Seven new giant pill-millipede species and numerous new records of the genus Zoosphaerium from Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae) Author Wesener, Thomas 86DEA7CD-988C-43EC-B9D6-C51000595B47 Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Section Myriapoda, Adenauerallee 160, D- 53113 Bonn, Germany. t.wesener@leibniz-zfmk.de Author Sagorny, Christina 9C89C1B7-897A-426E-8FD4-C747DF004C85 Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Section Myriapoda, Adenauerallee 160, D- 53113 Bonn, Germany. & University of Bonn, Institute of Evolutionary Biology and Ecology, D- 53121 Bonn, Germany. csagorny@evolution.uni-bonn.de text European Journal of Taxonomy 2021 2021-07-06 758 1 1 48 http://dx.doi.org/10.5852/ejt.2021.758.1423 journal article 10.5852/ejt.2021.758.1423 de9c0c06-0e7d-44a6-a582-d41a346233f6 2118-9773 5075824 01BBC12C-E715-4393-A9F6-6EA85CB1289F Genus Zoosphaerium Pocock, 1895 Zoosphaerium Pocock, 1895: 410 . Globotherium Brolemann, 1922: 230 (formal synonymization in Wesener & Wägele 2008: 9 ). Heligmasoma Chamberlin, 1921: 58 (formal synonymization in Wesener & Wägele 2008: 9 ). See Wesener (2016) for a summary; five additional species have been described since then ( Sagorny & Wesener 2017 ; Wesener & Anilkumar 2020 ). Type species Sphaerotherium neptunus Butler, 1872 . Other species included 66 and 11 nomina dubia (including the species described below). Fig. 1. Maximum likelihood tree inferred from the COI dataset with 1000 bootstrap pseudoreplicates implementing the GTR+I+ G model. Colors representing newly described species of Zoosphaerium : orange = Z. nigrum sp. nov. ; green = Z. silens sp. nov. ; red = Z. ambatovaky sp. nov. ; yellow = Z. beanka sp. nov. ; purple = Z. voahangy sp. nov. ; blue = Z. masoala sp. nov. Round-cornered rectangles indicate well-supported sister group relationships. Distribution Endemic to Madagascar . Remarks The diverse genus Zoosphaerium has been divided into six species-groups ( Wesener & Wägele 2008 ; Wesener 2009). Six additional species are currently not assignable to any group ( Sagorny & Wesener 2017 ). Four species are only known from females and an additional 11 species are nomina dubia. A key to the species-groups has been published elsewhere (Wesener 2009).