Seven new giant pill-millipede species and numerous new records of the genus Zoosphaerium from Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae) Author Wesener, Thomas 86DEA7CD-988C-43EC-B9D6-C51000595B47 Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Section Myriapoda, Adenauerallee 160, D- 53113 Bonn, Germany. t.wesener@leibniz-zfmk.de Author Sagorny, Christina 9C89C1B7-897A-426E-8FD4-C747DF004C85 Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Section Myriapoda, Adenauerallee 160, D- 53113 Bonn, Germany. & University of Bonn, Institute of Evolutionary Biology and Ecology, D- 53121 Bonn, Germany. csagorny@evolution.uni-bonn.de text European Journal of Taxonomy 2021 2021-07-06 758 1 1 48 http://dx.doi.org/10.5852/ejt.2021.758.1423 journal article 10.5852/ejt.2021.758.1423 de9c0c06-0e7d-44a6-a582-d41a346233f6 2118-9773 5075824 01BBC12C-E715-4393-A9F6-6EA85CB1289F Unidentifiable Zoosphaerium spp. (mainly immatures) Material examined MADAGASCAR Antsiranana 1 spec. ; Marojejy Res. ; 4 km NNW of Manantenina ; 14°26′ S , 49°45′ E ; alt. 700 m ; 10–16 Nov. 1993 ; J. Coddington , N. Scharff , S. Larcher , C. Griswold and R. Andriamasimanana leg.; NHMD 1 ♀ ; Montagne dʼAmbre ; 12°32′ S , 49°10′ E ; alt. 1100 m ; 23–28 Nov. 1993 ; J. Coddington , N. Scharff , S. Larcher , C. Griswold and R. Andriamasimanana leg.; NHMD 1 spec. ; same collection data as for preceding; NHMD 2 specs ; N Ankarana Reserve , base of tsingy summit ; 12.8461° S , 49.15° E ; alt. 284 m ; dry deciduous forest on limestone ; 8 Apr. 2007 ; Ken Emberton leg.; coll. no. MBI-2038; FMNH-INS 000 103 908 1 spec. ; N Ankarana Reserve , local tsingy , summit; 12.8465° S , 49.1503° E ; alt. 294 m ; dry deciduous forest on limestone ; 8 Apr. 2007 ; Ken Emberton leg.; coll. no. MBI-2037; FMNH-INS 000 103 909 1 spec. ; NE Ankarana Reserve , roadside ; 12.8329° S , 49.2286° E ; alt. 331 m ; dry deciduous forest on lava ; 14 Apr. 2007 ; Ken Emberton leg.; coll. no. MBI- 2066; FMNH-INS 000 103 904 2 specs ; E Ankarana Reserve , talus and rocks below 20 m tsingy cliff ; 12.8609° S , 49.2267° E ; alt. 294 m ; dry deciduous forest on limestone ; 14 Apr. 2007 ; Ken Emberton leg.; coll. no. MBI-2068 ; FMNH-INS 000 103 911 1 spec. ; NE Ankarana Reserve , local limestone summit ; 12.8738° S , 49.2249° E ; alt. 315 m ; dry deciduous forest on limestone ; 14 Apr. 2007 ; Ken Emberton leg.; coll. no. MBI-2064; FMNH-INS 000 103 920 4 specs ; NE Ankarana Reserve , talus below tsingy visible from road ; 12.8617° S , 49.2266° E ; alt. 266 m ; dry deciduous forest on limestone ; 14 Apr. 2007 ; Ken Emberton leg.; coll. no. MBI-2062; FMNH-INS 000 103 922 2 spec. ; NE Analamerana, near summit ; 12.6833° S , 49.5333° E ; alt. 36 m ; litter-loam pockets in limestone brocks, dry deciduous forest on limestone ; 23 Apr. 2007 ; Ken Emberton leg.; coll. no. MBI-2072; FMNH-INS 000 103 929 3 specs ; NE Analamerana , 160 m downslope E from summit; 12.6833° S , 49.5333° E ; limestone outcrop, dry deciduous forest on limestone ; 25 Apr. 2007 ; Ken Emberton leg.; coll. no. MBI-2086 ; FMNH-INS 000 103 921 1 spec. ; NE Analamerana , on W bank of Antafiamantsina River, at waterfall, 3 rd rapid up ; 12.6833° S , 49.5333° E ; dry deciduous forest on limestone; 26 Apr. 2007 ; Ken Emberton leg.; coll. no. MBI-2099; FMNH-INS 000 103 903 7 specs ; NE Analamerana , small tsingy 180 m above river, ca 30 minute walk ; 12.6833° S , 49.5333° E ; dry deciduous forest on limestone ; 26 Apr. 2007 ; Ken Emberton leg.; coll. no. MBI-2094; FMNH-INS 000 103 917 2 specs ; N Cap d’Ambre , Pointe du Phoque ; 11.9698° S , 49.2845° E ; alt. 11 m ; spring-bottle tree stand, dry deciduous forest on limestone ; 4 May 2007 ; Ken Emberton leg.; coll. no. MBI-2108 ; FMNH-INS 000 103 910 2 specs ; NW of Antalaha, S of Mahadera, Mt. Ambohodrapaka ; 14.6805° S , 50.0325° E ; alt. 270 m ; partially cut rainforest on granite ; 29 Jul. 2007 ; Ken Emberton leg.; coll. no. MBI-2403; FMNH-INS 000 103 906 2 specs ; same collection data as for preceding; coll. no. MBI-2404; FMNH-INS 000 103 907 1 spec. ; NW of Antalaha, S of Mahadera, Mt Ambohodrapaka ; 14.6836° S , 50.031° E ; alt. 369 m ; partially cut rainforest on granite ; 29 Jul. 2007 ; Ken Emberton leg.; coll. no. MBI-2417; FMNH-INS 000 103 924 1 spec. ; NW of Antalaha ; 14. 4123° S , 50.1755° E ; alt. 1790 m ; 30 Jul. 2007 ; Ken Emberton leg.; coll. no. MBI-2439; FMNH-INS 000 103 905 2 specs ; NW of Antalaha, S of Mahadera, Mt Ambohodrapaka ; 14.6873° S , 50.0301° E ; alt. 499 m ; rainforest, on quartzite ; 30 Jul. 2007 ; Ken Emberton leg.; coll. no. MBI-2430; FMNH-INS 000 103 915 1 spec. ; NW of Antalaha, S of Mahadera, Mt. Ambohodrapaka ; 14.6872° S , 50.0293° E ; alt. 545 m ; 30 Jul. 2007 ; Ken Emberton leg.; coll. no. MBI-2434; FMNH- INS 000 103 918 1 spec. ; NW of Antalaha, SW of Andranofotsy, Mt Ambodivohitra ; 14.8157° S , 49.9634° E ; alt. 487 m ; rainforest on quartzite ; 10 Aug. 2007 ; Ken Emberton leg.; coll. no. MBI- 2459; FMNH-INS 000 103 913 2 specs ; NW of Antalaha, SW of Andranofotsy, Mt Ambodivohitra ; 14.8157° S , 49.9634° E ; alt. 487 m ; rainforest on quartzite ; 10 Aug. 2007 ; Ken Emberton leg.; coll. no. MBI-2459; FMNH-INS 000 103 919 2 specs ; NW of Antalaha, SW of Andranofotsy, Mt Ambodivohitra ; 14.8166° S , 49.9622° E ; alt. 575 m ; rainforest, pandanus, palms, sphagnum on quartzite ; 11 Aug. 2007 ; Ken Emberton leg.; coll. no. MBI-2472; FMNH-INS 000 103 912 1 spec. ; NW of Antalaha, S of Antsahanoro, riverside hill rainforest ; 14.8434° S , 50.1372° E ; alt. 82 m ; on quartzite; 15 Aug. 2007 ; Ken Emberton leg.; coll. no. MBI-2498; FMNH-INS 000 103 916 1 spec. ; NW of Antalaha, S of Antsahanoro ; 14.8445° S , 50.1359° E ; alt. 131 m ; riverside hill rainforest on quartzite ; 15 Aug. 2007 ; Ken Emberton leg.; coll. no. MBI-2512; FMNH-INS 000 103 934 2 specs ; Mt Bobanora , about 40 km W of Vohemar ; 13.2141° S , 49.7574° E ; alt. 320 m ; viny rainforest on basalt talus ; 26 Aug. 2007 ; Ken Emberton leg.; coll. no. MBI-2550 ; FMNH-INS 000 103 914 1 spec. ; Manongorivo Reserve , slope high above Berato ; 14.0366° S , 48.2882° E ; alt. 897 m ; rainforest on metamorphic boulders ; 17 Oct. 2007 ; Ken Emberton leg.; coll. no. MBI-2611; FMNH-INS 000 103 933 2 specs ; Manongorivo Reserve , Monongorivo River junction with small tributary ; 13.9846° S , 48.2891° E ; alt. 321 m ; rainforest on granite ; 20 Oct. 2007 ; Ken Emberton leg.; coll. no. MBI-2619 ; FMNH-INS 000 103 941 5 specs ; Manongorivo Reserve , riverside flat area below cliffs , Monongorivo River ; 13.9818° S , 48.2858° E ; alt. 281 m ; rainforest on granite ; 20 Oct. 2007 ; Ken Emberton leg.; coll. no. MBI-2620; FMNH-INS 000 103 943 . – Mahajanga 1 spec. ; Namoroka Reserve , southern valley on NE side of tsingy ; 16.469° S , 45.3647° E ; alt. 142 m ; dry deciduous forest on limestone ; 25 Sep. 2007 ; Ken Emberton leg.; coll. no. MBI-2567; FMNH-INS 000 103 936 2 specs ; Namoroka Reserve , southern isolated tsingy canyon ; 16.4674° S , 45.3806° E ; alt. 432 feet ; dry deciduous forest on limestone ; 27 Sep. 2007 ; Ken Emberton leg.; coll. no. MBI-2574 ; FMNH-INS 000 103 928 2 specs ; Namaroka Reserve , southern isolated tsingy ; 16.4871° S , 45.3371° E ; alt. 435 m ; dry deciduous forest on limestone ; 27 Sep. 2007 ; Ken Emberton leg.; coll. no. MBI-2572; FMNH-INS 000 103 927 9 spec. ; S Namaroka National Park , low tsingy baobobs ; 16.4565° S , 45.4011° E ; alt. 115 m ; dry deciduous forest on limestone ; 29 Sep. 2007 ; Ken Emberton leg.; coll. no. MBI-2578; FMNH-INS 000 103 923 1 spec. ; Anjajavy , isolated steep limestone hill near Antsangabe ; 15.0643° S , 47.2373° E ; alt. 30 m ; dry deciduous forest on limestone ; 8 Oct. 2007 ; Ken Emberton leg.; coll. no. MBI-2592; FMNH-INS 000 104 000 1 spec. ; Anjajavi , side of long tsingy hill near dry mudflat ; 15.026° S , 47.2712° E ; alt. 22 m ; dry deciduous forest on limestone; 11 Oct. 2007 ; Ken Emberton leg.; coll. no. MBI-2595; FMNH-INS 000 103 940 . – Toamasina 1 spec. ; Andriantanely Massif ; 18.6907° S , 48.8063° E ; alt. 589 m ; rainforest; 19 May 2008 ; Ken Emberton leg.; coll. no. MBI-2665; FMNH-INS 000 103 932 1 spec. ; Andriantantely Massif , streamside cliff base ; 18.6997° S , 48.806° E ; alt. 301 m ; in rainforest, on granite ; 22 May 2008 ; Ken Emberton leg.; coll. no. MBI-2693; FMNH-INS 000 103 930 1 spec. ; Makira Reserve , W of Andaparaty ; 15.2302° S , 49.5975° E ; alt. 430 m ; rainforest; 23 Nov. 2008 ; Ken Emberton leg.; coll. no. MBI-3152; FMNH-INS 000 103 939 2 specs ; N of Manona littoral forest ; 16.6028° S , 49.8037° E ; alt. 23 m ; 2 Dec. 2008 ; Ken Emberton leg.; coll. no. MBI-3158; FMNH-INS 000 103 937 . – Toliara 2 specs ; W of Ehasity ; 24.154° S , 47.1821° E ; alt. 347 m ; rainforest ; 18 Aug. 2008 ; Ken Emberton leg.; coll. no. MBI-2966; FMNH-INS 000 103 938 1 spec. ; Mount Ivohibe ; 24.5607° S , 47.2065° E ; alt. 457 m ; rainforest ; 31 Aug. 2008 ; Ken Emberton leg.; coll. no. MBI-2900; FMNH-INS 000 103 931 1 spec. ; W of Ehasity rainforest ; 24.1514° S , 47.178° E ; alt. 479 m ; 18 Sep. 2008 ; Ken Emberton leg.; coll. no. MBI-2964 ; FMNH-INS 000 103 942 .