<document id="11A083AA95DD72C62CCFAC222DACE6CA" ID-CLB-Dataset="6818" ID-DOI="10.5281/zenodo.4760479" ID-GBIF-Dataset="9417b10b-5d16-4f7a-9d9d-86d80f05c38b" ID-ISSN="1854-0392" ID-Zenodo-Dep="4760479" ID-ZooBank="79B7368A-29DC-44C2-B9AD-5AFA006FBC7F" IM.materialsCitations_requiresApprovalFor="plazi" IM.metadata_requiresApprovalFor="plazi" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1620963140635" checkinUser="felipe" docAuthor="Stark, Bill P." docDate="2014" docId="03CD878C8238FFBB0F9D5E6F0E66FA13" docLanguage="en" docName="Illiesia.10.2.6-16.pdf" docOrigin="Illiesia 10 (2)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Anacroneuria tempisquito Stark 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="14" masterDocId="FFF4FFF4823FFFB30C2C592C0C25FFA1" masterDocTitle="Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species" masterLastPageNumber="16" masterPageNumber="6" pageNumber="13" updateTime="1698939657618" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="335D761ED36B301BE0243E6847550259">Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species</mods:title>
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<mods:title id="E48F6DE7143013D39B5D52B3357E0E50">Illiesia</mods:title>
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<treatment id="03CD878C8238FFBB0F9D5E6F0E66FA13" ID-DOI="http://doi.org/10.5281/zenodo.4760087" ID-GBIF-Taxon="182137841" ID-Zenodo-Dep="4760087" LSID="urn:lsid:plazi:treatment:03CD878C8238FFBB0F9D5E6F0E66FA13" httpUri="http://treatment.plazi.org/id/03CD878C8238FFBB0F9D5E6F0E66FA13" lastPageId="8" lastPageNumber="14" pageId="7" pageNumber="13">
<subSubSection id="C37E65118238FFB40F9D5E6F0910F8FA" box="[945,1333,1859,1883]" pageId="7" pageNumber="13" type="nomenclature">
<paragraph id="8BDB369A8238FFB40F9D5E6F0910F8FA" blockId="7.[945,1333,1859,1883]" box="[945,1333,1859,1883]" pageId="7" pageNumber="13">
<heading id="D09381F68238FFB40F9D5E6F0910F8FA" bold="true" box="[945,1333,1859,1883]" fontSize="10" level="2" pageId="7" pageNumber="13" reason="2">
<emphasis id="B910EA888238FFB40F9D5E6F0910F8FA" bold="true" box="[945,1333,1859,1883]" pageId="7" pageNumber="13">
<taxonomicName id="4C644D198238FFB40F9D5E6F08C2F8FA" ID-CoL="DC67" authority="Stark, 2014" authorityName="Stark" authorityYear="2014" box="[945,1255,1859,1883]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="13" phylum="Arthropoda" rank="species" species="tempisquito" status="sp. nov.">
<emphasis id="B910EA888238FFB40F9D5E6F08C2F8FA" bold="true" box="[945,1255,1859,1883]" italics="true" pageId="7" pageNumber="13">Anacroneuria tempisquito</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22357F38238FFB408D85E6F0910F8FA" box="[1268,1333,1859,1883]" pageId="7" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37E65118238FFB408005E4B089FF8DE" box="[1068,1210,1895,1919]" pageId="7" pageNumber="13" type="description">
<paragraph id="8BDB369A8238FFB408005E4B089FF8DE" blockId="7.[1068,1210,1895,1919]" box="[1068,1210,1895,1919]" pageId="7" pageNumber="13">
(
<figureCitation id="135F2A1F8238FFB408185E4B0897F8DE" box="[1076,1202,1895,1919]" captionStart="Figs" captionStartId="7.[189,241,1646,1670]" captionTargetBox="[352,1230,215,1547]" captionTargetId="figure-124@7.[325,1308,164,1558]" captionText="Figs. 11-15. Anacroneuria tempisquito sp.n. male. 11. Head and pronotum. 12. Sternum 9 with hammer. 13. Aedeagus, ventral. 14. Aedeagus, lateral. 15. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4765168" httpUri="https://zenodo.org/record/4765168/files/figure.png" pageId="7" pageNumber="13">Figs. 11-15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37E65118237FFBB0C9159CD0EC5FEE1" pageId="8" pageNumber="14" type="materials_examined">
<paragraph id="8BDB369A8237FFBB0C9159CD0EC5FEE1" blockId="8.[189,798,225,1459]" pageId="8" pageNumber="14">
<emphasis id="B910EA888237FFBB0C9159CD0D9CFF58" bold="true" box="[189,441,225,249]" pageId="8" pageNumber="14">Material examined.</emphasis>
<materialsCitation id="3B0C3CC78237FFBB0DFC59CD0EC5FEE1" ID-GBIF-Occurrence="3111908331" collectingDate="1990-05-21" collectingMethod="light trap" collectionCode="USNM" country="Costa Rica" location="Rio Tempisquito" municipality="Estacion Biologica Maritza" pageId="8" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste Province" typeStatus="holotype">
<typeStatus id="54DF88388237FFBB0DFC59CD0E65FF58" box="[464,576,225,249]" pageId="8" pageNumber="14" type="holotype">Holotype</typeStatus>
<specimenCount id="9D62FD138237FFBB0E7A59CD0E4FFF58" box="[598,618,225,249]" pageId="8" pageNumber="14" type="male">♂</specimenCount>
, 
<emphasis id="B910EA888237FFBB0EAB59CD0F33FF58" bold="true" box="[647,790,225,249]" pageId="8" pageNumber="14">
<collectingCountry id="F373760A8237FFBB0EAB59CD0F33FF58" box="[647,790,225,249]" name="Costa Rica" pageId="8" pageNumber="14">Costa Rica</collectingCountry>
</emphasis>
, 
<collectingRegion id="49A0F8788237FFBB0C9158290D96FEBC" box="[189,435,261,285]" country="Costa Rica" name="Guanacaste" pageId="8" pageNumber="14">Guanacaste Province</collectingRegion>
, 
<collectingMunicipality id="6BBFACE08237FFBB0D9058290EC8FEBC" box="[444,749,261,285]" pageId="8" pageNumber="14">Estación Biológica Maritza</collectingMunicipality>
, 
<location id="8EBB60418237FFBB0EDA58290D70FEE1" LSID="urn:lsid:plazi:treatment:03CD878C8238FFBB0F9D5E6F0E66FA13:8EBB60418237FFBB0EDA58290D70FEE1" country="Costa Rica" municipality="Estacion Biologica Maritza" name="Rio Tempisquito" pageId="8" pageNumber="14" stateProvince="Guanacaste Province">Rio Tempisquito</location>
, 
<date id="FFDA105A8237FFBB0D4C58040DD3FEE1" box="[352,502,296,321]" pageId="8" pageNumber="14" value="1990-05-21">
<collectingDate id="EF9EE9B28237FFBB0D4C58040DD3FEE1" box="[352,502,296,321]" pageId="8" pageNumber="14" value="1990-05-21">21 May 1990</collectingDate>
</date>
, 
<collectingMethod id="52254E8D8237FFBB0E2D58040E4BFEE1" box="[513,622,296,320]" pageId="8" pageNumber="14">light trap</collectingMethod>
(
<collectionCode id="ED75AE5F8237FFBB0E5158040EF1FEE1" box="[637,724,296,320]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="14" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37E65118237FFBB0C9158670DA9FC61" pageId="8" pageNumber="14" type="description">
<paragraph id="8BDB369A8237FFBB0C9158670EB2FDFD" blockId="8.[189,798,225,1459]" pageId="8" pageNumber="14">
<emphasis id="B910EA888237FFBB0C9158670D49FEC2" bold="true" box="[189,364,331,355]" pageId="8" pageNumber="14">Adult habitus.</emphasis>
Head yellow except for diffuse brown area forward of ocelli and dark brown lappets and antennae (
<figureCitation id="135F2A1F8237FFBB0D1B58BF0DADFE0A" box="[311,392,403,427]" captionStart="Figs" captionStartId="7.[189,241,1646,1670]" captionTargetBox="[352,1230,215,1547]" captionTargetId="figure-124@7.[325,1308,164,1558]" captionText="Figs. 11-15. Anacroneuria tempisquito sp.n. male. 11. Head and pronotum. 12. Sternum 9 with hammer. 13. Aedeagus, ventral. 14. Aedeagus, lateral. 15. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4765168" httpUri="https://zenodo.org/record/4765168/files/figure.png" pageId="8" pageNumber="14">Fig. 11</figureCitation>
). Pronotum with pale mesal band; dark lateral bands cover most of disc. Wing membrane transparent with pale amber tint; veins darker except for pale costa. Femora pale except for narrow black apical band; tibiae dark in proximal fourth and with a small distal dark spot.
</paragraph>
<paragraph id="8BDB369A8237FFBB0C915B440E7AFCD8" blockId="8.[189,798,225,1459]" pageId="8" pageNumber="14">
<emphasis id="B910EA888237FFBB0C915B440D27FD21" bold="true" box="[189,258,616,640]" pageId="8" pageNumber="14">Male.</emphasis>
Forewing length 
<quantity id="4C9C9B7F8237FFBB0DE15B440E38FD21" box="[461,541,616,640]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="14" unit="mm" value="10.0">10 mm</quantity>
. Hammer reduced to a small, transparent, hairless disc (
<figureCitation id="135F2A1F8237FFBB0E695BA00EBDFD05" box="[581,664,652,676]" captionStart="Figs" captionStartId="7.[189,241,1646,1670]" captionTargetBox="[352,1230,215,1547]" captionTargetId="figure-124@7.[325,1308,164,1558]" captionText="Figs. 11-15. Anacroneuria tempisquito sp.n. male. 11. Head and pronotum. 12. Sternum 9 with hammer. 13. Aedeagus, ventral. 14. Aedeagus, lateral. 15. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4765168" httpUri="https://zenodo.org/record/4765168/files/figure.png" pageId="8" pageNumber="14">Fig. 12</figureCitation>
). Aedeagal apex pointed in ventral aspect and bearing a prominent pair of ventral, membranous lobes (
<figureCitation id="135F2A1F8237FFBB0EDC5BFF0CFFFCAE" captionStart="Figs" captionStartId="7.[189,241,1646,1670]" captionTargetBox="[352,1230,215,1547]" captionTargetId="figure-124@7.[325,1308,164,1558]" captionText="Figs. 11-15. Anacroneuria tempisquito sp.n. male. 11. Head and pronotum. 12. Sternum 9 with hammer. 13. Aedeagus, ventral. 14. Aedeagus, lateral. 15. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4765168" httpUri="https://zenodo.org/record/4765168/files/figure.png" pageId="8" pageNumber="14">Fig. 13</figureCitation>
); apex angled sharply ventrad in lateral aspect (
<figureCitation id="135F2A1F8237FFBB0CE95A360D3CFC93" box="[197,281,794,818]" captionStart="Figs" captionStartId="7.[189,241,1646,1670]" captionTargetBox="[352,1230,215,1547]" captionTargetId="figure-124@7.[325,1308,164,1558]" captionText="Figs. 11-15. Anacroneuria tempisquito sp.n. male. 11. Head and pronotum. 12. Sternum 9 with hammer. 13. Aedeagus, ventral. 14. Aedeagus, lateral. 15. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4765168" httpUri="https://zenodo.org/record/4765168/files/figure.png" pageId="8" pageNumber="14">Fig. 14</figureCitation>
), and bearing an obscure V-shaped dorsal keel composed of divergent, broken lines (
<figureCitation id="135F2A1F8237FFBB0E915A120F2AFCF4" box="[701,783,829,854]" captionStart="Figs" captionStartId="7.[189,241,1646,1670]" captionTargetBox="[352,1230,215,1547]" captionTargetId="figure-124@7.[325,1308,164,1558]" captionText="Figs. 11-15. Anacroneuria tempisquito sp.n. male. 11. Head and pronotum. 12. Sternum 9 with hammer. 13. Aedeagus, ventral. 14. Aedeagus, lateral. 15. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4765168" httpUri="https://zenodo.org/record/4765168/files/figure.png" pageId="8" pageNumber="14">Fig. 15</figureCitation>
). Hooks slender with long acute tips.
</paragraph>
<paragraph id="8BDB369A8237FFBB0C915AA90D85FC3C" blockId="8.[189,798,225,1459]" box="[189,416,901,925]" pageId="8" pageNumber="14">
<emphasis id="B910EA888237FFBB0C915AA90D3EFC3C" bold="true" box="[189,283,901,925]" pageId="8" pageNumber="14">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BDB369A8237FFBB0C915A850DA9FC61" blockId="8.[189,798,225,1459]" box="[189,396,936,961]" pageId="8" pageNumber="14">
<emphasis id="B910EA888237FFBB0C915A850D2DFC60" bold="true" box="[189,264,937,961]" pageId="8" pageNumber="14">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C37E65118237FFBB0C915AE00EF6FBA9" pageId="8" pageNumber="14" type="etymology">
<paragraph id="8BDB369A8237FFBB0C915AE00EF6FBA9" blockId="8.[189,798,225,1459]" pageId="8" pageNumber="14">
<emphasis id="B910EA888237FFBB0C915AE00D62FC45" bold="true" box="[189,327,972,996]" pageId="8" pageNumber="14">Etymology.</emphasis>
The species name, based on Rio Tempisquito, is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C37E65118237FFBB0C915D3F0E66FA13" pageId="8" pageNumber="14" type="diagnosis">
<paragraph id="8BDB369A8237FFBB0C915D3F0E66FA13" blockId="8.[189,798,225,1459]" pageId="8" pageNumber="14">
<emphasis id="B910EA888237FFBB0C915D3F0D1AFB8D" bold="true" box="[189,319,1043,1068]" pageId="8" pageNumber="14">Diagnosis.</emphasis>
This species does not appear to be closely related to others known from Mesoamerica, but is more similar to the Ecuadorian species, 
<taxonomicName id="4C644D198237FFBB0EED5D770DEFFB36" authority="Zuniga &amp; Vimos, 2006" authorityName="Zuniga &amp; Vimos" authorityYear="2006" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="14" phylum="Arthropoda" rank="species" species="cajas">
<emphasis id="B910EA888237FFBB0EED5D770F38FBD2" box="[705,797,1115,1139]" italics="true" pageId="8" pageNumber="14">A. cajas</emphasis>
<bibRefCitation id="EFF54B6B8237FFBB0C915D530DEFFB36" author="Zuniga, M. C. &amp; B. P. Stark &amp; J. J. Vasconez &amp; F. Bersosa &amp; L. D. Vimos" box="[189,458,1151,1175]" pageId="8" pageNumber="14" pagination="45 - 57" refId="ref5785" refString="Zuniga, M. C., B. P. Stark, J. J. Vasconez, F. Bersosa, &amp; L. D. Vimos. 2006. Colombian and Ecuadorian Anacroneuria (Plecoptera: Perlidae): seven new species, records and life stages. Studies on Neotropical Fauna and Environment, 41: 45 - 57." type="journal article" year="2006">Zúñiga &amp; Vimos, 2006</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFF54B6B8237FFBB0DF65D530E97FB36" author="Zuniga, M. C. &amp; B. P. Stark &amp; J. J. Vasconez &amp; F. Bersosa &amp; L. D. Vimos" box="[474,690,1151,1175]" pageId="8" pageNumber="14" pagination="45 - 57" refId="ref5785" refString="Zuniga, M. C., B. P. Stark, J. J. Vasconez, F. Bersosa, &amp; L. D. Vimos. 2006. Colombian and Ecuadorian Anacroneuria (Plecoptera: Perlidae): seven new species, records and life stages. Studies on Neotropical Fauna and Environment, 41: 45 - 57." type="journal article" year="2006">Zúñiga et al. 2006</bibRefCitation>
) and the Colombian species 
<taxonomicName id="4C644D198237FFBB0D825D8F0F38FB1B" authority="Stark &amp; Zuniga, 2007" authorityName="Stark &amp; Zuniga" authorityYear="2007" box="[430,797,1186,1210]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="14" phylum="Arthropoda" rank="species" species="otun">
<emphasis id="B910EA888237FFBB0D825D8F0E23FB1B" box="[430,518,1187,1210]" italics="true" pageId="8" pageNumber="14">A. otun</emphasis>
Stark &amp; 
<bibRefCitation id="EFF54B6B8237FFBB0E535D8E0F38FB1B" author="Zuniga, M. C. &amp; B. P. Stark &amp; W. Cardona &amp; O. E. Ortega" box="[639,797,1186,1210]" pageId="8" pageNumber="14" pagination="127 - 149" refId="ref5849" refString="Zuniga, M. C., B. P. Stark, W. Cardona, C. Tamaris- Turizo, &amp; O. E. Ortega. 2007. Additions to the Colombian Anacroneuria fauna (Plecoptera: Perlidae) with descriptions of seven new species. Illiesia, 3: 127 - 149." type="journal article" year="2007">Zúñiga, 2007</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFF54B6B8237FFBB0CE95DEA0DB0FB7F" author="Zuniga, M. C. &amp; B. P. Stark &amp; W. Cardona &amp; O. E. Ortega" box="[197,405,1222,1246]" pageId="8" pageNumber="14" pagination="127 - 149" refId="ref5849" refString="Zuniga, M. C., B. P. Stark, W. Cardona, C. Tamaris- Turizo, &amp; O. E. Ortega. 2007. Additions to the Colombian Anacroneuria fauna (Plecoptera: Perlidae) with descriptions of seven new species. Illiesia, 3: 127 - 149." type="journal article" year="2007">Zúñiga et al. 2007</bibRefCitation>
). The new species is distinguished from both on the basis of pigment pattern, the prominent ventral membranous lobes, the dorsal keel, and the obscure hammer. In 
<bibRefCitation id="EFF54B6B8237FFBB0E4C5C1C0ECBFAE9" author="Stark, B. P." box="[608,750,1328,1352]" pageId="8" pageNumber="14" pagination="551 - 603" refId="ref5572" refString="Stark, B. P. 1998. The Anacroneuria of Costa Rica and Panama (Insecta: Plecoptera: Perlidae). Proceedings of the Biological Society of Washington, 111: 551 - 603." type="journal article" year="1998">Stark (1998)</bibRefCitation>
the new species will key to couplet 9 where a choice of 
<taxonomicName id="4C644D198237FFBB0C915C540D0BFA2E" authorityName="Stark" authorityYear="1998" box="[189,302,1399,1423]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="14" phylum="Arthropoda" rank="species" species="varilla">
<emphasis id="B910EA888237FFBB0C915C540D0BFA2E" box="[189,302,1399,1423]" italics="true" pageId="8" pageNumber="14">A. varilla</emphasis>
</taxonomicName>
could be forced, despite conspicuous differences in aedeagal structure.
</paragraph>
</subSubSection>
</treatment>
</document>