<document id="A73D7D1AFD27B36F7A6BA9DC6F1A5D0A" ID-DOI="10.1080/00222930210155701" ID-ISSN="1464-5262" ID-Zenodo-Dep="10100437" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629979948711" checkinUser="felipe" docAuthor="Cantero, A. L. Peña &amp; Vervoort, W." docDate="2003" docId="03FD87E371470906FDDBFC55FDDFBCFB" docLanguage="en" docName="JNatHist.37.22.2653-2722.pdf" docOrigin="Journal of Natural History 37 (22)" docSource="http://dx.doi.org/10.1080/00222930210155701" docStyle="DocumentStyle:443144A73DAE3E092C51A378E4CD0138.2:JNatHist.2003-2004.journal_article.1cover" docStyleId="443144A73DAE3E092C51A378E4CD0138" docStyleName="JNatHist.2003-2004.journal_article.1cover" docStyleVersion="2" docTitle="Staurotheca vervoorti" docType="treatment" docVersion="3" lastPageNumber="2712" masterDocId="FFC4FF9B717D093AFFC2FFA7FFCEBE30" masterDocTitle="Species of Staurotheca Allman, 1888 (Cnidaria: Hydrozoa: Sertulariidae) from US Antarctic expeditions, with the description of three new species" masterLastPageNumber="2722" masterPageNumber="2653" pageNumber="2710" updateTime="1699603448332" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2C115DB046725D2B09A70889A81EDA42">Species of Staurotheca Allman, 1888 (Cnidaria: Hydrozoa: Sertulariidae) from US Antarctic expeditions, with the description of three new species</mods:title>
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<mods:namePart id="A5D63D971A64B6D1CAD11E0EFADBA669">Cantero, A. L. Peña</mods:namePart>
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<mods:namePart id="57029F17B10E5DB56842183171D7B2EA">Vervoort, W.</mods:namePart>
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<mods:title id="EF73F3EF9F2745EF90682F3ED5AA52B3">Journal of Natural History</mods:title>
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<mods:part id="1E0651DD48B9E807CBA47FFD2D36E17E">
<mods:date id="1AFDD766836D412B2B4C05F61ACDEEB7">2003</mods:date>
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<mods:number id="AB9FE4A9B1D74BA9F2930B68C04E5248">2003-11-30</mods:number>
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<mods:number id="CCB9A5227B12A9CDDE5A0DBF9854180D">37</mods:number>
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<mods:number id="CFDC17B410C3E0CBEC5CA3868E549606">22</mods:number>
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<mods:identifier id="6BB6118D79E9CA9883DB6780C1E6A380" type="ISSN">1464-5262</mods:identifier>
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<treatment id="03FD87E371470906FDDBFC55FDDFBCFB" LSID="urn:lsid:plazi:treatment:03FD87E371470906FDDBFC55FDDFBCFB" httpUri="http://treatment.plazi.org/id/03FD87E371470906FDDBFC55FDDFBCFB" lastPageId="60" lastPageNumber="2712" pageId="58" pageNumber="2710">
<subSubSection id="C34E657E71470900FDDBFC55FC9FBA1A" pageId="58" pageNumber="2710" type="nomenclature">
<paragraph id="8BEB36F571470900FDDBFC55FBDCBA3A" blockId="58.[313,1224,1010,1177]" box="[537,1042,1010,1034]" pageId="58" pageNumber="2710">
<heading id="D0A3819971470900FDDBFC55FBDCBA3A" box="[537,1042,1010,1034]" centered="true" fontSize="10" level="2" pageId="58" pageNumber="2710" reason="3">
<taxonomicName id="4C544D7671470900FDDBFC55FBDCBA3A" authority="(El Beshbeeshy, 1991)" baseAuthorityName="El Beshbeeshy" baseAuthorityYear="1991" box="[537,1042,1010,1034]" class="Hydrozoa" family="Sertulariidae" genus="Staurotheca" kingdom="Animalia" order="Leptothecata" pageId="58" pageNumber="2710" phylum="Cnidaria" rank="species" species="vervoorti">
<emphasis id="B920EAE771470900FDDBFC55FCC4BA3A" bold="true" box="[537,778,1010,1034]" italics="true" pageId="58" pageNumber="2710">Staurotheca vervoorti</emphasis>
(El Beshbeeshy, 1991)
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BEB36F571470900FD1EFBB5FC9FBA1A" blockId="58.[313,1224,1010,1177]" box="[732,849,1042,1066]" pageId="58" pageNumber="2710">(figure 19)</paragraph>
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<subSubSection id="C34E657E71470900FEFBFB93FC1EBAA9" pageId="58" pageNumber="2710" type="reference_group">
<paragraph id="8BEB36F571470900FEFBFB93FC54BA7A" blockId="58.[313,1224,1010,1177]" box="[313,922,1075,1098]" pageId="58" pageNumber="2710">
<taxonomicName id="4C544D7671470900FEFBFB93FCE1BA7A" authority="El Beshbeeshy, 1991: 250 - 253" authorityName="El Beshbeeshy" authorityPageNumber="250 - 253" authorityYear="1991" box="[313,815,1075,1098]" class="Hydrozoa" family="Sertulariidae" genus="Thuiaria" kingdom="Animalia" order="Leptothecata" pageId="58" pageNumber="2710" phylum="Cnidaria" rank="species" species="vervoorti">
<emphasis id="B920EAE771470900FEFBFB93FE21BA79" box="[313,495,1076,1097]" italics="true" pageId="58" pageNumber="2710">Thuiaria vervoorti</emphasis>
El Beshbeeshy, 1991: 250–253
</taxonomicName>
, figure 64.
</paragraph>
<paragraph id="8BEB36F571470900FEFBFBE9FC7CBA4E" blockId="58.[313,1224,1010,1177]" pageId="58" pageNumber="2710">
<taxonomicName id="4C544D7671470900FEFBFBE9FD2BBA53" authority=": Vervoort, 1972: 127" authorityName="Vervoort" authorityPageNumber="127" authorityYear="1972" box="[313,741,1102,1124]" class="Hydrozoa" family="Sertularellidae" genus="Sertularella" kingdom="Animalia" order="Leptothecata" pageId="58" pageNumber="2710" phylum="Cnidaria" rank="species" species="edentula">
<emphasis id="B920EAE771470900FEFBFBE9FDC7BA53" box="[313,521,1102,1123]" italics="true" pageId="58" pageNumber="2710">Sertularella edentula</emphasis>
: Vervoort, 1972: 127
</taxonomicName>
, figure 39b–d; Blanco, 1994a: 154; 1994b: 199. 
<taxonomicName id="4C544D7671470900FEFBFBCEFE24BA4E" box="[313,490,1129,1150]" class="Hydrozoa" family="Sertulariidae" genus="Thuiaria" kingdom="Animalia" order="Leptothecata" pageId="58" pageNumber="2710" phylum="Cnidaria" rank="species" species="edentula">
<emphasis id="B920EAE771470900FEFBFBCEFE24BA4E" box="[313,490,1129,1150]" italics="true" pageId="58" pageNumber="2710">Thuiaria edentula</emphasis>
</taxonomicName>
: Stepan’yants, 1979: 92, pl. 17 figure 9A–V.
</paragraph>
<paragraph id="8BEB36F571470900FEFBFB24FC1EBAA9" blockId="58.[313,1224,1010,1177]" box="[313,976,1155,1177]" pageId="58" pageNumber="2710">
<taxonomicName id="4C544D7671470900FEFBFB24FC94BAA9" authority=": Pena Cantero et al., 1997: 337" authorityName="Pena Cantero" authorityPageNumber="337" authorityYear="1997" box="[313,858,1155,1177]" class="Hydrozoa" family="Sertulariidae" genus="Staurotheca" kingdom="Animalia" order="Leptothecata" pageId="58" pageNumber="2710" phylum="Cnidaria" rank="species" species="vervoorti">
<emphasis id="B920EAE771470900FEFBFB24FDDFBAA9" box="[313,529,1155,1177]" italics="true" pageId="58" pageNumber="2710">Staurotheca vervoorti</emphasis>
: Peña Cantero 
<emphasis id="B920EAE771470900FD77FB23FD2FBAA8" box="[693,737,1155,1177]" italics="true" pageId="58" pageNumber="2710">et al</emphasis>
., 1997: 337
</taxonomicName>
; 1999: 160.
</paragraph>
</subSubSection>
<subSubSection id="C34E657E71470906FEA3FB16FDDFBCFB" lastPageId="60" lastPageNumber="2712" pageId="58" pageNumber="2710" type="materials_examined">
<paragraph id="8BEB36F571470900FEA3FB16FD61BB79" blockId="58.[313,1269,1201,1736]" pageId="58" pageNumber="2710">
<emphasis id="B920EAE771470900FEA3FB16FDBBBAFA" box="[353,629,1201,1226]" italics="true" pageId="58" pageNumber="2710">Material examined. 22</emphasis>
/ 
<emphasis id="B920EAE771470900FDBCFB15FD7DBAFA" box="[638,691,1202,1226]" italics="true" pageId="58" pageNumber="2710">1592</emphasis>
, three stem fragments up to 
<quantity id="4CAC9B1071470900FBDAFB16FBBEBAF9" box="[1048,1136,1201,1226]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="58" pageNumber="2710" unit="mm" value="150.0">150 mm</quantity>
long, with gonothecae (
<collectionCode id="ED45AE3071470900FE0DFB76FDEABAD9" box="[463,548,1233,1257]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="58" pageNumber="2710" type="Museum">USNM</collectionCode>
1003292; RMNH-Coel. 30328); 
<emphasis id="B920EAE771470900FC5DFB76FC62BAD9" box="[927,940,1233,1257]" italics="true" pageId="58" pageNumber="2710">6</emphasis>
/ 
<emphasis id="B920EAE771470900FC77FB76FC13BAD9" box="[949,989,1233,1257]" italics="true" pageId="58" pageNumber="2710">377</emphasis>
, one fragmented colony (fragments up to 
<quantity id="4CAC9B1071470900FDD0FB56FD93BB39" box="[530,605,1265,1289]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="58" pageNumber="2710" unit="mm" value="70.0">70 mm</quantity>
long) (
<collectionCode id="ED45AE3071470900FD03FB56FCD8BB39" box="[705,790,1265,1289]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="58" pageNumber="2710" type="Museum">USNM</collectionCode>
1003293; RMNH-Coel. 30329; 
<collectionCode id="ED45AE3071470900FB59FB56FB3ABB39" box="[1179,1268,1265,1289]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="58" pageNumber="2710" type="Museum">MNCN</collectionCode>
2.03/298); 
<emphasis id="B920EAE771470900FE75FAB5FE0ABB19" box="[439,452,1298,1321]" italics="true" pageId="58" pageNumber="2710">7</emphasis>
/ 
<emphasis id="B920EAE771470900FE0FFAB6FE3BBB19" box="[461,501,1297,1321]" italics="true" pageId="58" pageNumber="2710">551</emphasis>
, one stem 
<emphasis id="B920EAE771470900FDB8FAB5FD5DBB19" box="[634,659,1298,1321]" italics="true" pageId="58" pageNumber="2710">ca</emphasis>
<quantity id="4CAC9B1071470900FD5CFAB7FD27BB19" box="[670,745,1296,1321]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="58" pageNumber="2710" unit="mm" value="65.0">65 mm</quantity>
high (
<collectionCode id="ED45AE3071470900FCFFFAB6FC5CBB19" box="[829,914,1297,1321]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="58" pageNumber="2710" type="Museum">USNM</collectionCode>
1003294); 
<emphasis id="B920EAE771470900FBD5FAB6FBEABB19" box="[1047,1060,1297,1321]" italics="true" pageId="58" pageNumber="2710">9</emphasis>
/ 
<emphasis id="B920EAE771470900FBEFFAB6FB9BBB19" box="[1069,1109,1297,1321]" italics="true" pageId="58" pageNumber="2710">724</emphasis>
, one stem 
<emphasis id="B920EAE771470900FB19FAB5FB3ABB19" box="[1243,1268,1298,1321]" italics="true" pageId="58" pageNumber="2710">ca</emphasis>
<quantity id="4CAC9B1071470900FEFBFA96FE5FBB78" box="[313,401,1329,1353]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="58" pageNumber="2710" unit="mm" value="100.0">100 mm</quantity>
high (
<collectionCode id="ED45AE3071470900FE22FA96FDFBBB79" box="[480,565,1329,1353]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="58" pageNumber="2710" type="Museum">USNM</collectionCode>
1003295).
</paragraph>
<paragraph id="8BEB36F571470900FEA3FAF5FDDFBBF8" blockId="58.[313,1269,1201,1736]" pageId="58" pageNumber="2710">
<emphasis id="B920EAE771470900FEA3FAF5FE26BB59" box="[353,488,1362,1385]" italics="true" pageId="58" pageNumber="2710">Description.</emphasis>
Stems up to 
<quantity id="4CAC9B1071470900FD5DFAF7FD39BB58" box="[671,759,1360,1385]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="58" pageNumber="2710" unit="mm" value="150.0">150 mm</quantity>
long, polysiphonic over a great extension. Branching dichotomous and in one plane. Branches strongly contracted at their base and originating at a position corresponding to a hydrotheca. Both stems and branches flattened.
</paragraph>
<paragraph id="8BEB36F571470900FEA3FA77FB18B838" blockId="58.[313,1269,1201,1736]" pageId="58" pageNumber="2710">Hydrothecae arranged in subopposite or alternate pairs, forming two longitudinal series (figure 19A, B). In these series, hydrothecae may be closely approximated.</paragraph>
<paragraph id="8BEB36F571470906FEA3F9B7FDDABEDD" blockId="58.[313,1269,1201,1736]" lastBlockId="60.[313,1269,181,715]" lastPageId="60" lastPageNumber="2712" pageId="58" pageNumber="2710">Hydrothecae cylindrical and almost straight (figure 19A–E); usually adnate to stem for their entire adcauline length (figure 19B, C, E). In the material from Stn 22/1592 and Stn 6/377 the free adcauline hydrothecal wall is inconspicuous (figure 19A, D). Abcauline hydrothecal wall straight or slightly concave. Aperture circular, tilted downwards and forming an acute angle with the long axis of branch, provided with a dish-shaped operculum. Rim even or uneven and slightly laterally depressed, sometimes with a few short renovations. Hydrotheca without mushroomshaped diaphragm.</paragraph>
<caption id="DF2B667D71460901FEFBF988FD21B8A4" ID-DOI="http://doi.org/10.5281/zenodo.10100475" ID-Zenodo-Dep="10100475" httpUri="https://zenodo.org/record/10100475/files/figure.png" pageId="59" pageNumber="2711" startId="59.[313,329,1583,1604]" targetBox="[316,1268,179,1557]" targetPageId="59">
<paragraph id="8BEB36F571460901FEFBF988FD21B8A4" blockId="59.[313,1268,1583,1685]" pageId="59" pageNumber="2711">
F. 19. 
<taxonomicName id="4C544D7671460901FE62F988FCBAB875" authority="(El Beshbeeshy, 1991)" baseAuthorityName="El Beshbeeshy" baseAuthorityYear="1991" box="[416,884,1583,1605]" class="Hydrozoa" family="Sertulariidae" genus="Staurotheca" kingdom="Animalia" order="Leptothecata" pageId="59" pageNumber="2711" phylum="Cnidaria" rank="species" species="vervoorti">
<emphasis id="B920EAE771460901FE62F988FDB4B875" box="[416,634,1583,1605]" italics="true" pageId="59" pageNumber="2711">Staurotheca vervoorti</emphasis>
(El Beshbeeshy, 1991)
</taxonomicName>
. (A, B) Two fragments of branches showing arrangement of hydrothecae; (C–E) hydrothecae (in E with gonotheca); (F) gonotheca. (A, E, F: Stn 22/1592; B, C: Stn 9/724; D: Stn 6/377.) Scale bars: 1 mm (A, B), 600 Mm (F), 400 Mm (C–E).
</paragraph>
</caption>
<paragraph id="8BEB36F571410906FEA3FF52FB3ABF7D" blockId="60.[313,1269,181,715]" pageId="60" pageNumber="2712">Presumed male gonothecae usually developing inside the hydrothecae (figure 19E), though also originating from their base (figure 19F). Gonotheca fusiform with a small circular aperture on an inconspicuous distal neck (figure 19E, F).</paragraph>
<paragraph id="8BEB36F571410906FEA3FEF3FBF1BFBC" blockId="60.[313,1269,181,715]" pageId="60" pageNumber="2712">Cnidome composed of microbasic mastigophores in two size groups: a larger (19.6–21.7×4.2–4.9 Mm) and a smaller group (7–7.7×2.8–3.5 Mm).</paragraph>
<paragraph id="8BEB36F571410906FEA3FE33FCD9BFDC" blockId="60.[313,1269,181,715]" pageId="60" pageNumber="2712">
<emphasis id="B920EAE771410906FEA3FE33FE02BF9C" box="[353,460,404,428]" italics="true" pageId="60" pageNumber="2712">Remarks.</emphasis>
El Beshbeeshy (1991) described gonothecae of this species from his Patagonian material. These ‘gonothecae’ may actually represent mollusc eggs deposited on the branches of 
<taxonomicName id="4C544D7671410906FD4CFE73FCDEBFDC" baseAuthorityName="El Beshbeeshy" baseAuthorityYear="1991" box="[654,784,468,492]" class="Hydrozoa" family="Sertulariidae" genus="Staurotheca" kingdom="Animalia" order="Leptothecata" pageId="60" pageNumber="2712" phylum="Cnidaria" rank="species" species="vervoorti">
<emphasis id="B920EAE771410906FD4CFE73FCDEBFDC" box="[654,784,468,492]" italics="true" pageId="60" pageNumber="2712">S. vervoorti</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BEB36F571410906FEA3FE53FBF2BC7C" blockId="60.[313,1269,181,715]" pageId="60" pageNumber="2712">
<emphasis id="B920EAE771410906FEA3FE53FC4ABC3C" box="[353,900,500,524]" italics="true" pageId="60" pageNumber="2712">
Ecology and distribution. 
<taxonomicName id="4C544D7671410906FD56FE53FC4ABC3C" baseAuthorityName="El Beshbeeshy" baseAuthorityYear="1991" box="[660,900,500,524]" class="Hydrozoa" family="Sertulariidae" genus="Staurotheca" kingdom="Animalia" order="Leptothecata" pageId="60" pageNumber="2712" phylum="Cnidaria" rank="species" species="vervoorti">Staurotheca vervoorti</taxonomicName>
</emphasis>
had been found at depths from 310 (El Beshbeeshy, 1991) to 
<quantity id="4CAC9B1071410906FD57FDB4FD2ABC1B" box="[661,740,531,556]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="60" pageNumber="2712" unit="m" value="1500.0">1500 m</quantity>
(Stepan’yants, 1979). We found it from 
<quantity id="4CAC9B1071410906FB7DFDB3FE62BC7B" metricMagnitude="3" metricUnit="m" metricValue="2.187" metricValueMax="2.727" metricValueMin="1.6469999999999998" pageId="60" pageNumber="2712" unit="m" value="2187.0" valueMax="2727.0" valueMin="1647.0">1647 to 2727 m</quantity>
. In our material fertile colonies were collected in March.
</paragraph>
<paragraph id="8BEB36F571410906FEA3FDF3FDDFBCFB" blockId="60.[313,1269,181,715]" pageId="60" pageNumber="2712">
<taxonomicName id="4C544D7671410906FEA3FDF3FD9FBC5B" baseAuthorityName="El Beshbeeshy" baseAuthorityYear="1991" box="[353,593,596,620]" class="Hydrozoa" family="Sertulariidae" genus="Staurotheca" kingdom="Animalia" order="Leptothecata" pageId="60" pageNumber="2712" phylum="Cnidaria" rank="species" species="vervoorti">
<emphasis id="B920EAE771410906FEA3FDF3FD9FBC5B" box="[353,593,596,620]" italics="true" pageId="60" pageNumber="2712">Staurotheca vervoorti</emphasis>
</taxonomicName>
is a Patagonian species. It had been reported from off the Patagonian coasts (Stepan’yants, 1979; El Beshbeeshy, 1991) and south-east of the 
<collectingCountry id="F343766571410906FEFBFD34FE36BC9B" box="[313,504,659,683]" name="Falkland Islands" pageId="60" pageNumber="2712">Falkland Islands</collectingCountry>
(Vervoort, 1972). Our material comes from Burdwood Bank and off 
<collectingCountry id="F343766571410906FEA3FD14FDC3BCFB" box="[353,525,691,715]" name="South Georgia and the South Sandwich Islands" pageId="60" pageNumber="2712">South Georgia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>