4BA8F69FFFEFCD5A257FBC7F966" ID-CoL="S5FJ" box="[837,1067,1663,1686]" class="Liliopsida" family="Poaceae" genus="Cenchrus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="species" species="echinatus">
<emphasis id="B96A632B8F69FFFEFCD5A257FBC7F966" bold="true" box="[837,1067,1663,1686]" italics="true" pageId="8" pageNumber="67">Cenchrus echinatus</emphasis>
</taxonomicName>
</emphasis>
L.
</heading>
</paragraph>
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<taxonomicName id="4C1EC4BA8F69FFFEFCBBA285FC35F94F" box="[811,985,1709,1727]" class="Liliopsida" family="Poaceae" genus="Cenchrus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="species" species="echinatus">
<emphasis id="B96A632B8F69FFFEFCBBA285FC35F94F" box="[811,985,1709,1727]" italics="true" pageId="8" pageNumber="67">Cenchrus echinatus</emphasis>
</taxonomicName>
<materialsCitation id="3B76B5648F69FFFEFC4FA285FBBAF902" collectionCode="L, IDC" location="Royen" pageId="8" pageNumber="67" specimenCode="BT-341" specimenCount="1" stateProvince="Van" typeStatus="lectotype">
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. (1753) 1050; Cav. (1799b) 39, t. 462. — 
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: 
<emphasis id="B96A632B8F69FFFEFCAAA2EFFC15F929" box="[826,1017,1735,1753]" italics="true" pageId="8" pageNumber="67">
Herb. 
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s.n.
</emphasis>
(holo 
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, sh. 912.356-116; microfiche 
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), designated by Veldkamp (1993).
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</paragraph>
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<taxonomicName id="4C1EC4BA8F69FFFEFCBBA2D1FC3BF8FC" box="[811,983,1785,1804]" class="Liliopsida" family="Poaceae" genus="Cenchrus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="67" phylum="Tracheophyta" rank="species" species="echinatus">
<emphasis id="B96A632B8F69FFFEFCBBA2D1FC3BF8FC" box="[811,983,1785,1804]" italics="true" pageId="8" pageNumber="67">Cenchrus echinatus</emphasis>
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<collectionCode id="ED0F27FC8F69FFFEFC4BA2D2FC04F8FC" box="[987,1000,1786,1804]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="8" pageNumber="67" type="Herbarium">L</collectionCode>
. 
<taxonomicName id="4C1EC4BA8F69FFFEFC60A2D2FB21F8FC" authority="F. Br. (1931)" authorityName="F. Br." authorityYear="1931" box="[1008,1229,1786,1804]" pageId="8" pageNumber="67" rank="variety" variety="glabratus">
var. 
<emphasis id="B96A632B8F69FFFEFB84A2D2FB89F8FC" box="[1044,1125,1786,1804]" italics="true" pageId="8" pageNumber="67">glabratus</emphasis>
F.Br. (1931)
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66. — 
<typeStatus id="54A5019B8F69FFFEFA9BA2D2FA88F8FC" box="[1291,1380,1786,1804]" pageId="8" pageNumber="67" type="lectotype">Lectotype</typeStatus>
: 
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<collectionCode id="ED0F27FC8F69FFFEFAFAA2D2FA9BF8FC" box="[1386,1399,1786,1804]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="8" pageNumber="67" type="Herbarium">F</collectionCode>
. Brown 745
</emphasis>
(holo 
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), designated by St. 
<bibRefCitation id="EF8FC2C88F69FFFEFBFFA33BFB37F8D5" author="St John H." box="[1135,1243,1811,1829]" pageId="8" pageNumber="67" pagination="419" refId="ref24192" refString="St John H. 1976. Lectotypes for Brown's Flora of Southeastern Polynesia. Phytologia 33: 419." type="journal article" year="1976">John (1976)</bibRefCitation>
.
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</paragraph>
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Annuals. 
<emphasis id="B96A632B8F69FFFEFC19A314FC27F8A3" box="[905,971,1852,1875]" italics="true" pageId="8" pageNumber="67">Culms</emphasis>
0.15–0.6(–1) m long,erect to geniculate at base, rooting in the decumbent nodes (sometimes). 
<emphasis id="B96A632B8F69FFFEFA8CA372FAB0F881" box="[1308,1372,1882,1905]" italics="true" pageId="8" pageNumber="67">Ligule</emphasis>
<quantity id="4CE612DC8F69FFFEFAF5A372FCBFF87E" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.7" metricValueMin="0.7" pageId="8" pageNumber="67" unit="mm" value="1.2" valueMax="1.7" valueMin="0.7">0.7–1.7 mm</quantity>
long. Leaf blades 4–26(–35) cm by 3.5–8(–12) mm. Inflorescence 3–7(–10) cm long. Common axis scaberulous, internodes 
<quantity id="4CE612DC8F69FFFEFC31A39AFC10F839" box="[929,1020,1970,1993]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="8" pageNumber="67" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long. Stipe 1–3 by 
<quantity id="4CE612DC8F69FFFEFB58A39AFAA3F839" box="[1224,1359,1970,1993]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.6" metricValueMin="2.2" pageId="8" pageNumber="67" unit="mm" value="2.9" valueMax="3.6" valueMin="2.2">2.2–3.6 mm</quantity>
, base obconical, pubescent. Burs loosely spaced, 4–7(–10) by 
<quantity id="4CE612DC8F69FFFEFAE4A3E7FCBBF7F4" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="6.0" metricValueMin="3.5" pageId="8" pageNumber="67" unit="mm" value="4.75" valueMax="6.0" valueMin="3.5">3.5–6 mm</quantity>
, becoming purple. Outer main bristles shorter than the inner spines, retrorsely barbed. Inner spines c. 10, connate for more than halfway above the base, forming a closed bur, flat, erect to sometimes interlocking, subequal, puberulous, margins densely pilose. 
<emphasis id="B96A632B8F68FFFFFE9AA488FE87FF47" box="[266,363,160,183]" italics="true" pageId="9" pageNumber="68">Spikelets</emphasis>
(1–)2–4(–6) per bur, 
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long. Lower glume present, 
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long, 1-nerved; upper glume 
<quantity id="4CE612DC8F68FFFFFFF4A4F4FF06FF03" box="[100,234,220,243]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.7" metricValueMin="3.8" pageId="9" pageNumber="68" unit="mm" value="4.75" valueMax="5.7" valueMin="3.8">3.8–5.7 mm</quantity>
long, 3–5-nerved. First lemma paleate, sterile, 
<quantity id="4CE612DC8F68FFFFFFF4A4D1FF09FEE0" box="[100,229,249,272]" metricMagnitude="-3" metricUnit="m" metricValue="5.45" metricValueMax="6.4" metricValueMin="4.5" pageId="9" pageNumber="68" unit="mm" value="5.45" valueMax="6.4" valueMin="4.5">4.5–6.4 mm</quantity>
long, 3–5-nerved. Second lemma 
<quantity id="4CE612DC8F68FFFFFDDDA4D1FD5AFEE0" box="[589,694,249,272]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="7.0" metricValueMin="4.7" pageId="9" pageNumber="68" unit="mm" value="5.85" valueMax="7.0" valueMin="4.7">4.7–7 mm</quantity>
long. Anthers 
<quantity id="4CE612DC8F68FFFFFF2DA53FFEADFEDE" box="[189,321,279,302]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="2.4" metricValueMin="0.8" pageId="9" pageNumber="68" unit="mm" value="1.6" valueMax="2.4" valueMin="0.8">0.8–2.4 mm</quantity>
long. 2n = 34, 68, also 70.
</paragraph>
<paragraph id="8BA1BF398F68FFFFFF12A51CFD53FE2F" blockId="9.[100,751,160,568]" pageId="9" pageNumber="68">
Distribution — Warmer regions of the New World, introduced elsewhere, e.g. 
<collectingCountry id="F309FFA98F68FFFFFE83A57AFE86FE99" box="[275,362,338,361]" name="Malaysia" pageId="9" pageNumber="68">Malesia</collectingCountry>
: Malay Pen. (
<collectingRegion id="49DA71DB8F68FFFFFD90A57AFD84FE99" box="[512,616,338,361]" country="Malaysia" name="Selangor" pageId="9" pageNumber="68">Selangor</collectingRegion>
), 
<collectingCountry id="F309FFA98F68FFFFFDEAA57AFD07FE99" box="[634,747,338,361]" name="Singapore" pageId="9" pageNumber="68">Singapore</collectingCountry>
, Lesser Sunda Isl. (Bali), Borneo (
<collectingCountry id="F309FFA98F68FFFFFE29A558FDE8FE77" box="[441,516,368,391]" name="Brunei" pageId="9" pageNumber="68">Brunei</collectingCountry>
, E Kalimantan), 
<collectingCountry id="F309FFA98F68FFFFFD3CA558FF71FE54" name="Philippines" pageId="9" pageNumber="68">Philippines</collectingCountry>
(
<collectingRegion id="49DA71DB8F68FFFFFF3EA5A5FF01FE54" box="[174,237,397,420]" country="Philippines" name="Cebu" pageId="9" pageNumber="68">Cebu</collectingRegion>
, Luzon, 
<collectingRegion id="49DA71DB8F68FFFFFEDBA5A5FE47FE54" box="[331,427,397,420]" country="Philippines" name="Palawan" pageId="9" pageNumber="68">Palawan</collectingRegion>
, 
<collectingRegion id="49DA71DB8F68FFFFFE28A5A5FDE9FE54" box="[440,517,397,420]" country="Philippines" name="Samar" pageId="9" pageNumber="68">Samar</collectingRegion>
), 
<collectingCountry id="F309FFA98F68FFFFFD87A5A5FD02FE54" box="[535,750,397,420]" name="Papua New Guinea" pageId="9" pageNumber="68">Papua New Guinea</collectingCountry>
(Central, 
<collectingRegion id="49DA71DB8F68FFFFFF53A583FF1EFE32" box="[195,242,427,450]" country="Papua New Guinea" name="Gulf" pageId="9" pageNumber="68">Gulf</collectingRegion>
, 
<collectingRegion id="49DA71DB8F68FFFFFF69A583FEA1FE32" box="[249,333,427,450]" country="Papua New Guinea" name="Morobe" pageId="9" pageNumber="68">Morobe</collectingRegion>
, 
<collectingRegion id="49DA71DB8F68FFFFFEC4A583FDE2FE32" box="[340,526,427,450]" country="Papua New Guinea" name="New Ireland" pageId="9" pageNumber="68">New Ireland Prov.</collectingRegion>
, 
<collectingRegion id="49DA71DB8F68FFFFFD85A583FDB0FE32" box="[533,604,427,450]" country="Papua New Guinea" name="Manus" pageId="9" pageNumber="68">Manus</collectingRegion>
Isl.), 
<collectingCountry id="F309FFA98F68FFFFFD00A583FD03FE32" box="[656,751,427,450]" name="Ashmore and Cartier Islands" pageId="9" pageNumber="68">Ashmore</collectingCountry>
Reef, N 
<collectingCountry id="F309FFA98F68FFFFFF2BA5E0FEF0FE2F" box="[187,284,456,479]" name="Australia" pageId="9" pageNumber="68">Australia</collectingCountry>
, Polynesia (Carolines to Easter Island).
</paragraph>
<paragraph id="8BA1BF398F68FFFFFF12A5CEFD80FDEB" blockId="9.[100,751,160,568]" pageId="9" pageNumber="68">
Habitat — Waste places, beaches, roadsides, fields, grassy slopes, 
<taxonomicName id="4C1EC4BA8F68FFFFFF27A62CFEC8FDEB" box="[183,292,516,539]" class="Magnoliopsida" family="Myrtaceae" genus="Melaleuca" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="68" phylum="Tracheophyta" rank="genus">
<emphasis id="B96A632B8F68FFFFFF27A62CFEC8FDEB" box="[183,292,516,539]" italics="true" pageId="9" pageNumber="68">Melaleuca</emphasis>
</taxonomicName>
savannah, 
<quantity id="4CE612DC8F68FFFFFE30A62CFDFCFDEB" box="[416,528,516,539]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="9" pageNumber="68" unit="m" value="1000.0" valueMax="2000.0" valueMin="0.0">0–2000 m</quantity>
altitude.
</paragraph>
<paragraph id="8BA1BF398F68FFFFFF12A609FD07FDC8" blockId="9.[100,751,160,568]" box="[130,747,545,568]" pageId="9" pageNumber="68">Vernacular names — Hedgehog grass, Southern sandbur.</paragraph>
</subSubSection>
</treatment>
</document>