Seven new giant pill-millipede species and numerous new records of the genus Zoosphaerium from Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae)
Author
Wesener, Thomas
86DEA7CD-988C-43EC-B9D6-C51000595B47
Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Section Myriapoda, Adenauerallee 160, D- 53113 Bonn, Germany.
t.wesener@leibniz-zfmk.de
Author
Sagorny, Christina
9C89C1B7-897A-426E-8FD4-C747DF004C85
Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Section Myriapoda, Adenauerallee 160, D- 53113 Bonn, Germany. & University of Bonn, Institute of Evolutionary Biology and Ecology, D- 53121 Bonn, Germany.
csagorny@evolution.uni-bonn.de
text
European Journal of Taxonomy
2021
2021-07-06
758
1
1
48
http://dx.doi.org/10.5852/ejt.2021.758.1423
journal article
10.5852/ejt.2021.758.1423
de9c0c06-0e7d-44a6-a582-d41a346233f6
2118-9773
5075824
01BBC12C-E715-4393-A9F6-6EA85CB1289F
Genus
Zoosphaerium
Pocock, 1895
Zoosphaerium
Pocock, 1895: 410
.
Globotherium
Brolemann, 1922: 230
(formal synonymization in
Wesener & Wägele 2008: 9
).
Heligmasoma
Chamberlin, 1921: 58
(formal synonymization in
Wesener & Wägele 2008: 9
).
See
Wesener (2016)
for a summary; five additional species have been described since then (
Sagorny & Wesener 2017
;
Wesener & Anilkumar 2020
).
Type
species
Sphaerotherium neptunus
Butler, 1872
.
Other species included
66 and 11 nomina dubia (including the species described below).
Fig. 1.
Maximum likelihood tree inferred from the COI dataset with 1000 bootstrap pseudoreplicates implementing the GTR+I+ G model. Colors representing newly described species of
Zoosphaerium
: orange =
Z. nigrum
sp. nov.
; green =
Z. silens
sp. nov.
; red =
Z. ambatovaky
sp. nov.
; yellow =
Z. beanka
sp. nov.
; purple =
Z. voahangy
sp. nov.
; blue =
Z. masoala
sp. nov.
Round-cornered rectangles indicate well-supported sister group relationships.
Distribution
Endemic to
Madagascar
.
Remarks
The diverse genus
Zoosphaerium
has been divided into six species-groups (
Wesener & Wägele 2008
; Wesener 2009). Six additional species are currently not assignable to any group (
Sagorny & Wesener 2017
). Four species are only known from females and an additional 11 species are nomina dubia. A key to the species-groups has been published elsewhere (Wesener 2009).