Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups
Author
Gunter, Nicole L.
Author
Weir, Thomas A.
text
Zootaxa
2019
2019-11-08
4695
3
201
252
journal article
24935
10.11646/zootaxa.4695.3.1
256591b2-6d7b-465d-a818-9a158017ca09
1175-5326
3532515
32E17629-F29A-47C8-ACCB-69A32C17B147
Lepanus villosus
Matthews, 1974
(
Figs 16A, 16C
,
17D
,
18D
,
19D
)
Lepanus villosus
Matthews, 1974: 118
.
Types examined:
Holotype
:
♂
“ Palmerston N.P., Q.
1.iv.68
,
E. Matthews
” / “
Lepanus villosus Det. 1972
M, E.G. Matthews,
HOLOTYPE
” / “
ANIC
Database No. 25 057986” in
ANIC
[
17°34’0”S
,
145°45’0”E
]
.
Paratypes
:
“Palmerston N.P., Q.
1.iv.68
,
E. Matthews
” / “
Lepanus villosus
Det. 1972
M, E.G. Matthews, PARATYPE” / “
ANIC
Database No. 25 057987”
2 specimens ♀
in
ANIC
[
17°34’0”S
,
145°45’0”E
],
;
“Tully, N.Q.,
12.x.1969
,
R.J. Huppatz
” / “
Lepanus villosus Det. 1972
M, E.G. Matthews, PARATYPE” / “
ANIC
Database No. 25 057988”
1 specimen ♀
in
ANIC
[
17°36’0”S
,
145°55’0”E
]
;
“QLD. <
50m
Seymour Ra.
,
10/vii1971
, Taylor Feehan” / Berlesate anic 376, rainforest 17.26x146.0” / “
Lepanus villosus
Det. 1972
,
E.G. Matthews
, PARATYPE” / “
ANIC
Database No. 25 057989”
1 specimen ♂
in
ANIC
[
17°26’0”S
,
146°0’0”E
]
;
“
Gillies
H’way, N.Q. 2/10.
GB
.” / “
PARATYPE
” / “EM.117.” / “
Lepanus villosus Det. 1972
,
E.G. Matthews
, PARATYPE” / “
ANIC
Database No. 25 057990”
1 specimen ♂
in
ANIC
[
17°12’0”S
,
145°42’0”E
]
;
“ QLD.
750m
Koombooloomba
,
4/vii/1971
Taylor Feehan
” / “Ber- lesate
ANIC 259
rainforest” / “
Lepanus villosus Det. 1972
,
E.G. Matthews
, PARATYPE” / “
ANIC
Database No. 25 057991”
1 specimen ♂
in
ANIC
[
17°49’0”S
,
145°33’0”E
]
;
“
Davies Creek
,
Atherton
Tbld,
N.Qld
,
10 vii 1970
,
J.H. Connell
” / “
Lepanus villosus Det. 1972
M, E.G. Matthews, PARATYPE” (25-058496
1 specimen ♀
, 25-058497
1 specimen ♂
, 25-058498
1 specimen ♂
,
1 specimen ♀
(on same card), 25-058499
1 specimen ♀
, 25-058500 1 speci- men
♂
in
QM
) [
16°59’0”S
,
145°33’0”E
]
.
FIGURE 16
. Some diagnostic characters of the
Lepanus villosus
species group.
A
, lateral view of elytron of
Lepanus villosus
, white arrow points to deep pit at the base of stria 8;
B
, lateral view of elytron of
Lepanus vestitus
, note the absence of any deep pit;
C
, elytron of
Lepanus villosus
dotted line showing humeral callus between striae 6 and 7 and white arrow showing depression at base of striae 4 and 5;
D
, male metatibiae of
Lepanus vestitus
, notch on inner edge;
E
, male protibia of
Lepanus pseudovillosus
,
showing typical configuration for the species group including the small comb of setae (grey arrow);
F
, close up of elytral interval surface of
Lepanus reticulatus
, showing reticulation of short longitudinal striations;
G
, close up of elytral interval surface of
Lepanus vestitus
showing reticulations of fine meshes.
FIGURE 17
. Habitus of species within the
Lepanus villosus
species group.
A
,
Lepanus pseudovillosus
;
B
,
Lepanus reticulatus
;
C
,
Lepanus vestitus
;
D
,
Lepanus villosus
. All specimens in comparative scale. Scale bar 1 mm.
FIGURE 18
. Left, dorsal, and right view of aedeagi of species within the
Lepanus villosus
species group.
A
,
Lepanus pseudovillosus
;
B
,
Lepanus reticulatus
;
C
,
Lepanus vestitus
;
D
,
Lepanus villosus
. All specimens in comparative scale. Scale bar 0.2 mm.
FIGURE 19
. Pygidial configuration of species within the
Lepanus villosus
species group.
A
,
Lepanus pseudovillosus
;
B
,
Lepanus reticulatus
;
C
,
Lepanus vestitus
;
D
,
Lepanus villosus
. All specimens in comparative scale. Scale bar 1 mm.
Other material examined.
532 specimens
from 81 localities were examined.
Cape Tribulation
transect
Site
5A,
16°4’41”S
,
145°27’0”E
,
1–31.i.1983
, (T44339
1 specimen
in
QM
)
;
Cape Tribulation
transect
Site
5A,
16°4’41”S
,
145°27’0”E
,
5–9.i.1983
, (T44480–T44482
3 specimens
in
QM
)
;
Cape Tribulation
transect
Site
5A,
16°4’41”S
,
145°27’0”E
,
20–23.iv.1983
, (T44483–T44484
2 specimens
in
QM
)
;
1.5 km
Eby
, N
Mount Sorrow
, [
16°5’0”S
,
145°27’0”E
],
25.iii.1984
,
A. Calder
&
T.A. Weir
(25-059588
2 specimens
in
ANIC
)
;
Thompson Creek
, (
Grove
),
16°6’31”S
,
145°26’25”E
,
12.ii–15.iii.1998
, (T85668
1 specimen
in
QM
)
;
Thompson Creek
, (
Grove
),
16°6’31”S
,
145°26’25”E
,
15.iii–7.v.1998
, (T88828
1 specimen
in
QM
)
;
Cooper Creek
, (
Grove
),
16°9’10”S
,
145°24’19”E
,
11.i–10.ii.1998
, (T73113–T73119
7 specimens
in
QM
)
;
Cooper Creek
, (
Grove
),
16°9’10”S
,
145°24’19”E
,
10.ii– 16.iii.1998
, (T85663–T85667, T85669, T85670, T85672–T85673, T88823–T88825
12 specimens
in
QM
)
;
Cooper Creek
, (
Grove
),
16°9’10”S
,
145°24’19”E
,
16.iii –7.v.1998
, (T88819–T88821
3 specimens
in
QM
)
;
Cooper Creek
, (
Grove
),
16°9’10”S
,
145°24’19”E
,
11.ix–24.x.1998
, (T88822, T88826
2 specimens
in
QM
)
;
Pimm’s Block
, (
Grove
),
16°11’20”S
,
145°24’18”E
,
13.iii–8.v.1998
, (T88829, T88831–T88833
4 specimens
in
QM
)
;
Pimm’s Block
, (
Grove
),
16°11’20”S
,
145°24’18”E
,
10.ix–24.x.1998
, (T88830, T88834
2 specimens
in
QM
)
;
Hutchinson Creek
, (
Grove
),
16°13’4”S
,
145°23’57”E
,
13.i–11.ii.1998
, (T73120–T73121
2 specimens
in
QM
)
;
Hutchinson Creek
, (
Grove
),
16°13’4”S
,
145°23’57”E
,
11.ii–14.iii.1998
, (T85671
1 specimen
in
QM
)
;
Hutchinson Creek
, (
Grove
),
16°13’4”S
,
145°23’57”E
,
14.iii–8.v.1998
, (T88827
1 specimen
in
QM
)
;
Alexandra Bay
, [
16°12’0”S
,
145°28’0”E
],
24.vi.1971
,
R.W. Taylor
&
J. Feehan
, (25-057996
1 specimen
in
ANIC
)
;
Lync-Haven
,
Daintree area
,
16°12’7”S
,
145°24’32”E
,
35 m
,
2–10.xii.2012
,
T.A. Weir
&
G.B. Monteith
, (25-057655
1 specimen
in
ANIC
)
;
Lync-Haven
,
Daintree area
,
16°12’7”S
,
145°24’32”E
,
35 m
,
6–9.xii.2012
,
G.B. Monteith
(25-057653
1 specimen
in
ANIC
)
;
Thornton Range
near
Daintree
, [
16°14’0”S
,
145°24’0”E
],
200 m
,
12–18.vii.1982
,
S. & J. Peck
(25-059578
6 specimens
in
ANIC
)
;
Buchanan Creek
, (
Grove
),
16°14’39”S
,
145°25’54”E
,
12.iii–8.v.1998
, (T88818
1 specimen
in
QM
)
;
Humbug Landing
,
Daintree
,
16°15’13”S
,
145°20’54”E
,
14–18.v.1998
, (T66779–T66780
2 specimens
in
QM
)
;
Mount Gorton
,
16°57’0”S
,
145°53’0”E
,
18.xi.1993
, (T44315–T44321
7 specimens
in
QM
)
;
Mount Murray Prior
,
16°56’0”S
,
145°51’0”E
,
7.xii.1995
(T44324
1 specimen
in
QM
)
;
Mount Murray Prior
,
16°56’0”S
,
145°51’0”E
,
31.x.1995
–
8.xii.1995
, (T44419–T44423
5 specimens
in
QM
)
;
Mount Murray Prior
,
16°56’0”S
,
145°51’0”E
,
7–8.xii.1998
, (T70933–T70954
21 specimens
in
QM
)
;
Chujeba Peak
summit,
16°56’14”S
,
145°39’19”E
,
16.xii.1989
, (T44340
1 specimen
in
QM
)
;
Davies Creek Road
via
Mareeba
, [
17°1’0”S
,
145°35’0”E
],
9.x–17.x.1976
,
R.I. Storey
(25-059595
3 specimens
in
ANIC
,
2 specimens
in
QDPC
)
;
Davies Creek Road
, via
Mareeba
,
17°2’33”S
,
145°36’51”E
,
26.xi.1985
, (T44341
x
1
in
QM
)
;
Davies Creek Road
, via
Mareeba
,
17°2’33”S
,
145°36’51”E
,
4–13.xii.1988
, (T44414–T44415
2 specimens
in
QM
)
;
Isley Creek
, upper,
17°2’41”S
,
145°39’57”E
,
29–30.xi.1993
, (T44330– T44338
9 specimens
in
QM
)
;
Mount Tiptree
, [
17°3’0”S
,
145°37’0”E
],
13.vii.1984
,
R.B. Halliday
(25-059585
1 specimen
in
ANIC
)
;
Mount Tiptree
,
17°3’0”S
,
145°38’0”E
],
730 m
,
29.vi.1971
,
R.W. Taylor
&
J. Feehan
SAMA25- 027824
2 specimens
in
SAMA
);
North Bell Peak
summit,
17°5’19”S
,
145°52’44”E
,
16.ix.1981
, (T44328
1 specimen
in
QM
)
;
North Bell Peak
summit,
17°5’19”S
,
145°52’44”E
,
22.xi.1990
, (T44342–T44345
3 specimens
in
QM
)
;
North Bell Peak
summit,
17°5’19”S
,
145°52’44”E
,
20–22.xi.1990
, (T44365–T44370
6 specimens
in
QM
)
;
Lamb Range
,
6 km
NNE Tinaroo Falls
,
17°6’40”S
,
145°33’58”E
,
1159 m
,
16.xi.2009
,
G.B. Monteith
&
Turco
, (25-
060851
3 specimens
in
ANIC
aslso as COL960- COL962)
;
Danbulla State Forest Robson Creek
Road
1.5 km
N of road to
Tinaroo
, [
17°7’0”S
,
145°37’0”E
],
27.iv–2.v.1998
,
Miledge,G.
, (25-
060767
1 specimen
in
AM
)
;
Danbulla
S.F.,
13 km
NE of Yungaburra
[Danbulla Forestry Reserve], [
17°10’0”S
,
145°38’0”E
],
13.i–13.ii.1987
,
R.I. Storey
&
De Faveri
, (25-059594
1 specimen
in
QDPC
)
;
Danbulla State Forest
,
13 km
NE of Yungaburra
[Danbulla Forestry Reserve], [
17°10’0”S
,
145°38’0”E
],
23.iv–27.v.1987
,
R.I. Storey
&
De Faveri
, (25-059604
1 specimen
in
ANIC
)
;
Danbulla State Forest
,
13 km
NE of Yungaburra
[Danbulla Forestry Reserve], [
17°10’0”S
,
145°38’0”E
],
23.vi–28.vii.1987
,
R.I. Storey
&
De Faveri
, (25-
032825
1 specimen
in
QDPC
)
;
Danbulla State Forest
,
13 km
NE of Yungaburra
[Danbulla Forestry Reserve], [
17°10’0”S
,
145°38’0”E
],
28.vii–3.ix.1987
,
R.I. Storey
&
De Faveri
, (25-
032826
1 specimen
in
QDPC
)
;
Cathedral Fig
,
17°10’52”S
,
145°39’26”E
,
10.xii.1988
, (T44326
1 specimen
in
QM
)
;
Graham Range
, summit,
17°16’24”S
,
145°57’58”E
,
1.xi–8.xii.1995
, (T44404–T44405
2 specimens
in
QM
)
;
Graham Range
,
350 m
,
17°16’37”S
,
145°57’48”E
,
1.xi–8.xii.1995
, (T54656–T54657
2 specimens
in
QM
)
;
Boar
Pock- et
Road
, approximately
8 km
N of Gillies Highway
[Gillies Road], [
17°12’0”S
,
145°42’0”E
],
13.xii.1969
,
J.G. Brooks
, (25-057998
1 specimen
in
ANIC
)
;
Boar Pocket Road
, approximately
8 km
N of Gillies Highway
[Gillies Road], [
17°12’0”S
,
145°42’0”E
],
9.xii.1969
,
J.G. Brooks
, (25-057997
1 specimen
in
ANIC
)
;
Boar Pocket Road
3–9 mi
,
17°11’57”S
,
145°40’8”E
,
2500 feet
,
8.iv.1974
,
J.G. Brooks
, (CMNEN-00030586-00030588 in
CMNC
)
;
Bellenden Ker Range
,
0.5 km
S Cable Tower
7, [
17°16’0”S
,
145°55’0”E
],
500 m
,
25–31.x.1981
,
Earthwatch
,
Queensland
Museum
, (25-
032830
1 specimen
in
QDPC
, 25-059582
x
5
in
ANIC
)
;
Bellenden Ker Range
,
1 km
S Cable Tower
6,
17°16’0”S
,
145°53’0”E
],
500 m
,
17–24.x.1981
,
Earthwatch
,
Queensland
Museum
, (25-057999
2 specimens
in
ANIC
)
;
Bellenden Ker Range
,
1 km
S Cable Tower
6,
17°16’0”S
,
145°53’0”E
],
500 m
,
25–31.x.1981
,
Earthwatch
,
Queensland
Museum
, (25-059583
1 specimen
in
ANIC
)
;
Lake Barrine National Park
, [
17°16’0”S
,
145°39’0”E
],
7.vii.1984
,
R.B. Halliday
(25-059587
1 specimen
in
ANIC
)
;
Wright Creek
,
Lake Eacham
, [
17°16’0”S
,
145°37’0”E
],
750 m
,
31.vii.1982
,
S. & J. Peck
(25-058000
2 specimens
in
ANIC
)
;
Wright Creek
,
Lake Eacham
, [
17°16’0”S
,
145°37’0”E
],
720 m
,
26–30.vii.1982
,
S. & J. Peck
(25-059579
2 specimens
in
ANIC
)
;
Baldy Mountain Road Atherton
, [
17°17’0”S
,
145°24’0”E
],
12.xii.1976
,
R.I. Storey
(25-
032828
1 specimen
in
QDPC
)
;
Baldy Mountain
near
Atherton
[Mount Baldy], [
17°17’0”S
,
145°27’0”E
],
xii.1977
–
i.1978
,
R.I. Storey
(25-
032827
2 specimens
in
QDPC
)
;
Eacham National Park
[Lake Eacham National Park], [
17°17’0”S
,
145°37’0”E
],
760 m
,
21.iii.1975
,
R.W.Taylor
(25-057994
1 specimen
in
ANIC
)
;
Bartle Frere
,
Gourka Road
,
17°22’38”S
,
145°44’34”E
,
28–30.xi.1998
, (T70929–T70932
4 specimens
in
QM
)
;
Bartle Frere
,
West Base
,
17°22’47”S
,
145°45’51”E
,
10.i–31.iii.1995
, (T44447–T44452
6 specimens
in
QM
)
;
Bartle Frere
,
West Base
,
17°22’47”S
,
145°45’51”E
,
7.iii–15.v.1995
, (T44453–T44464, T55904
11 specimens
in
QM
)
;
Bartle Frere
,
West Base
,
17°22’47”S
,
145°45’51”E
,
25.xi.1994
–
10.i.1995
, (T44465–T44479
15 specimens
in
QM
)
;
Lamins Hill
,
17°22’56”S
,
145°42’7”E
,
1.xii.1993
–
25.ii.1994
, (T44413
1 specimen
in
QM
)
;
Bartle Frere
W
Base
, [
17°23’0”S
,
145°46’0”E
],
750 m
,
28.xi.1998
,
G.B. Monteith
(25-
060849
1 specimen
in
ANIC
)
;
Mount Bartle Frere
W side [Bartle Frere South Peak],
17°24’0”S
,
145°49’0”E
],
700 m
,
30.vii.1982
,
S. & J. Peck
(25-059580
2 specimens
in
ANIC
)
;
Mount Bartle Frere
W side [Bartle Frere South Peak], [
17°24’0”S
,
145°49’0”E
],
700 m
,
30.vii–4.viii.1982
,
S. & J. Peck
(25-059581
1 specimen
in
ANIC
)
;
Hugh Nelson Range
,
21 km
S of Atherton
, [
17°25’0”S
,
145°28’0”E
],
12.x–5.xi.1983
,
R.I. Storey
&
Brown
(25-059601
1 specimen
in
ANIC
)
;
Hugh Nelson Range
,
21 km
S of Atherton
, [
17°25’0”S
,
145°28’0”E
],
13.iii–1.v.1984
,
R.I. Storey
&
Brown
(25-059603
1 specimen
in
QDPC
)
;
Hugh Nelson Range
,
21 km
S of Atherton
, [
17°25’0”S
,
145°28’0”E
],
21.vi–26.vii.1984
,
R.I. Storey
&
Brown
(25-059602
x
1
in
ANIC
)
;
Hugh Nelson Range
,
21 km
S of Atherton
, [
17°25’0”S
,
145°28’0”E
],
5.xi–1.xii.1983
,
R.I. Storey
&
Brown
(25-
032829
1 specimen
in
QDPC
)
;
Topaz
,
Hughes Road
, (
Breeden
house),
17°26’4”S
,
145°42’2”E
,
1.vii–30.ix.1993
, (T44426
1 specimen
in
QM
)
;
21 km
S of
Atherton
,
17°26’41”S
,
145°28’35”E
,
5–14.xii.1988
, (T44416
1 specimen
in
QM
)
;
Hugh Nelson Range
, [
17°27’0”S
,
145°29’0”E
],
1150 m
,
3.vii–1.viii.1995
,
P. Zborowski
(25-058038
1 specimen
in
ANIC
)
;
Josephine Falls
,
Mount Bartle Frere
[Bartle Frere], [
17°27’0”S
,
145°52’0”E
],
10.vii.1984
,
R.B. Halliday
(25-059586
2 specimens
in
ANIC
)
;
Longlands Gap
,
17°27’14”S
,
145°28’26”E
,
1120 m
,
18–19.ix.2010
,
G.B. Monteith
(25-059606
1 specimen
, 25-
060866
3 specimens
in
ANIC
also as COL1123-COL1125)
;
tower near
The Crater
,
17°27’22”S
,
145°29’13”E
,
1235m
,
21–23.i.2012
,
G.B. Monteith
(25-055609
7 specimens
in
ANIC
, 25-053847
2 specimens
in
ANIC
)
;
Tower
S of Crater National Park
,
17°27’23”S
,
145°29’12”E
,
25.xi.1994
–
10.i.1995
, (T44346–T44351
6 specimens
in
QM
)
;
Tower
S of Crater National Park
,
17°27’23”S
,
145°29’12”E
,
10.i–31.iii.1995
, (T44352–T44359
8 specimens
in
QM
)
;
Tower
S of Crater National Park
,
17°27’23”S
,
145°29’12”E
,
7.iii–15.v.1995
, (T44360–T44364
5 specimens
in
QM
)
;
Longlands Gap
,
17°27’46”S
,
145°29’32”E
,
30.x.1991
–
23.vii.1992
, (T44389
1 specimen
in
QM
)
;
Millaa Millaa Falls
(below), [
17°29’0”S
,
145°36’0”E
],
760 m
,
18–20.iv.1999
,
C. & I. Reid
(25-059605
1 specimen
in
ANIC
)
;
Millaa Millaa Falls
,
17°29’53”S
,
145°36’32”E
,
6–9.ii.1999
, (T76229,T76231–T76232
3 specimens
in
QM
)
;
Kjellberg Road
turnoff,
17°31’57”S
,
145°36’27”E
,
7.iii–15.v.1995
, (T44380
1 specimen
in
QM
)
;
Mount Fisher
,
Kjellberg Road
,
17°32’34”S
,
145°33’31”E
,
27.iv.1982
, (T44322–T44323
2 specimens
in
QM
)
;
Mount Fisher
,
Kjellberg Road
,
17°32’34”S
,
145°33’31”E
,
3.v.1983
, (T44329
1 specimen
in
QM
)
;
Mount Fisher
,
Kjellberg Road
,
17°32’34”S
,
145°33’31”E
,
27–29.iv.1982
, (T44417–T44418, T44425
3 specimens
in
QM
)
;
Kjellberg Road
,
State Forest
boundary,
17°32’9”S
,
145°34’39”E
,
6–9.ii.1999
, (T76233
1 specimen
in
QM
)
;
Maalan State Forest
,
Ravenshoe Road
,
17°35’23”S
,
145°35’40”E
,
10.i–31.iii.1995
, (T44427–T44431
5 specimens
in
QM
)
;
Maalan State Forest
,
Ravenshoe Road
,
17°35’23”S
,
145°35’40”E
,
7.iii–15.v.1995
, (T44435–T44438
4 specimens
in
QM
)
;
Maalan State Forest
,
Ravenshoe Road
,
17°35’23”S
,
145°35’40”E
,
25.xi.1994
–
10.i.1995
, (T44439–T44446
8 specimens
in
QM
)
;
Maalan State Forest
,
Ravenshoe Road
,
17°35’23”S
,
145°35’40”E
,
6–9.ii.1999
, (T76219– T76227
9 specimens
in
QM
)
;
Henrietta Creek Palmerston National Park
, [
17°35’54”S
,
145°45’30”E
],
22–24.iv.1973
,
R.I. Storey
(CMNEN-00030582-00030585 in
CMNC
)
;
Millaa Millaa Falls
, [
17°30’0”S
,
145°37’0”E
],
760 m
,
11.v.1974
,
R.I. Storey
(25-
032831
1 specimen
in
QDPC
)
;
Millaa Millaa Falls
, [
17°30’0”S
,
145°37’0”E
],
760 m
,
13. vii–15.viii.1990
,
R.I. Storey
&
H.K. Halfpapp
, (25-
032818
3 specimens
in
QDPC
)
;
Millaa Millaa Falls
, [
17°30’0”S
,
145°37’0”E
],
760 m
,
13.iii–10.iv.1990
,
R.I. Storey
&
H.K. Halfpapp
, (25-
032816
3 specimens
in
QDPC
)
;
Millaa Millaa Falls
, [
17°30’0”S
,
145°37’0”E
],
760 m
,
15.viii–12.ix.1990
,
R.I. Storey
&
H.K. Halfpapp
, (25-
032819
2 specimens
in
QDPC
)
;
Millaa Millaa Falls
, [
17°30’0”S
,
145°37’0”E
],
760 m
,
4.i–7.ii.1990
,
R.I. Storey
&
H.K. Halfpapp
, (25-
032815
1 specimen
in
QDPC
)
;
Millaa Millaa Falls
, [
17°30’0”S
,
145°37’0”E
],
760 m
,
6.vi–13.vii.1990
,
R.I. Storey
&
H.K. Halfpapp
, (25-
032817
2 specimens
in
QDPC
)
;
BS2
Mount Fisher
, [
17°33’0”S
,
145°32’0”E
],
1150 m
,
31.i–27.ii.1996
,
L. Umback
(25-059610
1 specimen
in
ANIC
)
;
Mount Fisher
, [
17°33’0”S
,
145°32’0”E
],
1150 m
,
1.xii.1994
–
3.i.1995
,
P. Zborowski
(25-059613
1 specimen
in
ANIC
)
;
Mount Fisher
, [
17°33’0”S
,
145°32’0”E
],
1150 m
,
2.viii–4.ix.1995
,
L. Umback
(25-058007
3 specimens
in
ANIC
)
;
Mount Fisher
, [
17°33’0”S
,
145°32’0”E
],
1150 m
,
2.vi–4.vii.1995
,
P. Zborowski
(25-058036
1 specimen
in
ANIC
)
;
Mount Fisher
, [
17°33’0”S
,
145°32’0”E
],
1150 m
,
2–30.xi.1995
,
L. Umback
(25-058013
1 specimen
in
ANIC
)
;
Mount Fisher
, [
17°33’0”S
,
145°32’0”E
],
1150 m
,
21.iii–5.iv.1995
,
P. Zborowski
(25-058021
1 specimen
, 25-058022
2 specimens
, 25-058023
2 specimens
, 25-058024
3 specimens
in
ANIC
)
;
Mount Fisher
, [
17°33’0”S
,
145°32’0”E
],
1150 m
,
3.i–4.ii.1995
,
P. Zborowski
(25-059611
2 specimens
in
ANIC
)
;
Mount Fisher
, [
17°33’0”S
,
145°32’0”E
],
1150 m
,
3.v–2.vi.1995
,
P. Zborowski
(25-058034
3 specimens
, 25-058035
4 specimens
in
ANIC
)
;
Mount Fisher
, [
17°33’0”S
,
145°32’0”E
],
1150 m
,
3.x–2.xi.1995
,
L. Umback
(25-058011
2 specimens
in
ANIC
)
;
Mount Fisher
, [
17°33’0”S
,
145°32’0”E
],
1150 m
,
4.ii–21.iii.1995
,
P. Zborowski
(25-059612
2 specimens
in
ANIC
)
;
Mount Fisher
, [
17°33’0”S
,
145°32’0”E
],
1150 m
,
4.vii–2.viii.1995
,
P. Zborowski
(25-058001
2 specimens
, 25-058002
1 specimen
in
ANIC
)
;
Mount Fisher
, [
17°33’0”S
,
145°32’0”E
],
1150 m
,
4.ix–3.x.1995
,
L. Umback
(25-058008
1 specimen
, 25-058009
9 specimens
in
ANIC
)
;
Mount Fisher
, [
17°33’0”S
,
145°32’0”E
],
1150 m
,
5.iv–3.v.1995
,
P. Zborowski
(25-058025
3 specimens
, 25-058026
5 specimens
, 25-058027
3 specimens
, 25-058028
1 specimen
, 25-058029
3 specimens
, 25-058030
2 specimens
in
ANIC
)
;
Henrietta Creek Palmerston National Park
[Plamerston National Park], [
17°34’0”S
,
145°45’0”E
],
22–24.iv.1973
,
R.I. Storey
(25-
032824
6 specimens
in
QDPC
, 25-058501
8 specimens
in
QM
)
;
BS3
Massey Creek
, [
17°37’0”S
,
145°34’0”E
],
1000 m
,
1.xii.1994
–
3.i.1995
,
P. Zborowski
(25-059608
5 specimens
in
ANIC
)
;
BS3
Massey Creek
, [
17°37’0”S
,
145°34’0”E
],
1000 m
,
3.i–4.ii.1995
,
P. Zborowski
(25-059609
1 specimen
in
ANIC
)
;
BS3
Massey Creek
, [
17°37’0”S
,
145°34’0”E
],
1000 m
,
4.ii–6.iii.1995
,
P. Zborowski
(25-059614 3 spec- imens in
ANIC
)
;
Massey Creek
, [
17°37’0”S
,
145°34’0”E
],
1000 m
,
2.viii–4.ix.1995
,
L. Umback
(25-058006
6 specimens
in
ANIC
)
;
Massey Creek
, [
17°37’0”S
,
145°34’0”E
],
1000 m
,
2.vi–4.vii.1995
,
P. Zborowski
(25-058037
1 specimen
, 25-058039
5 specimens
, 25-058040
5 specimens
in
ANIC
)
;
Massey Creek
, [
17°37’0”S
,
145°34’0”E
],
1000 m
,
2.xi–30.xi.1995
,
L. Umback
(25-058014
1 specimen
, 25-058015
3 specimens
in
ANIC
)
;
Massey Creek
, [
17°37’0”S
,
145°34’0”E
],
1000 m
,
3.v–2.vi.1995
,
P. Zborowski
(25-058031
1 specimen
, 25-058032
2 specimens
, 25-058033
2 specimens
in
ANIC
)
;
Massey Creek
, [
17°37’0”S
,
145°34’0”E
],
1000 m
,
3.x–2.xi.1995
,
L. Umback
(25-058012
14 specimens
in
ANIC
)
;
Massey Creek
, [
17°37’0”S
,
145°34’0”E
],
1000 m
,
4.vii–2.viii.1995
,
P. Zborowski
(25-058003
6 specimens
, 25-058004
3 specimens
, 25-058005
2 specimens
in
ANIC
)
;
Massey Creek
, [
17°37’0”S
,
145°34’0”E
],
1000 m
,
4.ix–3.x.1995
,
L. Umback
(25-058010
9 specimens
in
ANIC
)
;
Massey Creek
, [
17°37’0”S
,
145°34’0”E
],
1000 m
,
6.iii–5.iv.1995
,
P. Zborowski
(25-058016
3 specimens
, 25-058017
1 specimen
, 25-058018
2 specimens
, 25-058019
12 specimens
, 25-058020
17 specimen
in
ANIC
)
;
South Johnstone Forestry Camp
,
5 km
W,
17°38’12”S
,
145°40’15”E
,
1.xii.1993
–
25.ii.1994
, (T44403
1 specimen
in
QM
)
;
Vine Creek
,
Majors Mountain
,
17°40’58”S
,
145°32’2”E
,
4–6.ii.1999
, (T76234–T76236
3 specimens
in
QM
)
;
Vine Creek
,
Majors Mountain
,
17°40’58”S
,
145°32’2”E
,
15–18.iv.1999
, (T79131
1 specimen
in
QM
)
;
Charmillin Creek
crossing,
17°42’3”S
,
145°31’21”E
,
8.xii.1989
–
5.i.1990
, (T44373–T44374
23 specimens
in
QM
)
;
Charmillin Creek
crossing,
17°42’3”S
,
145°31’21”E
,
5–11.ii.1998
, (T62197
1 specimen
in
QM
)
;
Ravenshoe State Forest
,
Tully Falls
Road, [
17°43’0”S
,
145°33’0”E
],
10.x–15.xi.1987
,
A. Walford-Huggins
(25-059584
2 specimens
in
ANIC
)
;
Ravenshoe State Forest
,
Tully Falls
Road, [
17°43’0”S
,
145°33’0”E
],
15.xi–12.xii.1987
,
A. Walford-Huggins
(25-059589
3 specimens
in
ANIC
)
;
Tully Falls State Forest
18 km
SSW Ravenshoe
, [
17°46’0”S
,
145°31’0”E
],
730 m
,
1.x– 5.xi.1987
,
R.I. Storey
&
Dickinson
, (25-
032820
1 specimen
in
QDPC
, 25-059590
6 specimens
in
ANIC
)
;
Tully Falls State Forest
18 km
SSW Ravenshoe
, [
17°46’0”S
,
145°31’0”E
],
730 m
,
20.iv–25.v.1988
,
R.I. Storey
&
Dickinson
, (25-059591
1 specimen
in
ANIC
)
;
Tully Falls State Forest
18 km
SSW Ravenshoe
, [
17°46’0”S
,
145°31’0”E
],
730 m
,
5.i–7.ii.1988
,
R.I. Storey
&
Dickinson
, (25-
032822
1 specimen
in
QDPC
, 25-059598
1 specimen
in
ANIC
)
;
Tully Falls State Forest
18 km
SSW Ravenshoe
, [
17°46’0”S
,
145°31’0”E
],
730 m
,
7.xii.1987
–
7.i.1988
,
R.I. Storey
&
Dickinson
, (25-059597
2 specimens
in
ANIC
)
;
Tully Falls State Forest
18 km
SSW Ravenshoe
, [
17°46’0”S
,
145°31’0”E
],
730 m
,
7.xi–7.xii.1987
,
R.I. Storey
&
Dickinson
, (25-
032821
2 specimens
in
QDPC
, 25-059592
4 specimens
in
ANIC
)
;
Tully Falls State Forest
18 km
SSW Ravenshoe
, [
17°46’0”S
,
145°31’0”E
],
730 m
,
9.ii– 3.iii.1988
,
R.I. Storey
&
Dickinson
, (25-059600
2 specimens
in
ANIC
)
;
Tully Falls State Forest
11 km
SSW Ravenshoe
, [
17°47’0”S
,
145°34’0”E
],
900 m
,
1.x–5.xi.1987
,
R.I. Storey
&
Dickinson
, (25-
032823
1 specimen
in
QDPC
, 25-059593
1 specimen
in
ANIC
)
;
Tully Falls State Forest
11 km
SSW Ravenshoe
, [
17°47’0”S
,
145°34’0”E
],
900 m
,
20.iv–25.v.1988
,
R.I. Storey
&
Dickinson
, (25-059599
1 specimen
in
ANIC
)
;
Tully Falls State Forest
11 km
SSW Ravenshoe
, [
17°47’0”S
,
145°34’0”E
],
900 m
,
7.xii.1987
–
7.i.1988
,
R.I. Storey
&
Dickinson
, (25-059596
2 specimens
in
ANIC
)
;
Tully Falls State Forest
via
Ravenshoe
, [
17°47’0”S
,
145°34’0”E
],
20–21.iv.1973
,
R.I. Storey
(25-058502
1 specimen
in
QM
)
;
Tully Falls
,
17°47’0”S
,
145°33’27”E
,
8.xii.1989
-
5.i.1990
, (T44379
1 specimen
in
QM
)
;
Red Road
turnoff on
Tully Falls Road
,
17°49’19”S
,
145°32’42”E
,
8.xii.
198
9–5.i.1990
, (T44371
1 specimen
in
QM
)
;
Boulder Creek
track,
200–600 m
,
17°51’2”S
,
145°54’29”E
,
24–27.x.1983
, (T44390-T44397
8 specimens
in
QM
)
;
Laceys Creek
, [
17°52’0”S
,
146°4’0”E
],
40 m
,
11.vii.1971
,
R.W. Taylor J.E. Feehan
, (25-057995
1 specimen
in
ANIC
)
;
Tully
, [
17°53’0”S
,
145°55’0”E
],
200 m
,
17.x–22.xi.1994
, (25-059607
1 specimen
in
ANIC
)
;
Tully River Crossing
,
10 km
S Koombooloomba Dam
,
17°55’32”S
,
145°37’38”E
,
8.xii.1989
–
4.i.1990
, (T44375–T44378
4 specimens
in
QM
)
;
Mount Tyson
,
17°55’35”S
,
145°54’34”E
,
7.v.1983
, (T44327
1 specimen
in
QM
)
Description.
Uniformly brown in colour. Antennal clubs yellow to white.
Total length:
2.2–2.8 mm
.
Holotype
measurements: total length
2.4 mm
, elytral width
1.5 mm
.
Male
. Head. Surface smooth and nitid, with dense ocellate punctures each bearing a seta. Very broad U-shaped between clypeal teeth that are short but not upturned. Genal angle weakly defined. Dorsal part of the eyes narrow, separated by an interocular space approximately 11 times the eye width (34:3). Head 1.5 x wider than long (45:30).
Prothorax: Pronotal surface smooth and nitid, with dense ocellate punctures each bearing an obvious posteiorly directed seta. Punctures along the basal margin horseshoe shaped differing from those on disc. Anterior angles obtuse. Basal margin slightly angulate. Hypomeral surface reticulate and punctate. Hypomeral stria about two thirds the lenth of hypomeron with a row of fine setae present between hypomeral stria and lateral edge. Pronotum 1.8
x wider than long (68:38).
Elytra: Intervals reticulate with fine meshes, and with a double row of recurved setae. Striae geminate, appearing double. A well-developed humeral callus between striae 6 and 7, and a basal depression between striae 4 and 5. Stria 8 ending basally in a deep pit and lateral carina indented at this point. Elytra 1.05 x wider than long (87:82).
Legs: Apical digit of foretibiae broad with a small comb of setae on inner apex and very small apical spur on outer edge. Metatibiae with a notch on the inner edge.
Abdomen: Pygidium flat, reticulate, punctate and with setae and with an overhanging basal edge. Abdominal ventrites reticulate right across.
Pterothorax: Medial lobe of metaventrite punctate and narrowly margined between the mesocoxae. Lateral lobe of metaventrite with ocellate punctures. Meso-metaventral suture very slightly arched. Mesoventrite smooth with punctation. Mesepimeron reticulate and punctate. Metanepisternum reticulate and punctate.
Hind wings: Fully developed.
Female
. Metatibae without a notch on the inner edge, otherwise as per the
L
.
villosus
species group.
Distribution.
Known from rainforests of the Australian Wet Tropics bioregion from Cape Tribulation to Tully (
Fig. 20
). The majority of specimens have been collected from mid to high elevation forest (
500–1200 m
) but some are known from forest at sea level.
Comments.
Almost
550 specimens
were examined for this revision of which almost all have been collected in flight intercept traps, pitfall traps, or in berlesed leaf and log litter. The few specimens that have data for baited pitfalls indicate a preference of kangaroo dung, but other attractants include human faeces, carrion, mushroom, and fish baits.
Ebert
et al.
(2019)
only collected a single specimen of
L. villosus
in their 10-way food preference study and it came to a kangaroo dung baited pitfall trap. Specimens collected from Lambs Range and Longlands Gap (25-
060866
and 25-
060851
) have been sequenced and included in the phylogenetic analyses (
Gunter & Weir 2017
; Gunter
et al
. 2019). In the latter more comprehensively sampled study,
L. villosus
is recovered as sister to
L. pseudovillosus
new species
, in a monophyletic clade that also contained
L. vestitus
.