New species, new locality records and descriptions of calling songs of Phaneropterinae from Sabah Author Tan, Ming Kai Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore Author Damit, Dayang Fazrinah Binti Awg Forest Research Centre (Sepilok), Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia Author Japir, Razy Forest Research Centre (Sepilok), Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia Author Chung, Arthur Y. C. Forest Research Centre (Sepilok), Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia Author Robillard, Tony Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, SU, EPHE-PSL, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France text Zootaxa 2025 2025-03-17 5604 4 505 528 https://doi.org/10.11646/zootaxa.5604.4.4 journal article 309256 10.11646/zootaxa.5604.4.4 625b53e9-c749-487c-be35-667794003c3a 1175-5326 15214059 43CDF92A-7B39-4926-8FC0-EA813B85DF0E Tapiena ? incisa Karny, 1923 ( Figs 2G , 13–15 ) Tapiena incisa Karny, 1923: 141 — Tan et al. , 2015: 42 , 43 (key); Tan et al. , 2019: 328 (wrong identification); Tan et al. , 2024a: 481 . New material examined. EAST MALAYSIA ; Sabah State • 2♂ ; Sinipung Hill , near Long Pasia and Meligan ; N4.44538 E115.71494 , 1556.0± 10.1 m .a.s.l.; 21 November 2024 , 21h30; attracted to light trap ; coll. M.K. Tan ; SBH.24.205, 206 ( ZRC ) • 2♂ ; Meligan (P28 a.k.a. pt 2); N4.57619 E115.73923 , 1266.0± 6.8 m .a.s.l.; 23 November 2024 , 22h00; attracted to light trap on the road; coll. M.K. Tan ; SBH.24.268, 269 ( Fig. 13 ) ( ZRC ) . FIGURE 13. Tapiena ? incisa Karny, 1923 when alive, in captivity. FIGURE 14. Tapiena ? incisa Karny, 1923 : head, pronotum and anterior parts of tegmina in dorsal (A), lateral (B) and anterior (C) views; stridulatory area of the left tegmen in ventral view (D); abdominal apex in dorsal (E) and lateral (F, G) views. Scale bars: 1 mm. Remarks. Our specimens resemble those collected from Mount Trus Madi by Tan et al. (2024a) and northwestern Kinabalu ( Muhammad & Tan, 2025 ) in the important species characters, including the shapes of the male cercus, tenth abdominal tergite and subgenital plate, as well as the shape of the stridulatory areas ( Fig. 14 ). Our specimens also resemble Tapiena bullata Karny, 1923 from Malay Peninsula, particularly in the shape of the apex of their cercus. Type locality. EAST MALAYSIA : Sarawak : Baram River , Gunong Tamabo . Calling song ( Fig. 15 ). The calling song is an isolated echeme made up of an average of 11–12 closely-spaced syllables. The average echeme duration is 0.70± 0.10 s (0.58– 0.79 s ). Within each echeme, the amplitudes of the syllables increase to a maximum; the average syllable duration is 13.7±1.4 ms (12.2–15.3 ms) and the average down time (silent interval between the two syllables) is 48.6±7.1 ms (40.7–54.5 ms). The average period of syllable is 62.3±5.8 ms (55.7–67.4 ms). The frequency spectrum is broadband and is not symmetrical and has a peak frequency of 13.82±0.83 kHz (13.41–14.16 kHz).