<document id="D69AE0546C1D88CB6496EF667E48EED1" ID-DOI="10.5281/zenodo.10114352" ID-ISSN="0253-116X" ID-Zenodo-Dep="10114352" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630359673913" checkinUser="valdenar" docAuthor="Assing, V." docDate="2007" docId="03DA3C3AFFF8FFA71F06FE31FD3CFD1A" docLanguage="en" docName="LinzerbiolBeitr.2007.39.2.761-775.pdf" docOrigin="Linzer biologische Beiträge 39 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Paratyphlus espunae Assing, 2007, nov.sp." docType="treatment" docVersion="4" lastPageNumber="765" masterDocId="FFE34442FFFBFFA31E52FF9CFFECFF9F" masterDocTitle="Four new species and additional records of Staphylinidae from Spain (Insecta: Coleoptera)" masterLastPageNumber="775" masterPageNumber="761" pageNumber="764" updateTime="1700278658990" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="082AE62002E4165EA719BD888A399B38">Four new species and additional records of Staphylinidae from Spain (Insecta: Coleoptera)</mods:title>
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<mods:namePart id="A2E6BCC5F68C7643BEC237BD2A98330C">Assing, V.</mods:namePart>
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<treatment id="03DA3C3AFFF8FFA71F06FE31FD3CFD1A" LSID="urn:lsid:plazi:treatment:03DA3C3AFFF8FFA71F06FE31FD3CFD1A" httpUri="http://treatment.plazi.org/id/03DA3C3AFFF8FFA71F06FE31FD3CFD1A" lastPageId="4" lastPageNumber="765" pageId="3" pageNumber="764">
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<taxonomicName id="4C73F6AFFFF8FFA01F06FE31FDC2FE5C" ID-CoL="4DHDT" authorityName="Assing" authorityYear="2007" box="[340,558,429,452]" class="Insecta" family="Staphylinidae" genus="Paratyphlus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="764" phylum="Arthropoda" rank="species" species="espunae" status="sp. nov.">
<emphasis id="B907513EFFF8FFA01F06FE31FDC2FE5C" bold="true" box="[340,558,429,452]" italics="true" pageId="3" pageNumber="764">Paratyphlus espunae</emphasis>
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<taxonomicNameLabel id="A234EC45FFF8FFA01C66FE32FD6DFE5B" box="[564,641,430,452]" pageId="3" pageNumber="764" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="134891A9FFF8FFA01CDCFE32FD1CFE5B" box="[654,752,430,452]" captionStart-0="Figs 1-9" captionStart-1="Fig" captionStartId-0="2.[340,380,1653,1673]" captionStartId-1="3.[340,373,1747,1767]" captionTargetBox-0="[343,1236,425,1618]" captionTargetBox-1="[340,1254,1155,1735]" captionTargetId-0="figure-10@2.[340,1253,689,1167]" captionTargetId-1="figure-325@3.[340,1254,1155,1735]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs 1-9: Paratyphlus espunae nov.sp.: (1) habitus; (2) forebody; (3) male tergite VIII; (4) male sternite VIII; (5-7) aedeagus in lateral view; (8) female tergite VIII; (9) female sternite VIII. Scale bars: 1-2: 0.2 mm; 3-9: 0.1 mm." captionText-1="Fig. 10: Type locality of Paratyphlus espunae nov.sp." figureDoi-0="http://doi.org/10.5281/zenodo.10114354" figureDoi-1="http://doi.org/10.5281/zenodo.10114356" httpUri-0="https://zenodo.org/record/10114354/files/figure.png" httpUri-1="https://zenodo.org/record/10114356/files/figure.png" pageId="3" pageNumber="764">Figs 1-10</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BCC8D2CFFF8FFA01F06FE41FD7EFDCE" blockId="3.[340,1248,477,593]" pageId="3" pageNumber="764">
<materialsCitation id="3B1B8771FFF8FFA01F06FE41FB4CFDBE" collectingDate="2007-03-29" collectionCode="OOLL" collectorName="C. Andujar" determinerName="V. Assing" elevation="1100" latitude="37.888336" location="Prado Mayor" longLatPrecision="19" longitude="1.5677778" municipality="Sierra de Espuna" pageId="3" pageNumber="764" specimenCount="6" stateProvince="Murcia" typeStatus="holotype">
H o l o t y p e: E - 
<collectingRegion id="49B743CEFFF8FFA01C6AFE41FD9BFE6E" box="[568,631,477,497]" country="Spain" name="Murcia" pageId="3" pageNumber="764">Murcia</collectingRegion>
[12], 
<collectingMunicipality id="6BA81756FFF8FFA01CE3FE41FCA1FE6E" box="[689,845,477,497]" pageId="3" pageNumber="764">Sierra de Espuña</collectingMunicipality>
, 
<location id="8EACDBF7FFF8FFA01D0BFE41FC3DFE6E" LSID="urn:lsid:plazi:treatment:03DA3C3AFFF8FFA71F06FE31FD3CFD1A:8EACDBF7FFF8FFA01D0BFE41FC3DFE6E" box="[857,977,477,497]" latitude="37.888336" longLatPrecision="19" longitude="1.5677778" municipality="Sierra de Espuna" name="Prado Mayor" pageId="3" pageNumber="764" stateProvince="Murcia">Prado Mayor</location>
, pasture, 
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, 
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, 
<geoCoordinate id="EE47EBEBFFF8FFA01FD3FE69FE32FD96" box="[385,478,501,521]" degrees="01" direction="east" minutes="34" orientation="longitude" pageId="3" pageNumber="764" precision="15" seconds="04" value="1.5677778">01°34'04E</geoCoordinate>
, 
<date id="FFCDABECFFF8FFA01FBAFE69FDA6FD96" box="[488,586,501,521]" pageId="3" pageNumber="764" value="2007-03-29">
<collectingDate id="EF895204FFF8FFA01FBAFE69FDA6FD96" box="[488,586,501,521]" pageId="3" pageNumber="764" value="2007-03-29">29.III.2007</collectingDate>
</date>
, 
<collectorName id="2686E8FAFFF8FFA01C06FE69FD5AFD96" box="[596,694,501,521]" pageId="3" pageNumber="764">C. Andújar</collectorName>
/ 
<typeStatus id="54C8338EFFF8FFA01C9AFE69FCCFFD96" box="[712,803,501,521]" pageId="3" pageNumber="764" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C73F6AFFFF8FFA01D10FE68FC15FD96" authorityName="Assing" authorityYear="2007" box="[834,1017,500,521]" class="Insecta" family="Staphylinidae" genus="Paratyphlus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="764" phylum="Arthropoda" rank="species" species="espunae" status="sp. nov.">
<emphasis id="B907513EFFF8FFA01D10FE68FC15FD96" box="[834,1017,500,521]" italics="true" pageId="3" pageNumber="764">Paratyphlus espunae</emphasis>
</taxonomicName>
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det. 
<determinerName id="7E1061E8FFF8FFA01A07FE69FB42FD96" box="[1109,1198,501,521]" pageId="3" pageNumber="764">V. Assing</determinerName>
2007 (cAss). P a r a t y p e s:7, 7, 
<specimenCount id="9D7546A5FFF8FFA01CB6FD91FCF7FDBE" box="[740,795,525,545]" pageId="3" pageNumber="764" type="generic">6 exs.</specimenCount>
: same data as holotype (
<collectionCode id="ED6215E9FFF8FFA01DBEFD91FBC6FDBE" box="[1004,1066,525,545]" pageId="3" pageNumber="764">OÖLL</collectionCode>
, cAnd, cAss)
</materialsCitation>
; 
<materialsCitation id="3B1B8771FFF8FFA01AFBFD91FD61FDCE" collectingDate="2007-03-29" collectorName="C. Andujar" elevation="1140" latitude="37.88639" location="Prado Mayor" longLatPrecision="19" longitude="1.5647222" municipality="Sierra de Espuna" pageId="3" pageNumber="764" specimenCount="1" stateProvince="Murcia" typeStatus="holotype">
1: E - 
<collectingRegion id="49B743CEFFF8FFA01FC4FDB9FE39FDA6" box="[406,469,549,569]" country="Spain" name="Murcia" pageId="3" pageNumber="764">Murcia</collectingRegion>
[13], 
<collectingMunicipality id="6BA81756FFF8FFA01C48FDB9FD53FDA6" box="[538,703,549,569]" pageId="3" pageNumber="764">Sierra de Espuña</collectingMunicipality>
, 
<location id="8EACDBF7FFF8FFA01C83FDB9FCA2FDA6" LSID="urn:lsid:plazi:treatment:03DA3C3AFFF8FFA71F06FE31FD3CFD1A:8EACDBF7FFF8FFA01C83FDB9FCA2FDA6" box="[721,846,549,569]" latitude="37.88639" longLatPrecision="19" longitude="1.5647222" municipality="Sierra de Espuna" name="Prado Mayor" pageId="3" pageNumber="764" stateProvince="Murcia">Prado Mayor</location>
, pasture, 
<quantity id="4C8B20C9FFF8FFA01DE2FDB9FC11FDA6" box="[944,1021,549,569]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="3" pageNumber="764" unit="m" value="1140.0">
<elevation id="005E6A1FFFF8FFA01DE2FDB9FC11FDA6" box="[944,1021,549,569]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="3" pageNumber="764" unit="m" value="1140.0">1140 m</elevation>
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, 
<geoCoordinate id="EE47EBEBFFF8FFA01A5CFDB9FB81FDA6" box="[1038,1133,549,569]" degrees="37" direction="north" minutes="53" orientation="latitude" pageId="3" pageNumber="764" precision="15" seconds="11" value="37.88639">37°53'11N</geoCoordinate>
, 
<geoCoordinate id="EE47EBEBFFF8FFA01A2DFDB9FB30FDA6" box="[1151,1244,549,569]" degrees="01" direction="east" minutes="33" orientation="longitude" pageId="3" pageNumber="764" precision="15" seconds="53" value="1.5647222">01°33'53E</geoCoordinate>
, 
<date id="FFCDABECFFF8FFA01FD3FDA1FE08FDCE" box="[385,484,573,593]" pageId="3" pageNumber="764" value="2007-03-29">
<collectingDate id="EF895204FFF8FFA01FD3FDA1FE08FDCE" box="[385,484,573,593]" pageId="3" pageNumber="764" value="2007-03-29">29.III.2007</collectingDate>
</date>
, 
<collectorName id="2686E8FAFFF8FFA01FBFFDA1FDA2FDCE" box="[493,590,573,593]" pageId="3" pageNumber="764">C. Andújar</collectorName>
(cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C369DEA7FFF8FFA71F06FDFDFD3CFD1A" lastPageId="4" lastPageNumber="765" pageId="3" pageNumber="764" type="description">
<paragraph id="8BCC8D2CFFF8FFA01F06FDFDFE90FD0C" blockId="3.[340,1247,608,659]" pageId="3" pageNumber="764">
D e s c r i p t i o n: 1.0- 
<quantity id="4C8B20C9FFF8FFA01C0BFDFCFD44FDE9" box="[601,680,608,630]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="764" unit="mm" value="1.3">1.3 mm</quantity>
(abdomen extended); habitus and forebody as in 
<figureCitation id="134891A9FFF8FFA01AE1FDFCFE9BFD0C" captionStart="Figs 1-9" captionStartId="2.[340,380,1653,1673]" captionTargetBox="[343,1236,425,1618]" captionTargetId="figure-10@2.[340,1253,689,1167]" captionTargetPageId="2" captionText="Figs 1-9: Paratyphlus espunae nov.sp.: (1) habitus; (2) forebody; (3) male tergite VIII; (4) male sternite VIII; (5-7) aedeagus in lateral view; (8) female tergite VIII; (9) female sternite VIII. Scale bars: 1-2: 0.2 mm; 3-9: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114354" httpUri="https://zenodo.org/record/10114354/files/figure.png" pageId="3" pageNumber="764">Figs 1-2</figureCitation>
.
</paragraph>
<paragraph id="8BCC8D2CFFF8FFA01F3BFD3FFD0CFC8F" blockId="3.[340,1247,674,784]" pageId="3" pageNumber="764">
: sternite VII with shallow median impression in posterior half; tergite VII weakly transverse and with truncate posterior margin (
<figureCitation id="134891A9FFF8FFA01D63FD23FC83FD4A" box="[817,879,703,725]" captionStart="Figs 1-9" captionStartId="2.[340,380,1653,1673]" captionTargetBox="[343,1236,425,1618]" captionTargetId="figure-10@2.[340,1253,689,1167]" captionTargetPageId="2" captionText="Figs 1-9: Paratyphlus espunae nov.sp.: (1) habitus; (2) forebody; (3) male tergite VIII; (4) male sternite VIII; (5-7) aedeagus in lateral view; (8) female tergite VIII; (9) female sternite VIII. Scale bars: 1-2: 0.2 mm; 3-9: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114354" httpUri="https://zenodo.org/record/10114354/files/figure.png" pageId="3" pageNumber="764">Fig. 3</figureCitation>
); sternite VIII with rather extensive impunctate median impression, posterior margin with excision of broadly triangular shape (
<figureCitation id="134891A9FFF8FFA01FC9FD66FE36FC8F" box="[411,474,762,784]" captionStart="Figs 1-9" captionStartId="2.[340,380,1653,1673]" captionTargetBox="[343,1236,425,1618]" captionTargetId="figure-10@2.[340,1253,689,1167]" captionTargetPageId="2" captionText="Figs 1-9: Paratyphlus espunae nov.sp.: (1) habitus; (2) forebody; (3) male tergite VIII; (4) male sternite VIII; (5-7) aedeagus in lateral view; (8) female tergite VIII; (9) female sternite VIII. Scale bars: 1-2: 0.2 mm; 3-9: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114354" httpUri="https://zenodo.org/record/10114354/files/figure.png" pageId="3" pageNumber="764">Fig. 4</figureCitation>
); aedeagus as in 
<figureCitation id="134891A9FFF8FFA01CD4FD66FD30FC8F" box="[646,732,762,784]" captionStart="Figs 1-9" captionStartId="2.[340,380,1653,1673]" captionTargetBox="[343,1236,425,1618]" captionTargetId="figure-10@2.[340,1253,689,1167]" captionTargetPageId="2" captionText="Figs 1-9: Paratyphlus espunae nov.sp.: (1) habitus; (2) forebody; (3) male tergite VIII; (4) male sternite VIII; (5-7) aedeagus in lateral view; (8) female tergite VIII; (9) female sternite VIII. Scale bars: 1-2: 0.2 mm; 3-9: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114354" httpUri="https://zenodo.org/record/10114354/files/figure.png" pageId="3" pageNumber="764">Figs 5-7</figureCitation>
.
</paragraph>
<paragraph id="8BCC8D2CFFF8FFA01F3BFCBCFC07FCCC" blockId="3.[340,1247,799,851]" pageId="3" pageNumber="764">
: tergite VIII oblong and with weakly concave posterior margin (
<figureCitation id="134891A9FFF8FFA01A59FC83FBA6FCAA" box="[1035,1098,799,821]" captionStart="Figs 1-9" captionStartId="2.[340,380,1653,1673]" captionTargetBox="[343,1236,425,1618]" captionTargetId="figure-10@2.[340,1253,689,1167]" captionTargetPageId="2" captionText="Figs 1-9: Paratyphlus espunae nov.sp.: (1) habitus; (2) forebody; (3) male tergite VIII; (4) male sternite VIII; (5-7) aedeagus in lateral view; (8) female tergite VIII; (9) female sternite VIII. Scale bars: 1-2: 0.2 mm; 3-9: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114354" httpUri="https://zenodo.org/record/10114354/files/figure.png" pageId="3" pageNumber="764">Fig. 8</figureCitation>
); sternite VIII oblong, its posterior margin acutely angled in the middle (
<figureCitation id="134891A9FFF8FFA01DF2FCA1FC33FCCC" box="[928,991,829,851]" captionStart="Figs 1-9" captionStartId="2.[340,380,1653,1673]" captionTargetBox="[343,1236,425,1618]" captionTargetId="figure-10@2.[340,1253,689,1167]" captionTargetPageId="2" captionText="Figs 1-9: Paratyphlus espunae nov.sp.: (1) habitus; (2) forebody; (3) male tergite VIII; (4) male sternite VIII; (5-7) aedeagus in lateral view; (8) female tergite VIII; (9) female sternite VIII. Scale bars: 1-2: 0.2 mm; 3-9: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114354" httpUri="https://zenodo.org/record/10114354/files/figure.png" pageId="3" pageNumber="764">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8BCC8D2CFFF8FFA01F06FCFDFC57FBEC" blockId="3.[340,1248,865,1139]" pageId="3" pageNumber="764">
C o m p a r a t i v e n o t e s: The genus 
<taxonomicName id="4C73F6AFFFF8FFA01D60FCFDFB7EFCE8" authority="BLACKWELDER 1952" authorityName="BLACKWELDER" authorityYear="1952" box="[818,1170,865,887]" class="Insecta" family="Staphylinidae" genus="Paratyphlus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="764" phylum="Arthropoda" rank="genus">
<emphasis id="B907513EFFF8FFA01D60FCFDFC42FCE9" box="[818,942,865,886]" italics="true" pageId="3" pageNumber="764">Paratyphlus</emphasis>
BLACKWELDER 1952
</taxonomicName>
is distributed in the Iberian Peninsula (including the Baleares) and North Africa from 
<collectingCountry id="F364CDBCFFF8FFA01AD6FCE2FB33FC0C" box="[1156,1247,894,915]" name="Morocco" pageId="3" pageNumber="764">Morocco</collectingCountry>
to 
<collectingCountry id="F364CDBCFFF8FFA01F24FC05FE29FC30" box="[374,453,921,943]" name="Tunisia" pageId="3" pageNumber="764">Tunisia</collectingCountry>
; it currently includes 29 species (
<bibRefCitation id="EFE2F0DDFFF8FFA01D19FC05FC02FC30" author="COIFFAIT H" box="[843,1006,921,943]" pageId="3" pageNumber="764" pagination="237 - 437" refId="ref6600" refString="COIFFAIT H. (1959): Monographie des Leptotyphlites (Col. Staphylinidae). - Revue Francaise d'Entomologie 26: 237 - 437." type="journal article" year="1959">COIFFAIT 1959</bibRefCitation>
, 
<bibRefCitation id="EFE2F0DDFFF8FFA01A53FC05FBDBFC30" author="COIFFAIT H" box="[1025,1079,921,943]" pageId="3" pageNumber="764" pagination="237 - 437" refId="ref6626" refString="COIFFAIT H. (1972): Coleopteres Staphylinidae de la region palearctique occidentale. I. Generalites. Sous-familles: Xantholininae et Leptotyphlinae. - Supplement a la Nouvelle Revue d'Entomologie 2 (2): 237 - 437." type="journal article" year="1972">1972</bibRefCitation>
; HERNANDO &amp; FANCELLO 2004; 
<bibRefCitation id="EFE2F0DDFFF8FFA01C5AFC29FD47FC54" author="SMETANA A" box="[520,683,949,971]" pageId="3" pageNumber="764" pagination="505 - 698" refId="ref7044" refString="SMETANA A. (2004): Staphylinidae, subfamilies Omaliinae-Dasycerinae, Phloecharinae- Apaticinae, Piestinae-Staphylininae, pp. 237 - 272, 329 - 495, 505 - 698. - In: LOBL I. &amp; A. SMETANA (editors). Catalogue of Palaearctic Coleoptera. II. Hydrophiloidea - Histeroidea - Staphylinoidea. - Apollo Books, Stenstrup, 942 pp." type="book chapter" year="2004">SMETANA 2004</bibRefCitation>
). 13 species have been recorded from mainland 
<collectingCountry id="F364CDBCFFF8FFA01ACDFC29FB30FC54" box="[1183,1244,949,971]" name="Spain" pageId="3" pageNumber="764">Spain</collectingCountry>
. The geographically closest representatives of the genus are 
<taxonomicName id="4C73F6AFFFF8FFA01D81FC4EFE5CFB9C" authority="COIFFAIT 1955" authorityName="COIFFAIT" authorityYear="1955" class="Insecta" family="Staphylinidae" genus="Paratyphlus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="764" phylum="Arthropoda" rank="species" species="mateui">
<emphasis id="B907513EFFF8FFA01D81FC4EFBD5FC78" box="[979,1081,978,999]" italics="true" pageId="3" pageNumber="764">P. mateui</emphasis>
COIFFAIT 1955 (Malaga)
</taxonomicName>
and 
<taxonomicName id="4C73F6AFFFF8FFA01FBFFC72FC69FB9C" authority="COIFFAIT 1957" authorityName="COIFFAIT" authorityYear="1957" box="[493,901,1005,1027]" class="Insecta" family="Staphylinidae" genus="Paratyphlus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="764" phylum="Arthropoda" rank="species" species="morandi">
<emphasis id="B907513EFFF8FFA01FBFFC72FD8AFB9D" box="[493,614,1005,1027]" italics="true" pageId="3" pageNumber="764">P. morandi</emphasis>
COIFFAIT 1957 (Alicante)
</taxonomicName>
. Species of 
<taxonomicName id="4C73F6AFFFF8FFA01A42FC71FB60FB9D" authorityName="BLACKWELDER" authorityYear="1952" box="[1040,1164,1005,1026]" class="Insecta" family="Staphylinidae" genus="Paratyphlus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="764" phylum="Arthropoda" rank="genus">
<emphasis id="B907513EFFF8FFA01A42FC71FB60FB9D" box="[1040,1164,1005,1026]" italics="true" pageId="3" pageNumber="764">Paratyphlus</emphasis>
</taxonomicName>
can be distinguished only based on the male primary sexual characters, which is why the external morphology of 
<taxonomicName id="4C73F6AFFFF8FFA01C45FBBAFD6AFBA4" authorityName="Assing" authorityYear="2007" box="[535,646,1062,1083]" class="Insecta" family="Staphylinidae" genus="Paratyphlus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="764" phylum="Arthropoda" rank="species" species="espunae">
<emphasis id="B907513EFFF8FFA01C45FBBAFD6AFBA4" box="[535,646,1062,1083]" italics="true" pageId="3" pageNumber="764">P. espunae</emphasis>
</taxonomicName>
is not explicitly described above, apart from the secondary sexual characters. For illustrations of the aedeagi of other 
<taxonomicName id="4C73F6AFFFF8FFA01D8DFBDDFB5DFBC8" box="[991,1201,1089,1111]" class="Insecta" family="Staphylinidae" genus="Paratyphlus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="764" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B907513EFFF8FFA01D8DFBDDFBB7FBC9" box="[991,1115,1089,1110]" italics="true" pageId="3" pageNumber="764">Paratyphlus</emphasis>
species
</taxonomicName>
see 
<bibRefCitation id="EFE2F0DDFFF8FFA01F06FBC1FE19FBEC" author="COIFFAIT H" box="[340,501,1117,1139]" pageId="3" pageNumber="764" pagination="237 - 437" refId="ref6600" refString="COIFFAIT H. (1959): Monographie des Leptotyphlites (Col. Staphylinidae). - Revue Francaise d'Entomologie 26: 237 - 437." type="journal article" year="1959">COIFFAIT (1959</bibRefCitation>
, 
<bibRefCitation id="EFE2F0DDFFF8FFA01C52FBC1FDD8FBEC" author="COIFFAIT H" box="[512,564,1117,1139]" pageId="3" pageNumber="764" pagination="237 - 437" refId="ref6626" refString="COIFFAIT H. (1972): Coleopteres Staphylinidae de la region palearctique occidentale. I. Generalites. Sous-familles: Xantholininae et Leptotyphlinae. - Supplement a la Nouvelle Revue d'Entomologie 2 (2): 237 - 437." type="journal article" year="1972">1972</bibRefCitation>
) and HERNANDO &amp; FANCELLO (2004).
</paragraph>
<caption id="DF0CDDA4FFF8FFA01F06F94FFCACF978" ID-DOI="http://doi.org/10.5281/zenodo.10114356" ID-Zenodo-Dep="10114356" box="[340,832,1746,1767]" httpUri="https://zenodo.org/record/10114356/files/figure.png" pageId="3" pageNumber="764" startId="3.[340,373,1747,1767]" targetBox="[340,1254,1155,1735]" targetPageId="3">
<paragraph id="8BCC8D2CFFF8FFA01F06F94FFCACF978" blockId="3.[340,832,1746,1767]" box="[340,832,1746,1767]" pageId="3" pageNumber="764">
<emphasis id="B907513EFFF8FFA01F06F94FFE7AF978" bold="true" box="[340,406,1747,1767]" pageId="3" pageNumber="764">Fig. 10</emphasis>
: Type locality of 
<taxonomicName id="4C73F6AFFFF8FFA01C6AF94EFD1BF978" authorityName="Assing" authorityYear="2007" box="[568,759,1746,1767]" class="Insecta" family="Staphylinidae" genus="Paratyphlus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="764" phylum="Arthropoda" rank="species" species="espunae" status="sp. nov.">
<emphasis id="B907513EFFF8FFA01C6AF94EFD1BF978" box="[568,759,1746,1767]" italics="true" pageId="3" pageNumber="764">Paratyphlus espunae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234EC45FFF8FFA01CAFF94FFCACF978" box="[765,832,1747,1767]" pageId="3" pageNumber="764" rank="species">nov.sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BCC8D2CFFFFFFA71D55FEF9FCC7FEE5" blockId="4.[775,811,357,378]" box="[775,811,357,378]" pageId="4" pageNumber="765">765</paragraph>
<paragraph id="8BCC8D2CFFFFFFA71F06FE25FD87FE74" blockId="4.[340,1247,440,491]" pageId="4" pageNumber="765">
E t y m o l o g y: The name (noun, genitive) is derived from the mountain range where the 
<typeStatus id="54C8338EFFFFFFA71F2BFE4AFE49FE74" box="[377,421,470,491]" pageId="4" pageNumber="765">type</typeStatus>
locality is situated.
</paragraph>
<paragraph id="8BCC8D2CFFFFFFA71F06FE67FD3CFD1A" blockId="4.[340,1247,506,645]" pageId="4" pageNumber="765">
D i s t r i b u t i o n a n d b i o n o m i c s: As can be inferred from the restricted distributions of the vast majority of leptotyphlines, 
<taxonomicName id="4C73F6AFFFFFFFA71D0EFD85FC20FDB1" authorityName="Assing" authorityYear="2007" box="[860,972,537,558]" class="Insecta" family="Staphylinidae" genus="Paratyphlus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="765" phylum="Arthropoda" rank="species" species="espunae">
<emphasis id="B907513EFFFFFFA71D0EFD85FC20FDB1" box="[860,972,537,558]" italics="true" pageId="4" pageNumber="765">P. espunae</emphasis>
</taxonomicName>
is probably endemic to the Sierra de Espuña, where the 
<typeStatus id="54C8338EFFFFFFA71CDBFDAAFD52FDD4" box="[649,702,566,587]" pageId="4" pageNumber="765">types</typeStatus>
were collected in a pasture by washing sandy soil between stones and grass roots (
<figureCitation id="134891A9FFFFFFA71CF0FDCEFD02FDF7" box="[674,750,594,616]" captionStart="Fig" captionStartId="3.[340,373,1747,1767]" captionTargetBox="[340,1254,1155,1735]" captionTargetId="figure-325@3.[340,1254,1155,1735]" captionTargetPageId="3" captionText="Fig. 10: Type locality of Paratyphlus espunae nov.sp." figureDoi="http://doi.org/10.5281/zenodo.10114356" httpUri="https://zenodo.org/record/10114356/files/figure.png" pageId="4" pageNumber="765">Fig. 10</figureCitation>
) at altitudes of 1100 and 
<quantity id="4C8B20C9FFFFFFA71DA8FDCEFBA7FDF7" box="[1018,1099,594,616]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="4" pageNumber="765" unit="m" value="1140.0">1140 m</quantity>
, together with 
<taxonomicName id="4C73F6AFFFFFFFA71F06FDECFDE8FD1A" authorityName="Assing" authorityYear="2007" box="[340,516,624,645]" class="Insecta" family="Staphylinidae" genus="Meotica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="765" phylum="Arthropoda" rank="species" species="andujari">
<emphasis id="B907513EFFFFFFA71F06FDECFDE8FD1A" box="[340,516,624,645]" italics="true" pageId="4" pageNumber="765">Meotica andujari</emphasis>
</taxonomicName>
nov.sp (see below).
</paragraph>
</subSubSection>
</treatment>
</document>