Revision of the Australian species of Figulus MacLeay, 1819 (Coleoptera: Lucanidae)
Author
Monte, Cinzia
Author
Zilioli, Michele
Author
Bartolozzi, Luca
text
Zootaxa
2016
4189
3
447
484
journal article
37842
10.11646/zootaxa.4189.3.2
c55b8fbe-5e88-4dae-89fb-38df76c0b839
1175-5326
166170
3B503E4F-7748-4AC4-965E-B010E225255B
Figulus regularis
Westwood, 1834
(
Figs. 16–19
)
Figulus regularis
Westwood, 1834
: 120
;
Dejean 1836
: 194
;
Westwood 1838
: 263
;
Westwood 1845
: 7
;
Burmeister 1847
: 437
;
Reiche 1853
: 84
;
Thomson 1856
: 8
;
Thomson 1862
: 402
, 432;
Parry 1863
: 450
;
Parry 1864a
: 6
;
Parry 1864b
: 99
;
Harold 1868
: 965
;
Macleay 1871
: 173
;
Parry 1875
: 22
;
Gestro 1880
: 335
;
Albers 1883
: 225
;
Masters 1888
: 301
;
Boileau 1898
: 433
;
Felsche 1898
: 78
;
Heyne & Taschenberg 1908
: 57
;
Van Roon 1910
: 52
;
Boileau 1913
: 265
;
Mjöberg 1916
: 13
;
Kriesche 1922
: 133
;
Nagel 1928
: 294
;
Didier 1937
: 30
;
Nagel 1939
: 1
;
Didier & Séguy 1953
: 182
;
Benesh 1960
: 27
;
Holloway 1960
: 343
;
De Lisle 1964
: 4
;
Holloway 1969
: 974
;
Tiberghien & Lacroix 1969
: 535
;
Williams 1977
: 4
;
Klausnitzer 1982
: 16
;
Hawkeswood 1985
: 200
;
Bomans 1986
: 9
(
lectotype
designation);
Hawkeswood 1988
: 270
–272;
Maes 1992
: 52
;
Moore & Cassis 1992
: 10
;
Mizunuma & Nagai 1994
: 300
;
Mizunuma 2000
: 85
;
Krajcik 2001
: 12
;
Krajcik 2003
: 82
;
Klausnitzer & Sprecher-Uebersax 2008
: 33
;
Fujita 2010
: 382
;
Bartolozzi
et al
. 2011
: 45
;
Hawkeswood 2014
: 1
;
Maes 2014
;
Schoolmeesters 2015
.
Figulus australicus
Thomson, 1862
: 402
, 432; synonymized by
Benesh 1960
: 27
.
Figulus laevithorax
Bomans, 1987
: 6
;
Maes 1992
: 38
;
Moore & Cassis 1992
: 9
;
Krajcik 2001
: 11
;
Krajcik 2003
: 78
;
Maes 2014
;
Schoolmeesters 2015
.
New synonymy
.
Type material
.
Figulus regularis
Westwood
, 1Ƌ lectotype (designation by
Bomans 1986
): “N.H. [handwritten]”, “
Figulus regularis
/ Westw. in ann. Sc. nat. [handwritten]”, “TYPE / WESTWOOD / Ann. Sci. Nat. / 1834 P.120 / Coll. Hope Oxon. [printed, handwritten]”, “TYPE COL: 327
1
/
2
/
Figulus
/
regularis
Westw.
/ HOPE DEPT. OXFORD [printed, handwritten]”, [OUMNH];
Figulus regularis
Westwood
, 1♀ paralectotype: “N.H. [handwritten]”, “TYPE / WESTWOOD / Ann. Sc. Nat. / 1834 P.120 / Coll. Hope Oxon. [printed, handwritten]”, “TYPE COL: 327
2
/
2
/
Figulus
/
regularis
Westw.
/ HOPE DEPT. OXFORD [printed, handwritten]”, [OUMNH].
Figulus australicus
Thomson
:
Lectotype here designated
:
Lectotype male
: “Th. / TYPE [printed]”, “
Australicus
/ Type Thoms / Austr. [handwritten]”, “Ex musaeo / JAMES THOMSON [printed]”, “MUSÉUM PARIS / 1952 / COLL R OBERTHÜR [printed]” [MNHN].
Figulus laevithorax
Bomans
(Ƌ): “Holotype [printed]”, “Cap York [handwritten]”, “
Figulus
/
regularis
Ƌ / Australien [handwritten]”, “1431 [handwritten]”, “H.E. Bomans / BMNH (E) / 1999-248 [handwritten] \ 401431 [handwritten]”, “H.Bomans det., 1974 /
Figulus
/
laevithorax
n.sp.
[printed, handwritten]” [NHM];
Figulus laevithorax
Bomans
(Ƌ): “Para- / type [printed]”, “P. Darwin [handwritten]”, “Macleay Museum / University of Sydney / 2006 Australia [printed]”, “11382 [handwritten]”, “H.E. Bomans / BMNH (E) / 1999-248 [printed] \ 411382 [handwritten]”, “H.Bomans det., 1986 /
Figulus
/
laevithorax
Bom.
[printed, handwritten]” [NHM].
Other material
: 1 specimen “
Figulus
/
regularis
/ Westwood NH [handwritten]”, “Ex Musaeo / Parry [printed]”, “MUSÉUM PARIS / 1952 / Coll. R. OBERTHÜR [printed]”, [MNHN]; 1 specimen “
F. re g ul ar i s
/
v. minor
[handwritten]”, “Ex Musaeo / Parry [printed]”, “MUSÉUM PARIS / 1952 / Coll. R. OBERTHÜR [printed]”, [MNHN]; 1 specimen “Moreton / Bay [handwritten]”, “Ex Musaeo / Parry [printed]”, “MUSÉUM PARIS / 1952 / Coll. R. OBERTHÜR [printed]”, [MNHN]. Note: this material was indicated as “syntypes” in the loan form, but their status as type material is uncertain.
Diagnosis.
Figulus regularis
is similar to
F. boileaui
, from which it can be distinguished by the shape of labrum, which is strongly protruded and subtriangular, instead of broad, short, and slightly convex at the anterior margin; by the medial groove on the pronotal disc, which is superficial instead of deep and lanceolate; by the absence of microreticulation on pronotal disc, and by the different shape of the male genitalia.
Redescription.
Measurements:
size range (
n
= 777): TL:
10.91–18.25 mm
; PL:
2.92–4.83 mm
; EL:
6.25– 10.08 mm
; PW:
3.50–6.25 mm
; EW:
3.75–6.08 mm
.
Male
(
Fig. 16
). Body elongate, with subparallel sides, black or dark brown, glossy.
Head
: transverse, about half and two times wider than long, slightly depressed. Frontoclypeal region slightly concave at the anterior margin densely and coarsely punctate, with two large, glossy areas situated before the eyes and a medial ridge of frons that varies from weakly developed to absent. Labrum strongly protruding, narrow and almost equilateral triangular, with apex blunt and long, golden setae. Vertex slightly concave in the middle, inclined anteriorly, with two high transversal elevations. Two slightly elevated, longitudinal carinae are present along the inner margin of eyes. Eyes large, divided into 2 parts by the canthi, the upper part smaller than the lower with distinct facets. Canthus largely protruding, rounded laterally and directed obliquely inwards behind; upper surface slightly depressed and smooth, with sparse, minute punctures on the lateral margin. Head surface densely punctate with some smooth areas, on canthi, on frons and between the transversal elevations on vertex. Mandibles longer than head, densely and minutely punctate, strong and asymmetrical, rounded laterally, apices slightly blunted and strongly upwards; upper surface slightly elevated at base. Internal margin of the right mandible with a moderately developed basal tooth and a medial strong tooth, with apex blunted directed inwards. Internal margin of the left mandible with a moderately developed basal tooth, a well developed medial tooth and a preapical angulation, moderately developed. Antenna with 10 antennomeres, scape slightly shorter than the remaining antennomeres; club of last three antennomeres, which are flatter and broader than the remaining antennomeres of the funicle.
Mentum
subrectangular (
Fig. 17
), medially with 2 smooth irregular concavities almost always confluent; anterior margin slightly bilobed and sides from almost straight to slightly rounded. Surface strongly punctate at sides, at base and along anterior margin.
FIGURES 16‒19.
Figulus regularis
Westwood.
16, dorsal view of adult (scale bar: 5 mm); 17, mentum; 18, male genitalia, dorsal view (scale bar: 0.5 mm); 19, male genitalia, lateral view.
Thorax
: pronotum slightly wider than long with subparallel sides; anterior margin slightly convex, slightly to strongly elevated, with a small, blunt medial tubercle; posterior margin slightly bisinuate; subparallel sides; anterior angles produced and almost right angled with a narrow rim; the posterior angles largely rounded, with slightly crenulate margin. Longitudinal medial groove deep or sometimes shallow on disc, with some large, circular punctures. Pronotal surface glossy, with dense, minute, superficial punctures on disc and with patch of large close punctures at the anterolateral margin and at basal 2/3. Scutellum long, wedge shaped; surface smooth and glossy.
Elytra
: about twice as long and as wide as pronotum. Base concave, subparallel sides slightly crenulate at the base; apex regularly rounded. Humeral angles acute, with a small, acute tubercle. Elytral surface glossy, with 9 visible subparallel striae, the dorsal striae deeper than the lateral striae; each stria formed by a row of large, adjacent punctures, except for the first stria, where the punctures are confluent and is the only stria than reaches at apex. Striae 3‒6 and 4‒5 converging towards apex. Interstriae much wider than striae, slightly convex and minutely punctate. Elytral margins and apex strongly punctate, with short, erect, red setae. Wings present. Legs thin and straight. Protibiae with elongate apical fork, and a long, strong spur internally at apex; lateral margin with 4‒5 small teeth, increasing in size distally. Mesotibiae with 4 long, acute apical teeth, and 2 apical spurs, 1 larger than the other; lateral margin with strong submedial tooth, preceded by 1‒2 smaller teeth. Metatibiae with 4 acute apical teeth, and 2 long, apical spurs, 1 larger than the other; lateral margin with strong submedial tooth, preceded by a smaller tooth. Prosternum subpentagonal in shape behind the coxae; surface with large and deep punctures at anterior edge and with smaller punctures medially and at posterior edge. Metasternum smooth, with a long medial groove, deep at posterior edge.
Abdomen
: first abdominal ventrite with some horseshoe-shaped, superficial hollows bringing each a short, golden seta in the middle. The other ventrites have some large and deep punctures along the anterior margin and some horseshoe-shaped, superficial hollows at sides, punctures are fine in the middle. Last ventrite medially with dense, small punctures, becoming smaller towards the posterior margin.
Male genitalia
: penis tubular, short, evenly tapering towards apex in dorsal view (
Fig. 18
); in lateral view completely covered by a dorsal translucid, membranous extension of the basal piece, nearly straight, more curved proximally close to articulation with basal piece (
Figs. 18–19
). Permanently everted internal sac drawn out into a long and broad, colorless or yellow flagellum tapering towards apex. Parameres slightly shorter than the basal piece, apically broadly spatulate, slightly angulate at inner margin, with marginal fringe of short setae entirely enclosed in the cuticle (not represented in figure). Basal piece elongate, sclerotized, nearly parallel-sided; ventrally mostly membranous, dorsally with a moderately broad, nearly parallel-sided sclerotic area along midline (
Figs. 18– 19
). A pair of long, slender struts run along basal piece, from penis articulation to slightly beyond the base of the short, truncate expansion of basal piece.
Female.
All external morphological characters as in male.
Female genitalia
: hemisternites elongate, apically with long, golden setae surrounding the styli; moderately broad, enlarged medially; outer edge almost straight or slightly sinuose, medial edge convex; styli slender with apex straight, slightly longer than wide. Bursa copulatrix represented by a slight expansion of the wall of spermathecal duct near junction with the median oviduct. Spermathecal duct long, ending in a bulbous, pearshaped expansion; colorless in the proximal half, pigmented in the distal half.
Intraspecific variation.
The medial ridge of frons varies from weakly developed to absent; the prothorax can be nearly parallel or slightly broadened posteriorly. For the body size see the “Measurements” section.
Additional material examined.
Australia
[state unknown]:
1 specimen
,
Nouv
.
Galles du Sud, Mniszech 1856 (
MNHN
) [note: at this date
New
South Wales
included
Queensland
, the two states split up in 1859; so we put this specimen in this section]
. 1 specimen (IRSNB); 1 specimen, R. Mus. Hist. Nat. Belg. I.G. 12.595 (IRSNB); 3 specimens, R.I.Sc.N.B. 21.418, Collection P. de Moffarts (IRSNB);
1 specimen
,
Muséum Paris
1935
Collection Sedillot
(
MNHN
)
;
1 specimen
, ex
Musaeo Mniszech
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
1 specimen
, 1853,
Muséum Paris
ex
Collection Perroud
(
MNHN
)
;
1 specimen
,
Australia
Or.,
Act
,
X.1870
,
H.Deyr
(
MSNG
)
; 2 specimens (GTC); 1 specimen,
30.VIII.1946
(GTC); 2 specimens (MEL);
3 specimens
,
Muséum Paris J.
Verreaux 2-47 (
MNHN
)
;
3 specimens
,
Muséum Paris
(
MNHN
)
; 4 specimens
15.VII.1927
,
Lina Rasim
ded. (SMNS);
1 specimen
, Nouvelle Hollande, ex
Musaeo Van Lansberge
,
Muséum Paris
1982
Collection Oberthür
(
MNHN
)
;
1 specimen
, Nouvelle Hollande,
Muséum Paris
1935
Collection M.
Sedillot (
MNHN
)
;
1 specimen
, Nouvelle Hollande,
Vict.
,
Edwards
,
Fry Collection
1905-100,
BMNH
(E) # 607564 (
NHM
)
;
3 specimens
, Nouvelle Hollande,
Collection E.
Candèze (
IRSNB
)
;
2 specimens
, Nouvelle Hollande, (
MEL
,
IRSNB
)
.
NEW SOUTH WALES
:
1 specimen
,
New
South Wales
, ex
Musaeo Van Lansberge
, Muséum Paris 1952 Collection Oberthür (
MNHN
)
;
1 specimen
,
New
South Wales
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
1 specimen
,
New
South Wales
, (
GTC
)
;
1♀
,
6 specimens
,
New
South Wales
, COL-85935, COL-85936, COL-85937, COL-85938, COL-85939, COL-85858, and COL-85859 (
NMVM
)
;
1 specimen
,
New
South Wales
,
Nat. Mus
.
Victoria
,
C.French’s Collection
6.I.08
, COL-85903 (
NMVM
)
;
1 specimen
,
New
South Wales
,
Collection Ogier de Baulny
, 8114 (
IRSNB
)
;
1 specimen
,
Northern
N.S.W., 1972,
G.May
,
C.G.L.Gooding
collection, donated to
ANIC
1979
(
ANIC
)
;
2 specimens
,
Albury
,
4.VII.1937
,
F.E.Wilson
,
F.E.Wilson Collection
, COL-85908 (
NMVM
)
;
1 specimen
,
Baan Baa
,
29.X.1908
,
G.E.Bryant
,
G.E.Bryant Collection
1919-147, # 607613 (
NHM
)
;
2 specimens
,
Broad Sound
,
Queensland
,
24.V.1957
,
Banksia
, G.P.Witley, K
293171 and K 293172 (AMS)
;
1 specimen
,
Banyabba Nature Reserve
, near
Coaldale
,
8.I.1921
,
R.Hardie
,
on
Xanthorrhoea
,
University
of New England Collection, donated 1983 (
ANIC
)
;
1 specimen
,
Belubula
,
21.IV.1923
,
R. Musgrave
,
K 48145
(handwritten), K 293099 (AMS)
;
1 specimen
,
Bingara
,
5.VI.1992
,
O.Hillert
, ex
Collection Werner
(
LBC
)
;
1 specimen
,
Blandford
,
IX.1908
, C.S.,
K 22618
(handwritten), K 293121 (AMS)
;
1 specimen
,
Bogan Gate
,
Blow Glearwest State Forest
, R.H.M.
, K 293145,
R.H.Mulder Collection
(AMS)
;
1 specimen
,
Bugaldie
,
25.II.1972
,
D.A.Doolan
, K 293136,
D.A.Doolan Collection
(AMS)
;
2 specimens
,
Burma
Road,
Pilliga East State Forest
,
5–7.XI.1999
, R.deKeyzer &
A.Sundholm
, K 293309, K 293310, R.deKeyzer
Insect Collection
(AMS)
;
1 specimen
,
Burrier
,
29.XII.1933
,
Rodway
,
University
of New England Collection, donated 1983 (
ANIC
)
; 1 specimen, Caldwell, V.Robb, K 293107 (AMS);
1 specimen
,
Capella
,
Retro Station
,
VIII.1929
,
J.R.Sievin
,
K 61074
(handwritten), K293162 (AMS)
; 3 specimens, Caragabol, P.H.Taylor, K 293115, K 293116 and K 293120 (AMS); 2 specimens, Clarence R.,
I.1904
, R.Helms, K 293098 (AMS);
1 specimen
,
Condobolin
,
11.II.1972
,
D.A.Doolan
,
D.A.Doolan Collection
, K 293108 (AMS)
;
1 specimen
,
40 km
N of
Coolah
,
1.X.1971
,
S.Misko
(
ANIC
)
;
1 specimen
,
Coonabarabran
,
XII.1929
,
H.J.Carter
(
ANIC
)
;
2 specimens
,
Coonabarabran
,
25.II.1972
,
D.A.Doolan
, K 293134 and K 293135,
D.A. Doolan Collection
(AMS)
;
1 specimen
,
Coonabarabran
,
16.I.1981
,
D.A.Doolan
, K 293111,
D.A.Doolan Collection
(AMS)
;
3 specimens
,
Coonabarabran
,
21.X.1957
,
F.E.Wilson
,
F.E.Wilson Collection
, COL-85170 (
NMVM
)
;
1 specimen
,
CSIRO
Parkes Radio Telescope
,
16.XI.1996
, in soil at base of live
Eucalyptus
, G.W.Ulrich
Collection
(
ANIC
)
;
1 specimen
,
Curlewis
,
29.XI.1933
,
T.Iredale
, K 293105 (AMS)
; 1 specimen, Dabee,
K 28023
(handwritten), K 293097 (AMS);
1 specimen
,
Dorrigo
,
I.1931
,
C.Oke
, COL-85924 (
NMVM
)
;
2 specimens
,
Dubbo
,
29.VIII.
1925
, 149 m (
ANIC
)
;
1 specimen
,
Dubbo
,
25.IV.1975
, D
.
Lester
, K 293142 (AMS); 1Ƌ,
Dunmore
,
I.1988
,
J.Hasenpusch
,
Graeme Krake Collection
, donated
Feb. 2007
, COL-80304 (
NMVM
)
;
1 specimen
,
Emmaville
,
27.III.1973
,
R.H.M.
, K 293109 (AMS)
;
1 specimen
,
Fletcher
,
29.V.1967
,
E.Sutton
(
ANIC
)
; 2 specimens, Fletcher, 1945, E.Sutton, C.G.L.Gooding Collection, donated to ANIC 1979 (ANIC);
1Ƌ,
Fortis Creek State Forest
, N
Grafton,
5.X.1996
, mv light,
S.G.Watkins
, 96:247
Ƌ, S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
3ƋƋ,
2♀♀
,
Fortis Creek State Forest
, N
Grafton,
10.X.1996
, mv light,
S.G.Watkins
, 96:281 Ƌ, 96:282 Ƌ, 96:283
Ƌ, S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
2ƋƋ,
6♀♀
,
Fortis Creek State Forest
, N
Grafton,
13.XI.1996
, mv light,
S.G.Watkins
, 96:540 Ƌ, 96:541
Ƌ, S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
3ƋƋ,
4♀♀
,
Fortis Creek State Forest
, N
Grafton,
6.XII.1996
, mv light,
S.G.Watkins
, 96:709 Ƌ, 96:710 Ƌ, 96:711
Ƌ, S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
1♀
,
Fortis Creek State Forest
, N
Grafton,
9.XII.1996
, mv light,
S.G.Watkins
,
S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
2ƋƋ,
1♀
,
Fortis Creek State Forest
, N
Grafton,
3I.
XII.1996
, mv light,
S.G.Watkins
, 96:1079 Ƌ, 96:1080
Ƌ, S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
5ƋƋ,
4♀♀
,
Fortis Creek State Forest
, N
Grafton,
16.II.1997
, mv light,
S.G.Watkins
, 97:346 Ƌ, 97:347 Ƌ, 97:348 Ƌ, 97:349 Ƌ, 97:350
Ƌ, S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
4ƋƋ,
10♀♀
,
Fortis Creek State Forest
, N
Grafton,
22.II.1997
, mv light,.
G.Watkins
, 97:376 Ƌ, 97:377 Ƌ, 97:378 Ƌ, 97:379
Ƌ, S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
3ƋƋ,
1♀
,
Fortis Creek State Forest
, N
Grafton,
26.II.1997
, mv light,
S.G.Watkins
, under log, 97:412 Ƌ, 97:413 Ƌ, 97:414
Ƌ, S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
2ƋƋ,
Fortis Creek State Forest
, N
Grafton,
4.III.1997
, mv light,
S.G.Watkins
, 97:457 Ƌ, 97:458
Ƌ, S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
3ƋƋ,
Fortis Creek
National Park, SE
boundary,
N Grafton
,
6–7.II.1998
,
S.G.Watkins
, under log,
60 m
, 98:207 Ƌ, 98:217 Ƌ, 98:305
Ƌ, S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
1Ƌ,
Fortis Creek
National Park
, seast boundary,
N Grafton
,
50 m
,
16.XI.1998
,
S.G.Watkins
, 98:643 Ƌ, mv light,
S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
2ƋƋ,
2♀♀
,
Fortis Creek
National Park
, seast boundary,
N Grafton
,
50 m
,
17–20.XII.1998
,
S.G.Watkins
, 98:836 Ƌ, 98:852 Ƌ, mv light,
S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
2ƋƋ,
1♀
,
Fortis Creek
National Park, SE
boundary,
N Grafton
,
50 m
,
13.I.1999
,
S.G.Watkins
, 99:105 Ƌ, 98:106 Ƌ, mv light,
S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
2ƋƋ,
Fortis Creek
National Park
, seast boundary,
N Grafton
,
50 m
,
4.I.2000
, 00:3, 00:4, mv light,
Copmanhurst
9438-1-N 56JMN968328,
S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
1 specimen
,
Gilgandra
,
Biddon State Forest
,
3.XII
–
I.1986
, R.H.M., K 293148, R.H.Mulder Collection (AMS);
1 specimen
,
Glenview
,
12 km
S Inverell
,
29.VIII.1985
,
N.S. & S.G.Hunter
, K 293 147 (AMS)
; 1 specimen, SE boundary of Goobang National Park, Circuit Road, 33°11'19.7''S, 148°25'13.7''E, R.deKeyzer & A.Scott, K 293316, R.deKeyzer Insect Collection (AMS);
2 specimens
,
Goonoo State Forest
,
12–18.I.1981
,
D.A.Doolan
, K 293139 and
1 specimen
,
17.I.1981
,
R.H.Mulder
, K 293119, (AMS)
;
1 specimen
,
Goonoo State Forest
,
6 km
SW Mendooran
,
6.IV.1979
,
D.K.McAlpine
&
B.J.Day
, K 293110 (AMS)
;
1 specimen
,
Goonoo State Forest
,
Mogriguy
,
24.I.1980
, K 293144 (AMS)
;
1 specimen
,
Griffith
,
22.XI.1955
,
G.F.Bornemissza
(
ANIC
)
;
1 specimen
,
Gulgong
,
I.1986
,
F.E.Wilson
,
F.E.Wilson Collection
, COL- 85164 (
NMVM
)
;
1 specimen
,
Gunnedah
,
X.1959
,
F.E.Wilson
,
F.E.Wilson Collection
, COL-85165 (
NMVM
)
;
1 specimen
,
Mount Kaputar
,
Bullawa Creek
,
28.XI.1984
,
George Hangay
(
ANIC
)
;
3 specimens
,
Lambton
, ex
Musaeo Möllenkamp
1913,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
; 2 specimens, Macdonald R.,
23.II.1971
, D.A.Doolan, K 293122 and K 293123, D.A.Doolan Collection (AMS);
1 specimen
,
Maitland
,
26.XII.1962
, COL-86041 (
NMVM
)
;
1 specimen
,
24 km
NNE of
Mendooran
,
17.I.1981
,
D.A.Doolan
, K 293118,
D.A.Doolan Collection
(AMS)
;
1 specimen
,
Mogriguy
,
6.I.1979
,
R.H.M.
K 293143,
R.H.Mulder Collection
(AMS)
;
5 specimens
,
Mooney Mooney Range
near
Coolac
,
7.X.1974
, ex dead stem of
Xanthorrhoea australis
, K.L.Pullen
(
ANIC
)
;
2 specimens
,
73 km
N of
Moree
,
29.XII.1977
,
R.G.Thompson Collection
, COL-86044 and COL-86045 (
NMVM
)
; 1 specimen, Mulwala, COL-86033 (NMVM);
1 specimen
,
Munmorah State Conservation Area
, -33º12'34'', 151º34'59'',
1.V.1997
,
L.Wilkie
, K 136541 (AMS)
; 1 specimen, Murrurundi, 1841,
K 28030
(handwritten), K 293096 (AMS);
2 specimens
,
Narrabri
,
Sydney
,
30–31.X.1924
,
W.W.Froggatt
(
ANIC
)
;
1 specimen
,
Narrabri
,
Jacks Creek
,
5.XI.1932
,
K.C.McKeown
,
K 66664
(handwritten), K 293104 (AMS)
; 1 specimen, Narrabri, J.Armstrong, K.K.Spence Collection, K 293102 (AMS);
1 specimen
,
Native Dog Creek
Road
,
14.V.1999
,
W.W.Froggatt Collection
(
ANIC
)
;
1 specimen
,
Nymagee
,
16.XI.1962
,
H.Cogger
, K 293117 (AMS)
;
1 specimen
,
Ogilvie Trail
,
Banyabba Nature Reserve
, SSW
Whiporie
,
130 m
,
27.XI.2000
, 00:214, mv light (
ANIC
)
;
1Ƌ,
1♀
,
Olives Swamp Myrtle State Forest
,
28.X.1996
,
S.G.Watkins
, 96:360 Ƌ, mv light,
S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
1 specimen
,
Olives Swamp Myrtle State Forest
, SSE
Rappville
,
50 m
,
27.I.1999
,
S.G.Watkins
, mv light,
S.G.Watkins Collection
, donated 2001 (
ANIC
)
;
1 specimen
,
Ourimbah, C.
Schindler 1905 (
MNHN
)
;
2 specimens
,
Pilliga Scrub
,
XI.1999
R.deKeyzer (
LBC
) and (
MZUF
)
;
3 specimens
,
Pilliga Scrub
,
25 m
, W of
Narrabri
,
30.X.1967
,
C.W.Frazier
,
University
of New England Collection, donated 1983 (
ANIC
)
;
7 specimens
,
Pilliga Scrub
,
XI.1999
, R.deKeyzer,
Graeme Krake Collection
, donated
Feb. 2007
, COL-86871, COL- 86872, COL-86877, COL-86878, COL-86879, COL-86880, and COL-86881 (
NMVM
)
;
1♀
,
1 specimen
,
Pilliga State Forest
, 20
Foot Tk
,
4.xi.1994
(
MEL
)
;
1 specimen
,
Pilliga State Forest
,
Greens Road
,
Logging Site
,
2.XI.1991
, R.deKeyzer &
A.Sundholm
, K 293305, R.deKeyzer
Insect Collection
(AMS)
;
4 specimens
,
Pilliga State Forest
,
Pine Creek
Rd
,
XI.1991
, R.deKeyzer &
A.Sundholm
, K 293301, K 293302, K 293303 and K 293304, R.deKeyzer
Insect Collection
(AMS)
;
2 specimens
,
Pilliga
,
1.XII.1992
, R.deKeyzer, ex
Collection Bornemissza
(
SMNS
)
;
3 specimens
,
Pilliga
,
13.II.1994
,
J.A.Walker
, K 293306, K 293307 and K 293308, R.deKeyzer
Insect Collection
(AMS)
;
1 specimen
,
Quirindi
,
3.XI.1908
,
G.E.Bryant
,
G.E.Bryant Collection
1919-147, # 607597 (
NHM
)
;
1 specimen
,
Quirindi
,
V.1928
,
R.E.Jobson
,
K 58096
(handwritten), K 293101 (AMS)
; 1 specimen, Rooty Hill, E.W.Ferguson Collection (ANIC);
1 specimen
,
Salt Caves area
,
Pilliga East State Forest
,
5–7.XI.1999
, R.deKeyzer &
A.Sundholm
, K 293311, R.deKeyzer
Insect Collection
(AMS)
;
1 specimen
,
Shepherds Hill Quarry area
, east side of
Round Hill Nature Reserve
near
Euabalong West
,
23.X.2004
, R.deKeyzer, K 293319, R.deKeyzer
Insect Collection
(AMS)
;
1 specimen
,
Sutherland
,
X.1927
,
M.Fuller.
(
ANIC
)
;
1 specimen
,
Sydney
, (
MNHN
)
;
1 specimen
,
Sydney
,
VIII.1931
,
Dr. K.K.Spence, K.K.Spence Collection, K
293 103 (AMS)
;
12 specimens
, environs
de Sydney, H.
Bureau 1923,
Muséum Paris
(
MNHN
)
; 1 specimen, Turrella, K 293114 (AMS);
1 specimen
,
Upper Colo
,
1.I.1979
,
D.P.Carne
, at uv light (
ANIC
)
; 1 specimen, Wagga, C.Oke, COL-85907 (NMVM);
2 specimens
,
Warrumbungles
,
21–22.II.1972
,
D.A.Doolan
, K 293127 and K 293129,
D.A.Doolan Collection
(AMS)
;
1 specimen
,
Warrumbungles
,
3.X.1979
,
D.A.Doolan
, K 293138,
D.A.Doolan Collection
(AMS)
;
1 specimen
, near
Wellington
,
Ponto Falls
,
18.VII.1982
, in rotten logs,
J.F.Lawrence
(
ANIC
)
;
1 specimen
,
Wellington
Caves,
21.VII.1991
,
W.W.Froggatt Collection
(
ANIC
)
;
1Ƌ,
3 specimens
,
Wentworth
,
24.IV.1990
,
G.J.Krake
,
Graeme Krake Collection
, donated
Feb. 2007
, COL-86873, COL-86874, COL-86875 and COL-86876 (
NMVM
)
;
1 specimen
,
Wheogo
, near
Dunedoo
,
XII.1927
,
A.Musgrave
,
K58473
(handwritten), K 293100 (AMS)
;
2 specimens
,
Wyangala
dam,
24.VIII.2003
(
MEL
)
;
1 specimen
,
Wyee
,
7.VII.1973
,
D.A.Doolan
, K 2 93137,
D.A.Doolan Collection
(AMS)
;
1 specimen
,
Yalwal
, near
Nowra
,
29.xii.1933
,
G.Rodway
, K 293106 (AMS)
;
1 specimen
,
Yathong Nature Reserve
,
Mallee
woodland,
32°58'56''S
,
146°10'58''E
,
13.IX.2000
,
D.Eldridge
,
pitfall trap
., K 293113 (AMS)
;
1 specimen
,
Yathong NR
,
Semi-arid Grassland
,
32°35'58''S
,
145°32'28''E
,
20.XI.2001
,
G.Swan
,
Pitfall trap
., K 293140 (AMS)
;
1 specimen
,
Yathong Nature Reserve
,
Spinifex Grassland
,
32°35'13''S
,
145°29'51''E
,
XI–XII.1999
,
E.Wapstra
,
pitfall trap
., K 293112 (AMS)
.
NORTHERN
TERRITORY
:
18 specimens
,
Bullita
,
Gregory National Park
,
16°07'S
,
130°25'E
,
18–24.V.2001
,
T.Weir
&
P.Bouchard
, under log and bark (
ANIC
)
;
4 specimens
,
Curtin Springs
,
IX.1948
,
Geelong College Expedition
, COL-85158, COL-85159, COL- 85160, and COL-85161 (
NMVM
)
;
1♀
,
Herrmannsburg
,
Central
Australia
,
H.J.Hillier
1911-311,
BMNH
(E) # 607570 (
NHM
)
;
1 specimen
,
MacDonnell Ranges, S.A.
Wight
, ex
Collection Australian Museum
(
ANIC
)
;
1 specimen
,
Northern
Territory
,
Yulara
,
III.2007
(
CMC
)
;
1 specimen
,
Plenty Highway
,
268 km
ENE of Alice Springs
,
13.X.1978
,
Upton
&
Barrett
, 8637,
H.E.Bomans
BMNH
(E) 1999-248, 408637 (
NHM
)
;
1 specimen
,
Ti Tree
,
30.XII.1982
, at light,
D.P.Carne
(
ANIC
)
;
1 specimen
,
Tindal
,
1.XII.1967
,
W.J.Vestgens
(
ANIC
)
;
1 specimen
,
Victoria
River Crossing
,
29.XII.1994
(
MSNT
)
.
QUEENSLAND
:
1 specimen
,
Queensland
Expedition Range,
1.I.1968
,
G.Hangay
, K 293223,
Hangay Collection
(AMS)
;
3 specimens
,
Queensland
,
R. Mus. Hist. Nat. Belg.
I.G. 12.595 (
IRSNB
)
;
7 specimens
,
Queensland
,
Muséum Paris
(
MNHN
)
;
9 specimens
,
Queensland
,
Muséum Paris
1935
Collection Pic
(
MNHN
)
;
1♀
,
1 specimen
,
Queensland
,
Muséum Paris
1935
Collection Sedillot
(
MNHN
)
;
12 specimens
,
Queensland
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
1 specimen
,
Queensland
, ex
Musaeo Van Lansberge
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
1 specimen
,
Queensland
, ex
Musaeo Möllenkamp
1913,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
2 specimens
,
Queensland
,
A.H.Elston
, K 293163 and K 293165 (AMS)
;
1 specimen
,
Queensland
, 1905,
Rolle
(
MSNM
)
;
1 specimen
,
Queensland
,
Nat. Mus
.
Victoria
COL-85880 (
NMVM
)
;
1 specimen
,
Queensland
, COL-86036 (
NMVM
)
;
1 specimen
,
Queensland
,
Nat.
Mus.
Victoria
,
C.French’s Collection
6.I.08
, COL-85879 (
NMVM
)
;
1 specimen
,
Queensland
,
Dr. K.K.Spence, K
293160,
K.K.Spence Collection
(AMS)
;
2 specimens
,
Queensland
, near
Silver Plains
homestead,
24.VI.1960
,
C.N.Smithers
, K 293246 and K 293247 (AMS)
;
1 specimen
, N
Queensland
,
E.W.Ferguson Collection
(
ANIC
)
;
2 specimens
, N
Queensland
,
H.J.Carter Collection
P.
20.IV.1922
, COL-85889 and COL-86035 (
NMVM
)
;
1 specimen
, som. S. of
Ayr
,
11.IX.1950
,
E.F.Riek
(
ANIC
)
;
1 specimen
,
Allinga
,
Chinchilla
,
26°41'S
,
150°38'E
,
Grace Lithgow
(
QMB
)
;
1 specimen
,
Amiens
,
16.V.1954
,
G.L.Gooding
,
C.G.L.Gooding Collection
, donated to
ANIC
1979
(
ANIC
)
;
1Ƌ,
1♀
,
Amiens
,
10 mi
.
W Stanthorpe
,
25.III.1975
,
H. & A.Howden
(
LBC
)
;
1 specimen
,
Biggenden
,
25.31ºS
,
152.03ºE
, W of
Bluff Range
,
18.VIII.1972
,
H.Frauca
(
ANIC
)
;
1 specimen
,
Blackall Ranges
,
X.1920
,
F.E.Wilson F.E.Wilson Collection
, COL-85904 (
NMVM
)
;
2 specimens
,
Blair Athol
,
VIII.1960
,
C.V., C.G.L.Gooding Collection
, donated to
ANIC
1979
(
ANIC
)
;
2 specimens
,
Boggom
19, via
Taroom
,
XI.1996
‒
I.1997
, Baited Flight Intercept, 0 30, Monteith & Cook (QMB);
1Ƌ,
Bottle Tree Scrub
,
Taroom-Cracow Road
,
25°33'S
,
150°08'E
,
14.IX.1996
,
D.Cook
, night hand collection 0 70, CM Reg. No. T 34263 (
QMB
)
;
2 specimens
,
Braemar State Forest
,
27°11'S
,
150°47'E
,
2–4.III.
2000
, 380 m, brigalow & open forest, 9193,
D.J.Cook
(
QMB
)
;
1Ƌ,
Braemar State Forest
, via
Kogan
,
15–19.X.1979
,
G.B.Monteith
& QM (
QMB
)
;
1 specimen
,
Brigooda
,
Koy Property
,
26°16'S
,
151°25'E
,
23.I.1995
,
G.Monteith
(
QMB
)
;
1 specimen
,
Bringalily Fire Tower
,
2 km
S,
28°12.6'S
,
151°07.5'E
,
21.XI.
2007
, 340 m, mv lamp brigalow, 15599,
A.Ewart
(
QMB
)
; 1 specimen, Brisbane, 1637, ex Collection Australian Museum (ANIC);
1 specimen
,
Brisbane
,
29.X.1917
,
H.Hacker
(
QMB
)
;
2 specimens
,
Brisbane
,
Griffith University
,
15.II.1982
,
T.J.Hawkeswood
(
ANIC
)
;
2 specimens
,
Lake Broadwater
, via
Dalby
,
22–25.II.1986
,
Site
6 and
Site
9,
G.Monteith
&
G.Thompson
(
QMB
)
;
1 specimen
,
Bulimba Creek
,
Carindale
,
3.XI.2003
, 15 m,
QM Party
, day hand colin, forest red gum woodland, 51652 (
QMB
)
; 3 specimens, Bundaberg, 331(handwritten), K 293164, A.H.Elston Collection (AMS); 1 specimen, Cairns, 1912, R.Sunter,
K 31981
(handwritten), K 293152 (AMS); 1 specimen, Cairns, H.J.Carter Collection P.
20.IV.1922
., COL-85887 (NMVM);
1 specimen
,
Cairns
,
VIII.1965
, G.R. (
ANIC
)
; 1 specimen, Cairns, 1952, J.G.Brooks, C.G.L.Gooding Collection, donated to ANIC 1979 (ANIC);
1 specimen
,
Cairns
,
Edge Hill
,
18.IV.1965
‒
14.V.1965
, at indoor light, J.G.Brooks (ANIC);
1 specimen
,
Cape York
, ex
Musaeo Mniszech
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
1Ƌ
Cape York
, (
MEL
)
;
2 specimens
,
Cape York
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
2 specimens
,
Mount Carbine
,
30.XII.1973
and
10.II.1975
, (
GTC
)
;
1 specimen
,
Mount Carbine
,
I.1972
(
MSNM
)
;
1 specimen
,
Mount Carbine
,
30.XII.1973
,
Collection Bomans
6531-a,
H.E.Bomans
BMNH
(E) 1999-248, 411720-3 (
NHM
)
;
2 specimens
,
Mount Carbine
,
6.VI.1945
,
T.W.Gamble
, K 293169 and K 293170, (AMS)
;
1 specimen
,
Mount Carbine
, near
Mossman
,
IV.1945
,
T.W.Gamble
, 11720-c,
H.E.Bomans
BMNH
(E) 1999-248, 406531-1 (
NHM
)
;
4 specimens
,
Mount Carbine
, near
Mossman
,
5.V.1945
and
IV.1946
,
T.W.Gamble
, K 293174, K 293178, K 293185, and K 293187,
Hangay Collection
(AMS)
;
2 specimens
,
10 km
W by N of
Mount Carbine
,
Desailly Creek
,
16.30ºS
,
144.55ºE
,
19–21.V.1981
, by sweeping,
A.Calder
(
ANIC
)
;
1 specimen
,
Carnarvon Range
,
2.I.1939
,
R.Geary
, K 293167 (AMS)
;
2 specimens
,
Carnarvon National Park
,
Rotary
shelter shed, 24°54'52'',
148°01'33''E
,
8–21.IV.1999
,
M.Neave
,
E.D.Edwards
,
M.Yee
, &
J.Cardale
(
ANIC
)
;
1 specimen
,
Carnarvon National Park
, section
Mount Moffatt
, 25°01'312''S, 147°53'572''E,
710 m
, A5,
V.Malý
(
QMB
)
;
2ƋƋ,
1♀
, same locality,
16–20.XII.2004
,
V.Malý
(
CMC
)
;
1 specimen
,
Carnarvon National Park
,
Mount Moffat
,
Top Moffat Camp
,
13–15.XII.1987
,
Monteith
,
Thomson
, &
Yeates
(
QMB
)
;
4 specimens
,
Carnarvon Range
,
II.1944
,
R.Geary
, K 293168, K 293242, K 293243, and K 293244 (AMS)
;
3 specimens
,
Carnarvon National Park
,
Chimneys
[?],
25°05'54''S
,
147°52'04''E
,
8– 21.IV.1999
,
M.Neave
,
E.D.Edwards
,
M.Yee
, &
J.Cardale
(
ANIC
)
; 1 specimen, Clermont, Dr. K.K.Spence K 293166, K.K.Spence Collection (AMS);
1 specimen
,
Clermont
,
VIII.1929
,
K.K.Spence
(
ANIC
)
;
2 specimens
,
Clermont
,
X.1962
,
C.V., C.G.L.Gooding Collection
, donated to
ANIC
1979
(
ANIC
)
;
4 specimens
,
Clermont
,
VIII.1928
,
Dr. K.K.Spence, K
293156, K 293157, K 293158, and K 293159,
K.K.Spence Collection
(AMS)
; 2 specimens, Caves near R’Ton (=Rockhampton), R.L.Higgins (QMB);
1 specimen
,
Coen
, Nat. Mus.
Victoria
,
C.French’s Collection
6.I.08
, COL-85970 (
NMVM
)
;
2 specimens
,
Cooktown
, (
MNHN
)
;
1 specimen
,
Cooktown
,
19.V.1945
,
C.Olive
, COL-85905 (
NMVM
)
;
1 specimen
,
Cooktown
,
V.1951
,
C.Oke
, COL-85959 (
NMVM
)
;
1 specimen
,
Cooktown
,
19.V.1953
,
C.Oke
, COL-85918 (
NMVM
)
;
1 specimen
,
Cooktown
,
30.VIII.1946
, 10 (
GTC
)
; 1 specimen, Cunnamulla, Dr. A.Breyer, 8114 (IRSNB); 1 specimen, Cunnamulla, H.Hardcaste COL-86011 (NMVM);
1 specimen
, Curtis Island,
4.I.1957
, COL-85958 (
NMVM
)
;
1 specimen
,
Dalby
,
10.III.1990
,
S.Lamond
, K 293313, R.deKeyzer
Insect Collection
(AMS)
;
4 specimens
,
Dalby
,
21.II.1987
(
GTC
)
;
3 specimens
,
27 km
SW
Dalby
,
30.X.1982
, under wood,
P.Schroeder
,
Graeme Krake Collection
, donated
Feb. 2007
, COL-86883, COL- 86884, and COL-86885 (
NMVM
)
;
7 specimens
,
Davies Creek
National Park
,
5–12.IV.1998
,
A.A.Calder
(
ANIC
)
;
1 specimen
,
Dayboro
,
0.5 km
S,
27°12'S
,
152°49'E
,
X.2004
, 60 m, mv lamp, 52147,
S.G.Wright
(
QMB
)
;
1Ƌ,
Duaringa
,
I.1946
,
Smith
, 11721-a,
H.E.Bomans
BMNH
(E) 1999-248 (
NHM
)
;
4 specimens
,
Duaringa
,
I.1946
‒
II.1946
, C.W.Smith K 293190, K 293194, K 293201, and 293204, Hangay Collection (AMS);
2 specimens
,
Duaringa
,
8.III.1946
,
L.A.Smith
, K 29 3205 and K 293208,
Hangay Collection
(AMS)
;
3 specimens
,
Duaringa
,
4.IX.1946
,
C.W.Smith
, K 293173, K 293209, and K 293210,
Hangay Collection
(AMS)
;
2 specimens
,
Duaringa
,
26.XII.1946
,
C.W.Smith
, K 293214 and K 293221,
Hangay Collection
(AMS)
;
10 specimens
,
Dunmore State Forest
,
Trig
4 area,
23.I.1999
, R.deKeyzer &
A.Sundholm
, K 293290, K 293291, K 293292, K 293293, K 293294, K 293295, K 293296, K 293297, K 293298, and K 293299, R.deKeyzer
Insect Collection
(AMS)
;
2 specimens
,
Edungalba
,
26.XII.1958
,
E.E.A., C.G.L.Gooding Collection
, donated to
ANIC
1979
(
ANIC
)
;
5 specimens
,
Edungalba
,
1 mi
. N of
Mourangee Station
,
22.V.1969
,
T.Campbell
&
E.Adams
(
ANIC
)
;
5 specimens
,
1 mi
. W by N of
Edungalba
, on
Mourangee Station
,
22.V.1969
, under log,
T.Campbell
&
E.Adams
(
ANIC
)
;
2 specimens
,
Eidsvold
,
X.1929
‒
IV.1930
, T.L.Bancroft (ANIC);
2 specimens
,
Emu Creek
Road
, near Emu
Vale
,
28–29.X.2002
, R.deKeyzer, K 293314 and K 293315, R.deKeyzer
Insect Collection
(AMS)
;
1 specimen
,
Endeavour N
Queensland
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
1 specimen
,
Endeavour River
, H.
Rolle,
Berlin
S.W. (
MEL
)
;
1 specimen
,
Endeavour River
,
Muséum Paris
1935
Collection Pic
(
MNHN
)
;
6 specimens
,
Endeavour River
, C.
French’s Collection,
6.I.08
,
Nat. Mus.
Victoria
COL-85849, COL-85886, COL-85968, COL-86046, COL- 86047, and COL-86049 (
NMVM
)
;
1 specimen
,
Mount Eumamurrin
,
26°09'S
,
148°41'E
,
14.XII.
2001
, 500 m,
Ironbark
/cypress,
G.B.Monteith
&
D.J.Cook
(
QMB
)
;
1 specimen
,
Expedition Range National Park
,
25°12'S
,
148°59'E
,
24–26.IX.1997
,
Ironbark
woodland, 5007
D. & L. & T. & H. Cook
(
QMB
)
;
2 specimens
,
50 km
W of
Mount Garnet
,
15.I.2003
, K 293252 and
24.I.2003
, K 293253,
J.Hasenpusch
, at mv light, R.deKeyzer
Insect Collection
(AMS)
;
1♀
,
Mount Garnet
, N
Queensland
,
20.II.2010
(
CMC
)
;
1 specimen
,
9 mi
E of
Gatton
,
27.33ºS
,
152.17ºE
,
5.XI.1969
,
S.Misko
(
ANIC
)
;
1 specimen
,
Glen Aplin
,
XI.1935
,
B.Stoyles
,
C.E.Clarke Collection
B.M. 1957-24, #607616 (
NHM
)
;
2 specimens
,
38 miles
N
Goondiwindi
,
23.VI.1965
,
D.P.Carne
(
ANIC
)
; 1Ƌ, Haay, Collection E.Candèze (IRSNB);
1 specimen
,
Hann Tableland Radar Station
,
16°55'S
,
145°15'E
,
8.II.1996
, 800‒
900 m
,
G.B.Monteith
(
QMB
)
;
1 specimen
,
Herberton
,
6.I.1990
,
T.Woodger
K 293259, R.deKeyzer
Insect Collection
(AMS)
;
3 specimens
,
Inglewood
,
41 km
NE,
28°19'S
,
151°27'E
,
25.X.1991
, under rotten log,
Tom Gush
, 3194,
Tom Gush
1 specimen
,
Ipswich, K
293151 (AMS)
;
7 specimens
,
7 km
N of
Jondaryan
,
25.XII.1982
,
J.T.Doyen
(
ANIC
)
;
1 specimen
,
24 mi
. SW by W of
Kingaroy
,
26.44ºS
,
151.31ºE
,
26.XI.1968
,
Britton
&
Misko
(
ANIC
)
; 2 specimens, Kuranda, 2/48, J.G.Brooks Bequest 1976 (ANIC);
2 specimens
,
Kuranda
,
21.V.1970
(
GTC
)
; 1 specimen, Kuranda, 1972 (GTC);
1 specimen
,
Lansdown Research Station
,
19.40ºS
,
146.51ºE
,
7 km
S of Woodstock
,
16.I.1974
, at light,
R.A.Barrett
(
ANIC
)
;
7 specimens
,
Leyburn
,
29.I.1974
,
J.Macqueen
(
ANIC
)
; 1 specimen, Lindeman I., Whitsunday Grp. Q.,
XII.1928
, Mel. Ward,
K 60547
(handwritten), K 293161 (AMS);
1 specimen
,
Mackay
,
5–20.V.1991
,
W.J.Vestgens
(
ANIC
)
; 1 specimen, Mackay, COL-85963 (NMVM);
1 specimen
,
Mackay
,
Turner
, 284,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
1Ƌ,
Mackay
,
25.X.1971
, 2326 (
MSNM
)
;
1 specimen
, Magnetic Island,
H.J.Carter
,
H.J.Carter Collection
P.
20.IV.1922
, COL-85882 (
NMVM
)
;
1 specimen
, Magnetic Island,
24.IV.1953
,
C.Oke
, COL-85923 (
NMVM
)
;
1 specimen
, Magnetic Island,
H.J.Carter
, COL-85928 (
NMVM
)
;
1 specimen
, Magnetic Island,
G.F.Hill
, ex
Collection Australian Museum
(
ANIC
)
;
5 specimens
, Magnetic Island,
VII.21
,
H.J.Carter
(
ANIC
)
;
8 specimens
, Magnetic Island,
12.V.1981
,
D.P.Carne
, in
Xanthorrhoea
(
ANIC
)
;
1Ƌ,
Mount Malloy
,
7.I.1975
(
LBC
)
;
3 specimens
,
Mount Malloy
,
Station Creek
,
18.
II.1969
, X219,
J.G.Brooks
(
ANIC
)
;
4 specimens
,
N
Mount Malloy
,
Station Creek
,
11 m
,
2.XII.1972
,
J.A.G.Brooks
, 180 (
ANIC
)
; 1Ƌ, Mareeba, 6535-a, H.E.Bomans BMNH(E) 1999-248 (NHM); 1 specimen, Mareeba (GTC);
6 specimens
,
Mareeba
,
4–5.XII.1972
(
GTC
)
;
1Ƌ,
Mary Creek
,
16.33ºS
,
145.12ºE
,
4.XII.1968
, open forest at light,
E.Britton
&
S.Misko
, 8638,
H.E.Bomans
BMNH
(E) 1999-248 (
NHM
)
;
1 specimen
,
Millmerran
,
6.II.1974
(
ANIC
)
;
12 specimens
,
Millmerran
,
VII.1949
‒
VIII.1949
, G.L.Gooding, C.G.L.Gooding Collection, donated to ANIC 1979 (ANIC);
1 specimen
,
Morehead
,
10.I.1988
,
M.S.Moulds
, K 293249, R.deKeyzer
Insect Collection
(AMS)
;
2 specimens
, Moreton Island,
IV.1916
,
E.Allen
(
QMB
)
;
1 specimen
,
Moreton Bay
,
Waterhouse
, ex
Musaeo Van Lansberge
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
8 specimens
,
Mourangee
, near
Edungalba
,
50 mi
. SW of
Rockhampton
,
27.XI.1968
, under logs,
Britton
,
Misko
&
Adams
(
ANIC
)
;
2 specimens
,
Mutchilba
,
X.1959
,
A.D.Selby F.E.Wilson Collection
, COL-86003 (
NMVM
)
;
2 specimens
, National Park
Queensland
,
I.1928
,
H.J.Carter
,
K 60790
(handwritten), K 293154 and K 293155 (AMS)
;
2 specimens
,
20 km
SW of
Nebo
,
8.I.2006
,
T.M.S.Hanlon
, to light at night, K 293225 (AMS)
;
2 specimens
,
Nipping Gully
,
25°42'S
,
151°26'E
,
25.I.1996
,
Site
6, 240 m
, at mv light, open forest, 7442 and 7460,
Monteith
,
Gough
, &
Thompson
(
QMB
)
;
1 specimen
,
Nipping Gully
,
18–19.XII.
1998
, 200 m, 7532,
G.Monteith
,
C.Gough
, &
G.Maywald
(
QMB
)
;
1 specimen
,
North West
Island,
23.298°S
,
151.704°E
,
11.X.2008
,
Site
5, 17597, QM/QP WS party,
0.5 m
,
Day
hand,
Pisonia forest
Photog. Spm, PS
1174 (
QMB
)
;
2 specimens
,
Palm Creek
C.A.
, J.W.Hanks COL-85904 and COL-85904 (
NMVM
)
;
3 specimens
,
Parmentier Creek
,
10 km
W of Kuranda
,
Mareeba Road
,
26.II.1970
, m.v.,
R.Hardie
,
University
of New England Collection, donated 1983 (
ANIC
)
;
1 specimen
,
Prairie
,
VII.1904
,
W.Froggatt Collection
(
ANIC
)
;
2 specimens
,
13 km
N of
Quinalow
,
25.XII.1987
,
J.T.Doyen
(
ANIC
)
;
2 specimens
,
Rathdowney
,
28.12ºS
,
152.51ºE
,
21.XI.1982
(E.-Y: AU-82), dry logs on meadow,
Endrödy-Younga
(
TMP
)
;
1 specimen
,
Robert Wicks Research Station
,
1.3 km
SW,
28°17.9'S
,
151°08.4'E
,
22.XI.
2007
, 310 m, mv light, 15598,
A.Ewart.
(
QMB
)
;
1 specimen
,
Rockford Scrub
,
20°07.0'S
,
146°37.8'E
,
10–11.XII.2006
,
Site
1, 270 m
, vinescrub, bendee, hand collect,
Monteith
,
Cook
&
White
,
Reg. No T
147163 (
QMB
)
;
1 specimen
,
Rockford Scrub
,
20°06'S
,
146°37.1'E
,
11.II.2007
,
Site
5, 300 m
, QM party night hand collect, vinescrub, lancewood/bendee,
Monteith
,
Cook
, &
White
,
Reg. No T
147164 (
QMB
)
;
3 specimens
,
Rockhampton
,
III.1959
,
C.V., C.G.L.Gooding Collection
, donated to
ANIC
1979
(
ANIC
)
;
2 specimens
,
Rockhampton
,
XII.1964
,
C.V., C.G.L.Gooding Collection
, donated to
ANIC
1979
(
ANIC
)
;
1 specimen
,
Rockhampton
,
4.I.1999
,
T.M.S.Hanlon
, to light in town, K 293264, R.deKeyzer
Insect Collection
(AMS)
;
31 specimens
,
Rockhampton
,
Thozet
1870, 257 (
MNHN
)
;
1 specimen
,
Rockhampton
,
Meek
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
1 specimen
,
33 mi
. SW of
Rockhampton
,
26.XI.1968
,
Britton
&
Misko
(
ANIC
)
;
1 specimen
,
Mount Rose
, via
Taroom
,
25°25'S
,
149°58'E
,
15–16.XII.
1997
, 260 m, vine scrub remnant, 5057,
D. & L.Cook
(
QMB
)
;
1 specimen
,
5 km
W by N of
Rounded Hill
,
15.17ºS
,
145.13ºE
,
5–6.X.1980
,
T.Weir
(
ANIC
)
;
1 specimen
,
1 km
N of
Rounded Hill
,
15.17ºS
,
145.10ºE
,
7.XII.1980
,
T.Weir
(
ANIC
)
; 1 specimen, Scotts Peak, SE slopes, 22°51.7'S, 148°13.5'E,
9.III.
2006
, 480 m, day hand collect, rocky scree, Wright & Burwell, 13343, QM Reg. No T137257 (QMB);
1 specimen
,
Shipton’s Flat
,
15.47ºS
,
145.14ºE
,
17–19.X.1980
,
T.Weir
(
ANIC
)
;
1 specimen
,
Silver Valley
III,
R. Mus. Hist. Nat. Belg.
I.G. 12.595 (
IRSNB
)
;
1Ƌ,
Silver Valley
III (
MNHN
)
;
1 specimen
,
Somerset
Dam,
23.XII.1912
, at light,
R.J.Kohout
(
ANIC
)
;
1 specimen
,
Springsure
,
24.VII.1975
,
J.C.Lesouef.
(
ANIC
)
;
1 specimen
,
St Lucia
,
Brisbane
,
27.X.1984
, R.deKeyzer, K 293312, R.deKeyzer
Insect Collection
(AMS)
;
1 specimen
,
Stanthorpe
,
17.V.1923
, 0 91 (
QMB
)
;
3 specimens
,
Stanthorpe
,
19.X.1926
,
29.IX.1927
,
7.II.1929
,
E.Sutton
,
E.Sutton Collection Don.
-
Dec. 1964
(QMB);
1 specimen
,
Stanthorpe
,
I.1928
,
H.J.Carter
,
K 60713
(handwritten), K 293153 (AMS)
;
1 specimen
,
Stanthorpe
,
Muséum Paris
1935
Collection Pic
(
MNHN
)
;
3 specimens
,
Stanthorpe
,
5.II.1974
(
ANIC
)
; 3 specimens, Stanthorpe, E.Sutton, F.E.Wilson Collection, COL-85865, COL-85888, and COL-85895 (NMVM);
1 specimen
,
Station Creek
,
10 m
,
NW Cairns Cook Highway
,
28.II.1970
, m.v.,
R.Hardie
(
ANIC
)
;
1 specimen
,
Stockhaven
,
3 km
SE,
25°48'S
,
150°01'E
,
10.X.‒19.XII.
1998
, 450 m, rainforest intercept, 7516,
G.Monteith
&
C.Gough
(
QMB
)
;
1 specimen
,
Tara
,
16– 18.XII.2010
,
A.Urban
&
C.Dale
, K 293300, R.deKeyzer
Insect Collection
(AMS)
;
4 specimens
,
Taroom District
,
25°33'S
,
150°08'E
,
14.I.1997
,
Bottle
tree scrub, 210,
Burwell
,
Cook
&
Janetzki
(
QMB
)
;
1 specimen
,
Taroom District
,
Boggomoss No
19,
25°25'S
,
150°01'E
,
18.VI.1996
,
H.Janetzki
, QM Reg. No T34266 (
QMB
)
;
2 specimens
,
Taroom District
,
Boggomoss No
4,
25°25'S
,
150°01'E
,
9.IX.‒11.XII.1996
,
Pitfall
0 96, QM Reg. No T34264 and
XI.1996
‒
I.1997
, Pitfall 0 97, QM Reg. No T34265, P.Lawless, (QMB);
1 specimen
,
Taroom
,
6 km
N on
Highway
,
25°36'S
,
149°46'E
,
21.IX.‒17.XII.
1997
, 200 m, 5047,
G.Monteith
(
QMB
)
;
2 specimens
,
The Palms
via
Cooyar
,
25°29'S
,
149°42'E
,
6.XII.1992
,
G.J.Ingram
(
QMB
)
;
1 specimen
,
Toomba High School
(
Queensland
),
19°58.2'S
,
145°35.2'E
,
13–15.II.
2007
, 390 m, QM party dayhand, pasture, 14798,
QM Reg. No T
147165 (
QMB
)
; 1 specimen, Townsville, E.W.Ferguson Collection (ANIC); 1 specimen, Townsville, F.E.Wilson Collection, COL- 85864 (NMVM);
3 specimens
,
7 km
N of
Wallangarra
,
27.XII.1982
,
J.T.Doyen
(
ANIC
)
;
1 specimen
,
Warrego Highway
,
23 km
E Toowoomba
,
28.VII.1995
,
F.Douglas
, COL-86006 (
NMVM
)
;
2 specimens
,
Warrego Highway
,
25 km
E Toowoomba
,
28.VII.1995
,
F.Douglas
, ex pupa
12.VIII.1996
, K 293317 and K 293318, R.deKeyzer
Insect Collection
(AMS)
;
4 specimens
,
24 km
S of
Warwick
,
26.XII.1982
,
J.T.Doyen
(
ANIC
)
;
1 specimen
,
Westwood
, near
Rockhampton
,
22.II.
1924
, 500 feet,
G.H.Wilkins
, W
Australia
Exp.
, B.M. 1924-293, # 607612 (
NHM
)
;
1 specimen
,
Yarramulla
,
18
°
13'S
,
144°40'E
,
8–10.II.2003
, mv light, 11260,
G.B.Monteith
(
QMB
)
;
4 specimens
,
Yuleba State Forest
,
26°58'S
,
149°44'E
,
8–10.III.2002
,
Site
1, 270 m
, cypress/box, 10495,
Monteith
,
Cook
, &
Wright
(
QMB
)
;
1 specimen
,
Yuleba State Forest
,
26°58'S
,
149°45'E
,
8–10.III.2002
,
Site
2, 207 m
, 10496,
Monteith
,
Cook
, &
Wright
(
QMB
)
;
1 specimen
,
Yeppoon
,
3.III.1935
,
S.R.E. Brock Collection
, donated to
ANIC
1987
(
ANIC
)
.
SOUTH AUSTRALIA
:
5 specimens
,
South Australia
, Nat. Mus
.
Victoria
COL-85856, COL- 85860, COL-85862, COL-85909, and COL-85899 (
NMVM
)
;
1 specimen
,
South Australia
,
A.H.Elston
, A. K 293228,
H.Elston Collection
(AMS)
;
1 specimen
,
South Australia
,
G.H.Dalton
, ex
Collection Australian Museum
(
ANIC
)
; 1 specimen, Adelaide, 1867, Collection Castelnau (MSNG); 1 specimen, NW Davey, F.E.Wilson Collection, COL-85893 (NMVM);
1 specimen
,
Port Lincoln
,
K 28027
(handwritten), K 293226 and K 293227 (
Old Collection
) (AMS)
;
1 specimen
,
Willalo
,
IV.1942
,
H.Mincham
, K 293229 (AMS)
.
VICTORIA
:
1 specimen
,
Victoria
, ex
Musaeo Van Lansberge
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
1 specimen
,
Victoria
,
Muséum Paris
1936
Collection A.
Boucomont (
MNHN
)
;
1 specimen
,
Victoria
(
MNHN
)
;
1 specimen
,
Victoria
,
Muséum Paris
1935
Collection Sedillot
(
MNHN
)
;
1 specimen
,
Victoria
, COL-85964 (
NMVM
)
;
5 specimens
,
Victoria
,
J.E.Dixon
, COL-85930, COL-85931, COL-85932, COL-85933, and COL-85934 (
NMVM
)
;
1 specimen
, NW
Victoria
, 1934,
De Boulay, C.E.Clarke Collection B.M.
1957-24, # 607592 (
NHM
)
;
1 specimen
,
Cavendish
,
I.1950
,
B.Given
,
F.E.Wilson Collection
, COL-85162 (
NMVM
)
;
1 specimen
,
Echuca
, (
MNHN
)
; 2 specimens, Emerald, C.Oke, COL-85877 and COL-85897 (NMVM);
1 specimen
,
Ferntree Gully
,
XII.1920
,
C.Oke
, COL- 85927 (
NMVM
)
; 1 specimen, Grampian Ranges, COL-86034 (NMVM); 1 specimen, Gunbower, COL-85171 (NMVM); 2 specimens, Hattah, C.Oke, COL-86039 and D.Best, COL-85876 (NMVM); 1♀, Hattah, C.Oke, COL- 85854 (NMVM);
1 specimen
,
Hattah
,
6.X.1971
, 1335B, COL-86038 (
NMVM
)
;
1 specimen
,
Horsham
,
15.II.1945
, RB, COL-86043 (
NMVM
)
; 1 specimen, Kiata, 1335A, COL-86037 (NMVM);
1♀
,
1 specimen
,
Lake Hattah
,
XI.1924
,
C.Oke
, COL-85960 and COL-85926 (
NMVM
)
;
1 specimen
,
Kulkine Forest
, NW
Victoria
,
29.VI.1968
,
G.W.Anderson
, 8635,
H.E.Bomans
BMNH
(E) 1999-248, 408635 (
NHM
)
;
1 specimen
,
Linga
,
IV.1922
,
F.E.Wilson
,
F.E.Wilson Collection
, COL-85902 (
NMVM
)
;
1 specimen
,
Melbourne
, 24.63,
Muséum Paris
environs
de Melbourne
,
Miller
1863 (
MNHN
)
;
7 specimens
, environs
de Melbourne
,
Miller
1863, 2463 (
MNHN
)
;
1 specimen
,
Warburton
,
25.II.1922
,
C.Oke
, COL-85920 (
NMVM
)
.
WESTERN
AUSTRALIA
:
4 specimens
,
Australia
Orient, v. Müller 2-96, Muséum Paris (
MNHN
)
; 2 specimens, Fortescue, Hamersley Range, W.D.Dodd, ex Collection Australian Museum (ANIC).
TASMANIA
:
2 specimens
,
Tasmanie
, 4-46, Muséum Paris
Australie
,
J.Verreaux
1846 (
MNHN
)
;
1 specimen
,
Tasmanie
, Muséum Paris
Australie
,
J.Verreaux
2-47 (
MNHN
)
.
Distribution
.
Australia
:
Australian
Capital
Territory
,
New South Wales
,
Northern
Territory
,
Queensland
,
Victoria
,
South
Australia
,
Western
Australia
,
Tasmania
(?); New
Guinea
:
Papua New Guinea
; Moluccan Archipelago: Aru Islands (
Bartolozzi
et al.,
2011
).
Natural history data:
The adults have been collected year round, but mainly from November to February. They are most commonly found in rotten logs, in dead wood, under bark of rotten logs, rocky scree, and also in urbanized areas at light (Rockhampton,
I.1999
). The elevation ranges from
10 to
710 m
.
Hawkeswood (1985
,
1988
,
2014
) lists some host plants for the species:
Angophora bakeri
(Myrtaceae)
,
Camptostemon
(Bombacaceae)
,
Eucalyptus
(Myrtaceae)
, and
Xanthorrhoea johnsonii
(Xanthorrhoeaceae)
. Some data on host plants and habitats were obtained from labels, including brigalow (
Acacia harpophylla
,
Fabaceae
), bendee (
Acacia catenulata
,
Fabaceae
),
Banksia
(Proteaceae)
,
Spinifex
(Poaceae)
grassland, forest red gum woodland,
Xanthorrhoea australis
(Xanthorrhoeaceae)
, ironbark/cypress woodland, Mallee woodland, and vine scrub. Collecting methods include flight intercept trap and light trapping (mainly mv light, rarely uv light). Two pupae were collected in
July 1995
from which two adults emerged in
August 1996
. The larvae are unknown.
Remarks
.
We dissected the male of
F. laevithorax
labeled as "
Type
", and compared the genitalia with
F. regularis
. We found no significant differences between
F. laevithorax
and
F. re g ul ar i s
, both in external and in genitalia characters, except minor differences falling within the range of individual variations. Therefore we consider
F. levithorax
as a junior synonym of
F. regularis
(
new synonymy
).
We found in the collection of MNHN three old specimens labeled “Tasmanie” (J. Verreaux leg., 1846), but this locality is probably erroneous, since we did never see any other specimen of this species from Tasmania.