Synonymical Notes For Corylophidae And Cryptophagidae (Coleoptera: Cucujoidea)
Author
Leschen, Richard A. B.
Landcare Research Private Bag 92 170, 120 Mt Albert Road Auckland, NEW ZEALAND
LeschenR@landcare.cri.nz
Author
Bowestead, Stanley
Department of Entomology, The Manchester Museum The University, Manchester M 13 9 PL, ENGLAND
stan@bowestead.freeserve.co.uk
text
The Coleopterists Bulletin
2001
2001-09-01
55
3
312
316
journal article
10.1649/0010-065x(2001)055[0312:snfcac]2.0.co;2
1938-4394
4900371
Cryptophagidae
Circumtropical
Atomariinae
:
Cryptafricini
Anitamaria
Leschen, 1996: 625
.
Type
species:
Anitamaria thayerorum
Leschen, 1996
(original designation).
Scytomaria
Lyubarsky, 1998a: 2
.
Type
species:
Scytomaria himalaica
Lyubarsky, 1998
(original designation).
New synonymy.
Notes.
Type
specimens of
Scytomaria himalaica
are very similar to those that
Leschen (1996)
examined for his generic description of
Anitamaria
,
which included an undescribed species similar to
S. himalaica
also from
Nepal
(see appendix in
Leschen 1996
, deposited in MNHG). The synonymy results in the
new combination
Anitamaria himalaica
(Lyubarsky)
.
The phylogenetic classification of
Leschen (1996)
was amended subsequently by
Lyubarsky (1998b)
to include several new familygroup taxa. Critical assessment of all of the steps of
Lyubarsky’s (1998b)
analysis was not possible, though
Leschen’s (1996)
data matrix was modified (10% of the original characters were changed) and four new characters and two additional taxa (
Scytomaria
and
Hypophagus
Lyubarsky
) were added. Remarkably, the relationships among the higher taxa in the one tree presented by
Lyubarsky (1998b)
were not significantly different from those presented by
Leschen (1996)
with the exception of
Atomariinae
.
Lyubarsky (1998
b
)
splits the tribe
Cryptafricini Leschen
into three separate tribes:
Cryptafricini
, Microphagini Lyubarsky and Scytomariini Lyubarsky. Surprisingly,
Cryptafricini
was found to be paraphyletic because this tribe was strongly supported by 12 characters in the reference tree (and other trees) discussed by
Leschen (1996)
. Possibly, the paraphyly of
Cryptafricini
is the result of wrongly duplicating
Anitamaria
and
Scytomaria
in the revised data matrix. A more detailed evaluation of Ljubarsky’s (1998
b
) methods is beyond the scope of this paper though his analytical work on
Cryptophagidae
has been criticised in the Russian literature (Kirejtshuk and Kryzhanovskij 1998).
Lyubarsky (1998b)
increased the number of higher taxa contained in the two subfamilies of
Cryptophagidae
from six tribes to four supertribes, nine tribes, and seven subtribes. Included are new familygroup names for south temperate Cryptosomatulini, a group for which specimens were not examined by
Lyubarsky (1998
b
)
and is poorly known anyway (
Leschen 1996
). The proliferation of familygroup names seems unwarranted especially when critical specimens have not been examined.
Material Examined
(
SMNS
).
3,
Nepal
,
Myagdi Distr.
1995,
Myagdi Khola N. Dobang
,
2,8003,100 m
, 22.24.
V
.
Martens
&
Schawaller
;
Paratypus
,
Scytomaria himalaica
n. sp.
Lyubarsky
.