<document id="1E7302108E7BB171253FF8503DF05275" ID-ISSN="2251-8169" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665287058592" checkinUser="felipe" docAuthor="Benavides-Montaño, Javier Antonio, Betancourt-Echeverri, Jesus Antonio, Valencia, Gustavo López &amp; Mesa-Cobo, Nora Cristina" docDate="2022" docId="03E45B658939E27EE04CFCA12A23FB53" docLanguage="en" docName="PersJourAcarology.11.3.397–437.pdf" docOrigin="Persian Journal of Acarology 11 (3)" docStyle="DocumentStyle:B3904D664642C9C0DBE4CD09E74105CD.9:PersJourAcarology.2017-.journal_article" docStyleId="B3904D664642C9C0DBE4CD09E74105CD" docStyleName="PersJourAcarology.2017-.journal_article" docStyleVersion="9" docTitle="Amblyomma varium Koch 1844" docType="treatment" docVersion="2" lastPageNumber="411" masterDocId="FFDD231D8937E270E20EFFBB2824FFB9" masterDocTitle="A review of hard ticks (Acari: Ixodidae) in Colombia: The risk of tick-borne diseases" masterLastPageNumber="437" masterPageNumber="397" pageNumber="411" updateTime="1699379006652" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-2.0">
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<mods:title id="A0BD205081CF8FEE0EE86283BA32FA8A">A review of hard ticks (Acari: Ixodidae) in Colombia: The risk of tick-borne diseases</mods:title>
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<mods:namePart id="755A3FAE8F4AD8296852B5B94CCF5266">Benavides-Montaño, Javier Antonio</mods:namePart>
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<mods:namePart id="E5D85497D0A6DAC51C0EC6D8EEADE65D">Betancourt-Echeverri, Jesus Antonio</mods:namePart>
<mods:affiliation id="8B47E442726F679507B1DD551056BCCA">. School of Microbiology and Bioanalysis, Faculty of Health, Industrial Universidad de Santander, Bucaramanga, Colombia; E-mail: jesantbet @ yahoo. com</mods:affiliation>
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<mods:namePart id="F4D8118FF1247088F8B0A412A984B062">Valencia, Gustavo López</mods:namePart>
<mods:affiliation id="C61765E4F600448736D5F78E1D497E83">. Asesor Científico Instituto Colombiano de Medicina Tropical, ICMT, Universidad CES, Medellín, Colombia; E-mail: glopez @ ces. edu. co</mods:affiliation>
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<mods:namePart id="32E88930CDAE8B3F1FE5A9E1AD1D8043">Mesa-Cobo, Nora Cristina</mods:namePart>
<mods:affiliation id="E6F2F53C3DA65BC0AD4D459E8BBD7582">. Acarology Research group, Immunology and infectious disease group, Agriculture Department, Faculty of Agriculture, Universidad Nacional de Colombia-Palmira, Palmira, Colombia; E-mail: ncmesac @ unal. edu. co</mods:affiliation>
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<taxonomicName id="4C4D91F08939E27EE04CFCA12BD5FC88" ID-CoL="CMK8" authority="Koch, 1844" authorityName="Koch" authorityYear="1844" box="[578,1009,794,817]" class="Arachnida" family="Ixodidae" genus="Amblyomma" kingdom="Animalia" order="Ixodida" pageId="14" pageNumber="411" phylum="Arthropoda" rank="species" species="varium">
<emphasis id="B93936618939E27EE04CFCA12BD5FC88" bold="true" box="[578,1009,794,817]" pageId="14" pageNumber="411">
<emphasis id="B93936618939E27EE04CFCA12B74FC88" bold="true" box="[578,848,794,817]" italics="true" pageId="14" pageNumber="411">Amblyomma varium</emphasis>
Koch, 1844
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</paragraph>
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It is reported from 
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, 
<collectingCountry id="F35AAAE38939E27EE059FCD82A8CFCC3" box="[599,680,867,890]" name="Brazil" pageId="14" pageNumber="411">Brazil</collectingCountry>
, 
<collectingCountry id="F35AAAE38939E27EE0B9FCD82B24FCC3" box="[695,768,867,890]" name="Chile" pageId="14" pageNumber="411">Chile</collectingCountry>
, 
<collectingCountry id="F35AAAE38939E27EE103FCD82BAAFCC3" box="[781,910,867,890]" name="Colombia" pageId="14" pageNumber="411">Colombia</collectingCountry>
, 
<collectingCountry id="F35AAAE38939E27EE192FCD82C08FCC3" box="[924,1068,867,890]" name="Costa Rica" pageId="14" pageNumber="411">Costa Rica</collectingCountry>
, 
<collectingCountry id="F35AAAE38939E27EE634FCD82CDFFCC3" box="[1082,1275,867,890]" name="French Guiana" pageId="14" pageNumber="411">French Guiana</collectingCountry>
, 
<collectingCountry id="F35AAAE38939E27EE704FCD82DB0FCC3" box="[1290,1428,867,890]" name="Guatemala" pageId="14" pageNumber="411">Guatemala</collectingCountry>
, 
<collectingCountry id="F35AAAE38939E27EE299FC33293AFC26" box="[151,286,904,927]" name="Nicaragua" pageId="14" pageNumber="411">Nicaragua</collectingCountry>
, 
<collectingCountry id="F35AAAE38939E27EE33EFC3329BCFC26" box="[304,408,904,927]" name="Panama" pageId="14" pageNumber="411">Panama</collectingCountry>
, 
<collectingCountry id="F35AAAE38939E27EE3A5FC3329CDFC26" box="[427,489,904,927]" name="Peru" pageId="14" pageNumber="411">Peru</collectingCountry>
, and 
<collectingCountry id="F35AAAE38939E27EE039FC332AE6FC26" box="[567,706,904,927]" name="Venezuela" pageId="14" pageNumber="411">Venezuela</collectingCountry>
; (
<bibRefCitation id="EFDC97828939E27EE0D1FC332BE8FC26" author="Jones, E. K. &amp; Clifford, C. M. &amp; Keirans, J. E. &amp; Kohls, G. M." box="[735,972,904,927]" pageId="14" pageNumber="411" pagination="1 - 40" refId="ref22147" refString="Jones, E. K., Clifford, C. M., Keirans, J. E. &amp; Kohls, G. M. (1972) The ticks of Venezuela (Acarina: Ixodoidea) with a key to the species of Amblyomma in the western hemisphere. Brigham Young University Science Bulletin, 17 (4): 1 - 40." type="journal article" year="1972">
Jones 
<emphasis id="B93936618939E27EE13AFC332B56FC26" box="[820,882,904,927]" italics="true" pageId="14" pageNumber="411">et al</emphasis>
. 1972
</bibRefCitation>
; 
<bibRefCitation id="EFDC97828939E27EE1D0FC332D02FC26" author="Guglielmone, A. A. &amp; Estrada-Pena, A. &amp; Keirans, J. E. &amp; Robins, R. G." box="[990,1318,904,927]" pageId="14" pageNumber="411" refId="ref21404" refString="Guglielmone, A. A., Estrada-Pena, A., Keirans, J. E. &amp; Robins, R. G. (2003) Ticks (Acari: Ixodida) of the Neotropical Zoogeographic Region. International Consortium on Ticks and Tick-Borne Diseases (ICTTD- 2). Atlanta, Houten, The Netherlands, 173 pp." type="book" year="2003">
Guglielmone 
<emphasis id="B93936618939E27EE69DFC332CF4FC26" box="[1171,1232,904,927]" italics="true" pageId="14" pageNumber="411">et al</emphasis>
. 2003
</bibRefCitation>
; BarrosBattesti 
<emphasis id="B93936618939E27EE30AFC16291AFC7D" box="[260,318,941,964]" italics="true" pageId="14" pageNumber="411">et al</emphasis>
. 2006; 
<bibRefCitation id="EFDC97828939E27EE3ABFC162A81FC7D" author="Onofrio, V. &amp; Labruna, M. &amp; Pinter, A. &amp; Giacomin, F. &amp; Barros-Battesti, D. M." box="[421,677,941,964]" pageId="14" pageNumber="411" pagination="53 - 113" refId="ref25842" refString="Onofrio, V., Labruna, M., Pinter, A., Giacomin, F. &amp; Barros-Battesti, D. M. (2006) Comentarios e chaves para as especies do genero Amblyomma. In: Barros-Battesti, D. M., Arzua, M. &amp; Bechara, G. (Eds.), Carrapatos de importancia medico veterinaria da regiao neotropical. Um guia ilustrado para identificacao de especies. Sao Paulo: Butantan, pp. 53 - 113." type="book chapter" year="2006">
Onofrio 
<emphasis id="B93936618939E27EE01BFC162A74FC7D" box="[533,592,941,964]" italics="true" pageId="14" pageNumber="411">et al</emphasis>
. 2006
</bibRefCitation>
). 
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(anteater), armadillos and sloths are the main hosts. This tick has been reported in 
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on sloths, 
<taxonomicName id="4C4D91F08939E27EE1CFFC692D4FFC50" authority="(Lopez 2017)" baseAuthorityName="Lopez" baseAuthorityYear="2017" box="[961,1387,978,1001]" class="Mammalia" family="Bradypodidae" genus="Bradypus" kingdom="Animalia" order="Pilosa" pageId="14" pageNumber="411" phylum="Chordata" rank="species" species="griseus">
<emphasis id="B93936618939E27EE1CFFC692C83FC50" box="[961,1191,978,1001]" italics="true" pageId="14" pageNumber="411">Bradypus griseus</emphasis>
(Lopez 2017)
</taxonomicName>
, in 
<taxonomicName id="4C4D91F08939E27EE299FC4D2980FBB4" authorityName="Linnaeus" authorityYear="1758" box="[151,420,1014,1037]" class="Mammalia" family="Bradypodidae" genus="Bradypus" kingdom="Animalia" order="Pilosa" pageId="14" pageNumber="411" phylum="Chordata" rank="species" species="tridactylus">
<emphasis id="B93936618939E27EE299FC4D2980FBB4" box="[151,420,1014,1037]" italics="true" pageId="14" pageNumber="411">Bradypus tridactylus</emphasis>
</taxonomicName>
in Condoto – 
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, and in 
<collectingCountry id="F35AAAE38939E27EE112FC4D2BA0FBB4" box="[796,900,1014,1037]" name="Panama" pageId="14" pageNumber="411">Panama</collectingCountry>
, where it is commonly found parasitizing 
<taxonomicName id="4C4D91F08939E27EE299FBA0298FFB8B" authorityName="Peters" authorityYear="1858" box="[151,427,1051,1074]" class="Mammalia" family="Megalonychidae" genus="Choloepus" kingdom="Animalia" order="Pilosa" pageId="14" pageNumber="411" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="B93936618939E27EE299FBA0298FFB8B" box="[151,427,1051,1074]" italics="true" pageId="14" pageNumber="411">Choloepus hoffmanni</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFDC97828939E27EE3B0FBA02A9CFB8B" author="Osorno-Mesa, E." box="[446,696,1051,1074]" pageId="14" pageNumber="411" pagination="57 - 103" refId="ref26001" refString="Osorno-Mesa, E. (1942) Las garrapatas de la republica de Colombia. Revista Facultad Nacional de Agronomia, 5: 57 - 103." type="journal article" year="1942">Osorno-Mesa 1942</bibRefCitation>
; 
<bibRefCitation id="EFDC97828939E27EE0C8FBA02B49FB8B" author="WRAMC &amp; Washington, D. C. &amp; Washington, DC" box="[710,877,1051,1074]" pageId="14" pageNumber="411" refId="ref29624" refString="WRAMC (1998) Walter Reed Army Medical-Center. Disease vector ecology profile. Colombia. U. S., Armed Forces Pest Management Board. Forest Glen Section. Washington, D. C. Washington, DC, Defense Pest Management Information Analysis Center, Armed Forces Pest Management Board, Forest Glen Section, 88 pp." type="book" year="1998">Wramc 1998</bibRefCitation>
; 
<bibRefCitation id="EFDC97828939E27EE171FBA02C07FB8B" author="Voltzit, O. V." box="[895,1059,1051,1074]" pageId="14" pageNumber="411" pagination="3 - 134" refId="ref29431" refString="Voltzit, O. V. (2007) A review of neotropical Amblyomma species (Acari: Ixodidae) Acarina, 15 (1): 3 - 134." type="journal article" year="2007">Voltzit 2007</bibRefCitation>
). Recently, a new update has mentioned its presence on 
<emphasis id="B93936618939E27EE3F8FBFB2A92FBEE" box="[502,694,1088,1111]" italics="true" pageId="14" pageNumber="411">
C. l. 
<taxonomicName id="4C4D91F08939E27EE034FBFB2A92FBEE" baseAuthorityName="Lopez" baseAuthorityYear="2017" box="[570,694,1088,1111]" class="Gastropoda" family="Tomichiidae" genus="Coxiella" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="411" phylum="Mollusca" rank="species" species="familiaris">familiaris</taxonomicName>
</emphasis>
from Ibague, 
<collectingRegion id="498924918939E27EE17DFBFB2BF4FBEE" box="[883,976,1088,1111]" country="Colombia" name="Tolima" pageId="14" pageNumber="411">Tolima</collectingRegion>
(
<bibRefCitation id="EFDC97828939E27EE1ECFBFB2D0CFBEE" author="Rivera-Paez, F. A. &amp; Labruna, M. B. &amp; Martins, T. F. &amp; Perez, J. E. &amp; Castano-Villa, G. J. &amp; Ossa-Lopez, P. A. &amp; Gil, C. A. &amp; Sampieri, B. R. &amp; Aricapa-Giraldo, H. J. &amp; Camargo-Mathias, M. I." box="[994,1320,1088,1111]" pageId="14" pageNumber="411" pagination="57 - 66" refId="ref27050" refString="Rivera-Paez, F. A., Labruna, M. B., Martins, T. F., Perez, J. E., Castano-Villa, G. J., Ossa-Lopez, P. A., Gil, C. A., Sampieri, B. R., Aricapa-Giraldo, H. J. &amp; Camargo-Mathias, M. I. (2018 a) Contributions to the knowledge of hard ticks (Acari: Ixodidae) in Colombia. Ticks and Tick-Borne Diseases, 9: 57 - 66." type="journal article" year="2018">
Rivera-Paez 
<emphasis id="B93936618939E27EE685FBFB2CE2FBEE" box="[1163,1222,1088,1111]" italics="true" pageId="14" pageNumber="411">et al</emphasis>
. 2018a
</bibRefCitation>
). Except for these few reports in 
<collectingCountry id="F35AAAE38939E27EE3D1FBDE2A45FBC5" box="[479,609,1125,1148]" name="Colombia" pageId="14" pageNumber="411">Colombia</collectingCountry>
, nothing is known about the biology and ecology of this tick species, hence the need for research that describes it thoroughly; it is likely that the specimens reported by Rivera on a pet dog were an accidental parasitism since this species feeds mostly on sloths (
<bibRefCitation id="EFDC97828939E27EE2FCFB6829D2FB53" author="Marques, S. &amp; Barros-Battesti, D. M. &amp; Faccini, J. L. &amp; Onofrio, V. C." box="[242,502,1235,1258]" pageId="14" pageNumber="411" pagination="1141 - 1146" refId="ref23883" refString="Marques, S., Barros-Battesti, D. M., Faccini, J. L. &amp; Onofrio, V. C. (2002) Brazilian distribution of Amblyomma varium Koch, 1844 (Acari: Ixodidae), a common parasite of sloths (Mammalia: Xenarthra). Memorias do Instituto Oswaldo Cruz, 97: 1141 - 1146." type="journal article" year="2002">
Marques 
<emphasis id="B93936618939E27EE362FB682980FB53" box="[364,420,1235,1258]" italics="true" pageId="14" pageNumber="411">et al</emphasis>
. 2002
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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