Gomphidae of Guangdong & Hong Kong, China (Odonata: Anisoptera) 2177
Author
Wilson, Keith Duncan Peter
Author
Xu, Zaifu
text
Zootaxa
2009
2009-08-06
2177
1
62
journal article
11755334
Lamelligomphus formosanus
(
Matsumura 1926
)
Figures 15
(a–n)
Lindenia formosana
(Matsumura in Oguma 1926): Matsumura in Oguma (1926 97, type-loc. “
Formosa
”).
Onychogomphus micans
Needham 1930
:
Needham (1930: 41–42
, pl. 5, figs 10 & 10a, type-loc. “Lo-cheng-hsien, Kiangsi”
Guangxi
);
Chao (1954: 267–268
, figs 450–457,
Fujian
);
Sui & Sun (1984)
: 98
–100, figs 65a–f;
Fujian
,
Guangdong
&
Taiwan)
.
FIGURE 15(a–n):
Lamelligomphus formosanus
.
[a–f] ♂, Shimentai, Guangdong—(a) head, frontal, (b) synthorax, lateral, (c) caudal appendages, lateral, (d) caudal appendages, dorsal, (e) anterior hamulus, lateral, (f) posterior hamulus, lateral; [g–j] ♀, Shimentai, Guangdong—(g) head, frontal, (h) occiput, frontal, (k) caudal abdomen, ventral; [i–j] ♀, Nanling, Guangdong—(i) occiput, frontal, (j) head, frontal; ♀,
Lamelligomphus formosanus
as
Lamelligomphus jiuquensis
, from
Liu (1993)
—(l) synthorax, (m) occiput and vertex, frontal, (n) caudal abdomen, ventral.
FIGURE 16(a–t):
Lamelligomphus
.
[a–b] ♀,
Lamelligomphus hainanensis
, Hainan, from
Wilson & Reels (2001)
— (a) head, frontal, (b) postgenital plate; [c–f]
Lamelligomphus hainanensis
, Hong Kong, from Wilson 1995—(c) ♀, synthorax, lateral, (d) ♀, occiput, frontal, (e) ♂, anterior hamulus, (f) ♂, posterior hamulus; [g–h] ♀,
Lamelligomphus hanzhongensis
, Shaanxi from Yang & Zhu (2001)—(g) head, frontal, (h) synthorax, (i) postgenital plate; [j–l] ♀,
Lamelligomphus chaoi
, Yunnan, from Zhu (1999)—(j) head, frontal, (k) synthorax, (l) occiput, frontal; [m–q]
Lamelligomphus laetus
, Yunnan, from Yang & Davies (1993)—(m) ♂, synthorax, (n) ♀, occiput, frontal, (o) ♂, anterior hamulus, (p) ♂, posterior hamulus, (q) ♀, postgenital plate; [r–s]
Lamelligomphus laetus
, Yunnan, as
Lamelligomphus parvulus
, from
Zhou & Li (2000)
— (r) ♂, synthorax, lateral, (s) ♂, hamuli, (t) ♀, postgenital plate.
FIGURE 17(a–z):
Lamelligomphus
.
[a–e]
Lamelligomphus tutulus
, Guizhou, from Liu & Chao in
Chao (1990)
— (a) ♂, synthorax, (b) ♂, anterior hamulus, (c) ♂, posterior hamulus, (d) ♀, occiput and vertex, frontal, (e) ♀, postgenital plate; [f–g]
Lamelligomphus camelus
, from
Chao (1990)
—(f) ♂, synthorax, (g) ♂, caudal abdomen, (h) ♂, hamuli, (i) ♀, occipital margin, frontal, (j) ♂, dorsum of 8th abdominal segment, lateral; [k–m] ♀,
Lamelligomphus trinus
, Jiangxi, from
Chao (1990)
— (k) synthorax, (l) postgenital plate, (m) occiput and vertex, frontal; [n–r]
Lamelligomphus ringens
, Fujian, from
Chao (1990)
— (n) ♂, synthorax, (m) ♂, anterior hamulus, (p) ♂, posterior hamulus, (q) ♀, occiput, frontal, (r) ♀, postgenital plate; [s–t]
Lamelligomphus choui tienfuensis
, Sichuan, ♂, from
Chao (1995)
—(s) anterior hamulus, (t) posterior hamulus; [u–z]
Lamlligomphus motuoensis
, Xizang, from
Chao (1990)
— (u) ♂, synthorax, (v) ♂, occiput and vertex, frontal, (w) ♀, caudal abdomen and postgenital plate, (x) ♀, occiput and vertex, (y) ♂, anterior hamulus, (z) ♂, posterior hamulus.
Lamelligomphus formosanus
(Oguma 1926)
:
Laidlaw (1930: 194
,
Formosa
);
Chao (1990: 356–360
, figs 1–13,
Fujian
,
Guangxi
and
Taiwan)
;
Tsuda (2000: 100
,
China
,
Taiwan)
;
Hua (2000: 11
,
Fujian
,
Guangdong
,
Guangxi
,
Guizhou
,
Hainan
,
Jiangxi
and
Taiwan)
;
Bridges (1994
: VII.91, VIII.34,
Formosa
);
Zhang (1999: 238–239
, figs 24.65a–e, Fujian, Guangxi & Taiwan);
Wang (2000: 178–179
, 206–207, Taiwan).
Onychogomphus formosanus
(Matsumura)
:
Needham (1930: 43
,
Formosa
);
Chao (1954: 274–275
, Taiwan);
Matsuki (1978: 138
, 152, fig. 14, larval description);
Asahina (1961b: 60
, designation of
lectotype
from Hoppo = Peipu, Taiwan);
Asahina (1978a: 5–6
, figs 12–15, “Foochow”, Fujian, synonym of
micans
);
Lieftinck
et al
. (1984: 35
, Taiwan);
Davies and Tobin (1985: 43
,
Formosa
).
Lamelligomphus jiuquensis
Liu 1993
:
Liu (1993: 21–23
, figs 1–4, type-loc. Chong’an Xingcun,
Fujian
);
Zhang (1999: 239
, figs 24.66a–c,
Fujian
).
syn. nov.
Specimens:
3 ♂
,
4 ♀
, Shimentai,
6 viii 2003
, leg. KW;
2 ♂
,
1 ♀
, Shimentai,
7 viii 2003
, leg. KW;
1 ♂
, Nanling,
5 viii 2004
, leg. KW;
1 ♀
, Nanling,
9 viii 2005
, leg. KW.
Remarks:
Liu (1993)
described
Lamelligomphus jiuquensis
from a single female collected from Xingcun,
Fujian
(
Figs. 15l–n
).
Liu (1993)
considered
jiuquensis
to be similar to
L. formosanus
but differed in four points: (i) prothorax black, with yellow stripes, (ii) colour pattern of synthorax with 2
nd
and 3rd lateral yellow stripes broad and complete, but antehumeral stripe broken in the middle, (iii) 7th abdominal sternum not constricted subapically, (iv) a pair of occipital horns closing or parallel to each other. Lui's characters used to distinguish
L. jiuquensis
from
L. formosanus
also apply to female
L. formosanus
! Four
male and
one female
L. formosanus
specimens collected from Shimentai have an interrupted yellow stripe across the metepisternum and
one male
and
four females
are marked with an uniterrupted yellow stripe and although
one male
and
one female
possessed uninterrupted antehumerals stripes the remainder were interrupted (
Fig. 15b
). Three of the Shimentai females are marked with a pair of small yellow spots on the central dorsum of the prothorax, whereas the prothorax of the remaining
two females
and all
five males
are entirely black. There is slight variation in the structure of the occipital horns;
one female
features almost parallel sided horns (
Fig. 15i
). This same female from Nanling also featured an entirely black postclypeus and a slightly shallower groove dividing the yellow spots on the postfrons than Shimentai females. It is apparent that the differences described by
Liu (1993)
for the
type
female
L. jiuquensis
are merely phenotypic variation in colouration. There are no structural differences between
L. jiuquensis
(
Figs. 15j–l
) and
L. formosanus
(
Figs. 15a–i
). The undulating occiput, shape of the occipital horns and deep central dorsal groove on the top of the frons are identical. The two species are here considered synonymous.
The record of
Lamelligomphus formosanus
from
Hainan
(
Hua 2000
) requires confirmation.
Distribution:
China
(
Fujian
,
Guangdong
,
Guangxi
,
Guizhou
,
Hainan
[?] and
Taiwan)
.