Synopsis of the tribe Stipeae (Poaceae) in Nepal
Author
Nobis, Marcin
Author
Gudkova, Polina D.
Author
Pendry, Colin A.
text
PhytoKeys
2019
128
97
119
http://dx.doi.org/10.3897/phytokeys.128.34637
journal article
http://dx.doi.org/10.3897/phytokeys.128.34637
1314-2003-128-97
CFE1800CC09C53E78AFCB0D1E6864E45
3378135
Achnatherum
staintonii (Bor) M.Nobis & P.D.Gudkova, comb. nov.
Stipa staintonii
Bor, Bull. Bot. Surv. India 7: 133 (1965) (Basionym).
≡
Stipella staintonii
(Bor)
Roeser
& Hamasha, Pl. Syst. Evol. 298: 365 (2012), nom. inval.;
≡
Stipellula staintonii
(Bor)
Roeser
& H.R. Hamasha,
Schlechtendalia
24: 92 (2012).
Type.
Nepal, near Seng Khola, 12500 ft [3810 m.], exposed cliffs, 4 Okt 1954,
Stainton, Sykes & Williams 4677
(holotype K!, isotype BM!).
General distribution.
Nepal (endemic;
Bor 1965
;
Freitag 1985
).
Distribution in Nepal.
Baglung, Dolpa, Manang, Mustang, Rukum.
Habitat.
open rocky or stony sandy slopes and scrublands.
Altitude.
3000-4200 m.
Note.
Although
Achnatherum staintonii
has been confused with
Stipa przewalskyi
, the latter species does not occur in Nepal.
Achnatherum staintonii
is easily distinguished from
Stipa przewalskyi
by having maize-like vs. saw-like LEPs and in having unequal glumes and distinctly longer lemma than palea vs. glumes as well as lemma and palea subequal, respectively.
Roeser
(2012)
transferred five species of
Stipa
, including
Stipa staintonii
, into their new genus
Stipellula
on the basis of his earlier molecular analysis (
Hamasha et al. 2012
).
Stipellula
is characterized by its maize-like lemma epidermal pattern which clearly distinguishes it from
Stipa
and confirms that these species belong to the achnatheroid group of grasses within the
Stipeae
. However, there are no unique, diagnostic morphological characters to separate
Stipellula
from
Achnatherum
which is itself polymorphic and highly polyphyletic (
Romaschenko et al. 2012
;
Hamasha et al. 2012
). Thus, we prefer to treat
Stipa staintonii
as a member of
Achnatherum
.
Selected specimens studied.
Baglung
: Sing Khola, wet rocky cliff, 18 Sep. 1976,
H. Tabata, K.R. Rajbhandari & K. Tsuchiya 3711
(KATH).
Dolpa
: Ringmo, dry hillslope,
29°10'20"N
,
82°55'50"E
, 3400 m, 2 Aug. 1973,
S. Einarsson, L.
Skaerby
& B. Wetterhall 3128
(BM); Barbung Khola,
28°52'N
,
83°15'E
, 3030 m, 18 Jul. 1963,
J.D.A. Stainton 4417
(BM); Barbung Khola,
28°52'N
,
83°18'E
, 3030 m, 13 Jul. 1963,
J.D.A. Stainton 4417
(E); Suligad, Rhagaon, rocky slope,
29°28'N
,
82°55'E
, 2600 m, 25 Sep. 1982,
K.R. Rajbhandari & K.J. Malla 6740
(KATH); Karnali, Ringmigaon, on dry hilslope, 3400 m, 2 Aug. 1973,
S. Einarsson, L.
Skaerby
, B. Wetterhall 3126
(UPS); Karnali,
Manang
: Bhraka, Humde, open rocky slope,
28°38'24"N
,
84°5'36"E
, 3400 m, 3 Aug. 1983,
K.R. Rajbhandari 8814
(KATH); Marsyandi valley, Tangi above Manangbhot, on stony sandy places,
28°39'56"N
,
84°1'33"E
, 3800 m, 12 Oct. 1969,
T. Wraber 36427
(E); Tangi, above Manangbhot, Tangje, on stony sandy places,
28°39'22"N
,
84°2'2"E
, 3800 m, 12 Oct. 1969,
T. Wraber 479
(BM).
Mustang
: Entre Marpha et Syang; Syang, Marpha,
28°45'11"N
,
83°41'28"E
, 2650 m, 16 Sep. 1981,
M.A. Farille 81-340
(E); Entre Larjung et Tukuche, 2550 m, 14 Sep. 1981,
M.A. Farille 81-313
(E); Cha Lungpa, in E-facing Cupressus forest, 3030 m, 3 Oct. 1977,
G. Miehe 80
(BM); Phalyak, dry place
28°49'24"N
,
83°44'23"E
, 4110 m, 9
Aug
. 2002,
F. Miyamoto, N. Kurosaki, S. Akiyama, H. Ikeda, Y. Iokawa, Y. Takahashi, M. Tsusaka & M.N. Subedi 20210022
(KATH); Phalyak to Pongio Kharka,
28°49'0"N
,
83°45'0"E
, 3800-4000 m, 9 Aug. 2002,
F. Miyamoto, N. Kurosaki, S. Akiyama, H. Ikeda, Y. Iokawa, Y. Takahashi, M. Tsusaka & M.N. Subedi 20220034
(KATH); Chele - Samar,
28°57'43"N
,
83°48'6"E
, 3450-3670 m, 4 Aug. 2001,
S. Noshiro, M. Amano, Y. Iokawa, T. Kurosawa, & M.N. Subedi 20105005
(KATH); Tukuche, open rocky slope,
28°42'33"N
,
83°38'37"E
, 2630 m, 19 Jul. 1983,
K.R. Rajbhandari 7991
(KATH); Jomsom, open slope,
28°46'51"N
,
83°43'27"E
, 2760 m, 21 Jul. 1983,
K.R. Rajbhandari 8072
(KATH); Jomsom to Sayang, cliffs in dry valley with scrub (
Caragana
, Clematis, Rosa etc.),
28°46'51"N
,
83°43'27"E
, 2700 m, 19 Sep. 1999,
Shrestha et al. 1014
(E); Tukuche,
28°42'33"N
,
83°38'37"E
, 3180 m, 21 Aug. 1954,
J.D.A. Stainton, W.R. Sykes & L.H.J. Williams 7352
(BM).
Rukum
: Sen Khola,
28°42'54"N
,
82°57'21"E
, 3790 m, 4 Oct. 1954,
J.D.A. Stainton, W.R. Sykes & L.H.J. Williams 4677
(BM).