<document ID-CLB-Dataset="21337" ID-DOI="http://dx.doi.org/10.3897/zookeys.958.53865" ID-GBIF-Dataset="35a5c8a1-bf59-4069-9fb0-961971e7e6d2" ID-PMC="PMC7434803" ID-Pensoft-Pub="1313-2970-958-107" ID-Pensoft-UUID="377C3EF18E8951FD9599616E45E03C94" ID-PubMed="32863719" ID-ZooBank="AB196008154249D4B23E1892D2191C18" ModsDocID="1313-2970-958-107" checkinTime="1597178110450" checkinUser="pensoft" docAuthor="Ng, Ting Hui, Jeratthitikul, Ekgachai, Sutcharit, Chirasak, Chhuoy, Samol, Pin, Kakada, Pholyotha, Arthit, Siriwut, Warut, Srisonchai, Ruttapon, Hogan, Zeb S. &amp; Ngor, Peng Bun" docDate="2020" docId="0763C0E46C565B34B1B164E7325ABBA1" docLanguage="en" docName="ZooKeys 958: 107-141" docOrigin="ZooKeys 958" docSource="http://dx.doi.org/10.3897/zookeys.958.53865" docTitle="Pila pesmei" docType="treatment" docVersion="6" id="377C3EF18E8951FD9599616E45E03C94" lastPageNumber="107" masterDocId="377C3EF18E8951FD9599616E45E03C94" masterDocTitle="Annotated checklist of freshwater molluscs from the largest freshwater lake in Southeast Asia" masterLastPageNumber="141" masterPageNumber="107" pageNumber="107" updateTime="1732672174402" updateUser="ExternalLinkService">
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<mods:title id="6B5905EB0D80EA7845D64B12CFA5D417">Annotated checklist of freshwater molluscs from the largest freshwater lake in Southeast Asia</mods:title>
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<mods:namePart id="1D75CE3A6C9EFA31ED979427415DE0D3">Ng, Ting Hui</mods:namePart>
<mods:affiliation id="A764B2A0BAC6DFB5621BCB9EF2FBB84F">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, 117377, Singapore</mods:affiliation>
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<mods:namePart id="DF26B2785F4DCAA8EDBA32D5C599E729">Jeratthitikul, Ekgachai</mods:namePart>
<mods:affiliation id="371EFC206024DB7DF0B2A6A5BB404166">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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<mods:namePart id="01812302423BDA03EFB9F89F21EE35B6">Sutcharit, Chirasak</mods:namePart>
<mods:affiliation id="A01D2FAC231432B6A640EB4B2DD8C241">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, 254 Phayathai Road, Pathumwan, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="003EAD61F62FCF39DC127700E364726D">Chhuoy, Samol</mods:namePart>
<mods:affiliation id="367DE31C04E1ABB94ABA0272267513B0">Inland Fisheries Research and Development Institute (IFReDI), Fisheries Administration, No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia &amp; Wonders of the Mekong Project, c / o IFReDI, No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia</mods:affiliation>
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<mods:namePart id="95EE443BFE4AC2807605B52BB62C1B35">Pin, Kakada</mods:namePart>
<mods:affiliation id="E035E034908015F94A9B398994AC6354">Wonders of the Mekong Project, c / o IFReDI, No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia</mods:affiliation>
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<mods:namePart id="B30316B166D9C8BC778A4A7864EDE3E1">Pholyotha, Arthit</mods:namePart>
<mods:affiliation id="2A73E43F336757E12A189C70EFBDE835">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, 254 Phayathai Road, Pathumwan, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="0661922281085AC9DFED245431C8B2F1">Siriwut, Warut</mods:namePart>
<mods:affiliation id="1A297AC843BB7C0499B30A4E019A100E">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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<mods:namePart id="48E7CB5D9AE7064FF6E53D325670F838">Srisonchai, Ruttapon</mods:namePart>
<mods:affiliation id="9A9FC2C23757A6F483078641E12848BB">Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand</mods:affiliation>
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<mods:namePart id="C57F83829C384194BE51A8EE04A68073">Hogan, Zeb S.</mods:namePart>
<mods:affiliation id="9FC99B92B938280E79011D719EF499CE">Wonders of the Mekong Project, c / o IFReDI, No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia &amp; Department of Biology, University of Nevada, 1664 N. Virginia Street, Reno, NV 89557, USA</mods:affiliation>
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<mods:namePart id="2745F67B6C9524BD6DB9A2E63D537283">Ngor, Peng Bun</mods:namePart>
<mods:affiliation id="40FB394E64A92A9B5A74A7537EEE4CC4">Inland Fisheries Research and Development Institute (IFReDI), Fisheries Administration, No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia &amp; Wonders of the Mekong Project, c / o IFReDI, No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia</mods:affiliation>
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<treatment id="0763C0E46C565B34B1B164E7325ABBA1" ID-GBIF-Taxon="166239875" LSID="urn:lsid:plazi:treatment:0763C0E46C565B34B1B164E7325ABBA1" httpUri="http://treatment.plazi.org/id/0763C0E46C565B34B1B164E7325ABBA1" lastPageNumber="107" pageId="0" pageNumber="107" scope_class="Gastropoda" scope_family="Ampullariidae" scope_order="Architaenioglossa" scope_subClass="Caenogastropoda" scope_superFamily="Ampullarioidea">
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<paragraph id="FCA1C5E0EF112573CAA92D74DA5A98D2" pageId="0" pageNumber="107">
<taxonomicName id="9B529CF2F384F59B5811D5F3B2E19807" ID-CoL="7SQ2C" LSID="0763C0E4-6C56-5B34-B1B1-64E7325ABBA1" authority="(Morlet, 1889)" baseAuthorityName="Morlet" baseAuthorityYear="1889" class="Gastropoda" family="Ampullariidae" genus="Pila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pila pesmei" order="Architaenioglossa" pageId="0" pageNumber="107" phylum="Mollusca" rank="species" species="pesmei">Pila pesmei (Morlet, 1889)</taxonomicName>
<figureCitation id="8337CC7367E9B67AE342879180C0EA5B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Freshwater gastropods of the Tonle Sap basin, Cambodia A Pila gracilis B Pila pesmei C Pila virescens D Pomacea maculata E Filopaludina martensi cambodjensis F Idiopoma umbilicata G Mekongia rattei H Trochotaia trochoides I Bithynia siamensis goniomphalus J Wattebledia siamensis K Anentome cambojiensis L Anentome helena M Sulcospira housei N Melanoides tuberculata O Indoplanorbis exustus and P Radix rubiginosa. Scale bars: 10 mm, unless stated otherwise. Photographs by TH Ng." figureDoi="10.3897/zookeys.958.53865.figure5" httpUri="https://binary.pensoft.net/fig/441308" pageId="0" pageNumber="107">Fig. 5B</figureCitation>
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<subSubSection id="723811717A7E80872339BF6759E19FD3" pageId="0" pageNumber="107" type="reference_group">
<paragraph id="FEE86B4A7635D83F5CA575B4D10146D3" pageId="0" pageNumber="107">
<taxonomicName id="DB2BE462F28924B3A280D66D3E8E9D3B" authorityName="Morlet" authorityYear="1889" class="Gastropoda" family="Ampullariidae" genus="Ampullaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ampullaria pesmei" order="Architaenioglossa" pageId="0" pageNumber="107" phylum="Mollusca" rank="species" species="pesmei">Ampullaria pesmei</taxonomicName>
Morlet, 1899: 185, pl. 8, fig. 2. Type locality: &quot;Phnom-Penh (Cambodge)&quot;.
</paragraph>
<paragraph id="4FC678D12CD8CB6604715C1E888AE079" pageId="0" pageNumber="107">
<taxonomicName id="F76922325FE0511636F0776B37A33F71" baseAuthorityName="Morlet" baseAuthorityYear="1889" class="Gastropoda" family="Ampullariidae" genus="Pila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pila pesmei" order="Architaenioglossa" pageId="0" pageNumber="107" phylum="Mollusca" rank="species" species="pesmei">Pila pesmei</taxonomicName>
: 
<bibRefCitation id="85A20CEA2BEB86DAB56675FB767D010C" author="Brandt, RAM" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="107" pagination="1 - 423" refId="B8" refString="Brandt, RAM, 1974. The non-marine aquatic Mollusca of Thailand. Archiv fuer Molluskenkunde 105: 1 - 423" title="The non-marine aquatic Mollusca of Thailand." volume="105" year="1974">Brandt 1974</bibRefCitation>
: 51, pl. 5, figs 70, 71.
</paragraph>
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<subSubSection id="7F5A4543C8FC8235E42050FFFED9B2A3" pageId="0" pageNumber="107" type="materials_examined">
<paragraph id="1DB6BBE0C37B9B0C65ACC1220D0F0215" pageId="0" pageNumber="107">Material examined.</paragraph>
<paragraph id="33BD2345CEFD393728E1C0923C41E7B8" pageId="0" pageNumber="107">ZRC.MOL.015671.</paragraph>
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<subSubSection id="C2734F5CD7646D9472EA232EBACD59A0" pageId="0" pageNumber="107" type="distribution">
<paragraph id="C8C17947996B4F32843C290E6B1F035B" pageId="0" pageNumber="107">Distribution and habitat.</paragraph>
<paragraph id="788D99EE129DA15CA004965A69149D4C" pageId="0" pageNumber="107">Tonle Sap open area (locality no. 38).</paragraph>
</subSubSection>
<subSubSection id="247568F870547B0B384CB8503B1F5352" pageId="0" pageNumber="107" type="remarks">
<paragraph id="6109C43EBE8668A15C281994E3B32EB4" pageId="0" pageNumber="107">Remarks.</paragraph>
<paragraph id="A38201FB189208F408A60A3A5CA4437F" pageId="0" pageNumber="107">
Although 
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<emphasis id="1F41F631875976FCA8658A18C37D75CF" italics="true" pageId="0" pageNumber="107">Pila pesmei</emphasis>
</taxonomicName>
is widely distributed in Eastern and Northeastern Thailand (
<bibRefCitation id="3445CBC537DE48EC8569E4B93D392B6A" DOI="https://doi.org/10.1093/mollus/eyaa015" author="Ng, TH" journalOrPublisher="BioInvasions Records" pageId="0" pageNumber="107" refId="B65" refString="Ng, TH, Annate, S, Jeratthitikul, E, Sutcharit, C, Limpanont, Y, Panha, S, in press. . Disappearing freshwater apple snails (Ampullariidae): comprehensive update of their taxonomic status and distribution in Thailand. Journal of Molluscan Studies. https://doi.org/10.1093/mollus/eyaa015" title=". Disappearing freshwater apple snails (Ampullariidae): comprehensive update of their taxonomic status and distribution in Thailand. Journal of Molluscan Studies." url="https://doi.org/10.1093/mollus/eyaa015" year="in press">Ng et al. in press</bibRefCitation>
), the species does not appear to be as widespread as 
<taxonomicName id="3C26C1CE9ECA5DCC0233E2A60ADBCB93" baseAuthorityName="Lea" baseAuthorityYear="1856" class="Gastropoda" family="Ampullariidae" genus="Pila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pila gracilis" order="Architaenioglossa" pageId="0" pageNumber="107" phylum="Mollusca" rank="species" species="gracilis">
<emphasis id="6969A088E842C82258151F2E641E0CD6" italics="true" pageId="0" pageNumber="107">Pila gracilis</emphasis>
</taxonomicName>
in the Tonle Sap basin, especially during the dry season when our surveys were conducted. It is also uncertain if the species is still extant at its type locality of Phnom Penh. 
<taxonomicName id="01D30D9246C753141B12FF675979502D" baseAuthorityName="Morlet" baseAuthorityYear="1889" class="Gastropoda" family="Ampullariidae" genus="Pila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pila pesmei" order="Architaenioglossa" pageId="0" pageNumber="107" phylum="Mollusca" rank="species" species="pesmei">
<emphasis id="BB7D8EAF857593F757FB197440A7B74F" italics="true" pageId="0" pageNumber="107">Pila pesmei</emphasis>
</taxonomicName>
is harvested in abundance from the Tonle Sap Lake in the rainy season, with more than 380 tonnes recorded at five commercial landing sites in Kampong Chhnang Province within one year (as 
<taxonomicName id="48A19B73AD50E22B6221174AF1D4A84B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Gastropoda" family="Ampullariidae" genus="Pila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pila ampullacea" order="Architaenioglossa" pageId="0" pageNumber="107" phylum="Mollusca" rank="species" species="ampullacea">
<emphasis id="613179ED3E65DDBFB0ED072D2C62EAD6" italics="true" pageId="0" pageNumber="107">Pila ampullacea</emphasis>
</taxonomicName>
(Linneaus, 1758) in 
<bibRefCitation id="731F26DE06E9718E565417CBF113BE6D" author="Ngor, PB" journalOrPublisher="Catch and Culture" pageId="0" pageNumber="107" pagination="4 - 13" refId="B68" refString="Ngor, PB, Chhuon, K, Prak, LH, 2016. Cambodia completes first pilot study of Tonle Sap mollusc fishery. Catch and Culture 22: 4 - 13" title="Cambodia completes first pilot study of Tonle Sap mollusc fishery." volume="22" year="2016">Ngor et al. 2016</bibRefCitation>
, 
<bibRefCitation id="662446FC619ACA98F34517D2E36BE46A" DOI="https://doi.org/10.1051/limn/2018026" author="Ngor, PB" journalOrPublisher="Annales de Limnologie-International Journal of Limnology" pageId="0" pageNumber="107" pagination="34 - 10" refId="B72" refString="Ngor, PB, Sor, R, Prak, LH, So, N, Hogan, ZS, Lek, S, 2018d. Mollusc fisheries and length-weight relationship in Tonle Sap flood pulse system, Cambodia. Annales de Limnologie-International Journal of Limnology 54: 34 - 10, DOI: https://doi.org/10.1051/limn/2018026" title="Mollusc fisheries and length-weight relationship in Tonle Sap flood pulse system, Cambodia." url="https://doi.org/10.1051/limn/2018026" volume="54" year="2018 d">2018d</bibRefCitation>
). 
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<emphasis id="ACEF8500332B25EBC0F487F3EAB581FD" italics="true" pageId="0" pageNumber="107">Pila pesmei</emphasis>
</taxonomicName>
may be confused with 
<taxonomicName id="D962492D6F52CECB9D6B5A6B66AD8FD4" baseAuthorityName="Lea" baseAuthorityYear="1856" class="Gastropoda" family="Ampullariidae" genus="Pila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pila gracilis" order="Architaenioglossa" pageId="0" pageNumber="107" phylum="Mollusca" rank="species" species="gracilis">
<emphasis id="04FA2042F15F3DD8EAB0B1DC11BB7954" italics="true" pageId="0" pageNumber="107">Pila gracilis</emphasis>
</taxonomicName>
owing to morphological similarity among some populations (
<bibRefCitation id="BB6DF404DD2B46EDBAD5FEFDD1E8D71E" DOI="https://doi.org/10.1093/mollus/eyaa015" author="Ng, TH" journalOrPublisher="BioInvasions Records" pageId="0" pageNumber="107" refId="B65" refString="Ng, TH, Annate, S, Jeratthitikul, E, Sutcharit, C, Limpanont, Y, Panha, S, in press. . Disappearing freshwater apple snails (Ampullariidae): comprehensive update of their taxonomic status and distribution in Thailand. Journal of Molluscan Studies. https://doi.org/10.1093/mollus/eyaa015" title=". Disappearing freshwater apple snails (Ampullariidae): comprehensive update of their taxonomic status and distribution in Thailand. Journal of Molluscan Studies." url="https://doi.org/10.1093/mollus/eyaa015" year="in press">Ng et al. in press</bibRefCitation>
), and further extensive examination of both species in Cambodia would be necessary to resolve the true distribution of both species in the country. The 
<taxonomicName id="6B5B023E60CF94D268F9381FCABAEF31" baseAuthorityName="Morlet" baseAuthorityYear="1889" class="Gastropoda" family="Ampullariidae" genus="Pila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pila pesmei" order="Architaenioglossa" pageId="0" pageNumber="107" phylum="Mollusca" rank="species" species="pesmei">
<emphasis id="AAA2A3CE71B4B92586873D6F51E23FC7" italics="true" pageId="0" pageNumber="107">Pila pesmei</emphasis>
</taxonomicName>
morph from Tonle Sap Lake have relatively flat spires, compared to those in Thailand (
<bibRefCitation id="A023AD5B4073328CF3C39C1D54ADF20A" DOI="https://doi.org/10.1093/mollus/eyaa015" author="Ng, TH" journalOrPublisher="BioInvasions Records" pageId="0" pageNumber="107" refId="B65" refString="Ng, TH, Annate, S, Jeratthitikul, E, Sutcharit, C, Limpanont, Y, Panha, S, in press. . Disappearing freshwater apple snails (Ampullariidae): comprehensive update of their taxonomic status and distribution in Thailand. Journal of Molluscan Studies. https://doi.org/10.1093/mollus/eyaa015" title=". Disappearing freshwater apple snails (Ampullariidae): comprehensive update of their taxonomic status and distribution in Thailand. Journal of Molluscan Studies." url="https://doi.org/10.1093/mollus/eyaa015" year="in press">Ng et al. in press</bibRefCitation>
), and appear to also be found in Southern Vietnam (as 
<taxonomicName id="10F2E309BAD2928A7D532A71D46E614D" class="Gastropoda" family="Ampullariidae" genus="Pila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pila erythrochila" order="Architaenioglossa" pageId="0" pageNumber="107" phylum="Mollusca" rank="species" species="erythrochila">
<emphasis id="5BD8886A71EC3DC3AFA6CE9221C5CC20" italics="true" pageId="0" pageNumber="107">Pila erythrochila</emphasis>
</taxonomicName>
(Dautzenberg &amp; Fischer, 1905) in 
<bibRefCitation id="AFB5559E5024657617334FA42767ACF1" DOI="https://doi.org/10.1017/S0030605318000443" author="Ng, TH" journalOrPublisher="Oryx" pageId="0" pageNumber="107" pagination="230 - 238" refId="B66" refString="Ng, TH, Tan, SK, Ahmad, A, Do, VT, Joshi, RC, Wang, WY, Tan, HH, Yeo, DCJ, 2019. Not in the Least Concern: anthropogenic influences on a South-east Asian apple snail Pila scutata (Ampullariidae). Oryx 53: 230 - 238, DOI: https://doi.org/10.1017/S0030605318000443" title="Not in the Least Concern: anthropogenic influences on a South-east Asian apple snail Pila scutata (Ampullariidae)." url="https://doi.org/10.1017/S0030605318000443" volume="53" year="2019">Ng et al. 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>