Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA Author Ferro, Michael L. Author Gimmel, Matthew L. Author Harms, Kyle E. Author Carlton, Christopher E. text Insecta Mundi 2012 2012-11-30 2012 260 1 8 journal article 10.5281/zenodo.5175284 1942-1354 5175284 Caulophilus dubius (Horn) ( Fig. 25 ) Range: throughout eastern United States : New York to Florida , west to Michigan and Texas . Habitat: under bark of dead trees and Vitis (grape) vine, in leaf litter and tree holes. Collection Method: searching under bark and sifting/Berlese leaf litter and rotten wood, emergence. Biology: unknown. Ferro et al. (2012a) : significantly more abundant in leaf litter and spring, indifferent to forest. Present Study: significantly more abundant in FWD1 and primary forest. References: Blatchley and Leng 1916 (as Allomimus dubius Horn ); Downie and Arnett 1996 ; Peck and Thomas 1998 ; Ciegler 2010 .