A Revision of the Australian Funnel-web Spiders (Hexathelidae: Atracinae)
Author
Gray, Michael R.
text
Records of the Australian Museum
2010
2010-11-24
62
3
285
392
http://dx.doi.org/10.3853/j.0067-1975.62.2010.1556
journal article
10.3853/j.0067-1975.62.2010.1556
2201-4349
5298014
Genus
Hadronyche
—Key to species
1 Large spiders (male CL 10.0–12.0). Tibia II with a large, protuberant, rounded apophysis (
Fig. 30C
). PLS relatively long (PLSASL> 3×W) (
Figs. 1F
,
30D
) ..........................................................................
H. formidabilis
—— Spiders usually smaller. Tibia II apophysis less prominent or absent. PLS often shorter ............................................................................................................... 2
2 Tibia I and metatarsus I incrassate (
Fig. 106F
) ......................................................
H. adelaidensis
—— Tibia I and metatarsus I not incrassate ........................................................................................... 3
3 Palpal patella with many spines (12–16) (Fig. 103F). Leg II unmodified .....................................................................................................................
H. flindersi
—— Palpal patella spines absent or fewer (0–9). Leg II modified or unmodified ..................................................................................................................................... 4
4 Labium and sternum fused medially, labiosternal sigilla divided (Fig. 50A) .....................................................................................................................
H. mascordi
—— Labium and sternum not fused, labiosternal sigilla entire ............................................................. 5
5 Caput high and wide frontally, almost bulbous, rising relatively steeply from fovea (
CH
≥ 0.5×CW) (
Figs. 53A,C
; 54B). Metatarsus II unmodified (
Fig. 53E
) ......................................................................................
H. nimoola
—— Not in above combination .............................................................................................................. 6
6 Chelicerae with central tooth row short, basal (
Fig. 82D
). Labium almost as long as wide (LL/LW 0.96) (
Fig. 82A
) ................................................................................................................................
H. kaputarensis
—— Not in above combination .............................................................................................................. 7
7 Embolus short, with distal flanged part set at distinct angle to shaft (
Fig. 65B,C
) .........................................................................................................
H. jensenae
—— Distal embolus not as above .......................................................................................................... 8
8 Chelicerae with central tooth row short, basal. (
Fig. 97E
). Palpal middle haematodocha widely exposed (
Figs. 97H
,
100B
).
Caput
strongly raised (
Fig. 97B
). PMS apical segment short (
Fig. 97A
) (
lamingtonensis
species group) ...................................................................................................... 9
—— Chelicerae with central tooth row typically long (
Fig. 20G
), rarely short. Palpal middle haematodocha not or rarely exposed (
Fig. 48B
).
Caput
height and PMS length variable ........................................................................................ 13
9 Cuspules on labium relatively numerous (200–225). Cheliceral margins diverge distally (
Fig. 94C
) ..................................................................................
H. raveni
—— Cuspules on labium relatively sparse (40–100). Cheliceral margins subparallel (
Fig. 97E
) .................................................................................................................. 10
10 Male
tarsi
III
and
IV
unmodified ........................................................................
H. lamingtonensis
—— Male tarsi III and IV swollen (“boat-shaped”) than. tarsi I and II (
Fig. 3G
) ....................................................................................................................................... 11
11 Cheliceral promargin with few teeth (c. 3), in short basal row. Embolus straight ...............................................................................................................
H. anzses
—— Cheliceral promargin with several teeth (c. 6–12) in longer row (Figs. 99D, 102C). Embolus strongly curved (
Figs. 100A,B
; 102D) ............................................. 12
12 Male
tarsus I strongly spinose (24–43) .................................................................
H. annachristiae
—— Male tarsus I weakly spinose (5–7) .............................................................................
H. monteithi
13 Labium relatively short (LL/LW 0.70–0.80) (Figs. 57E, 60D). Tibia II and metatarsus II with apophyseal swellings ........................................................................... 14
—— Labium relatively long (LL/LW 0.85–1.05) (
Figs. 23A
, 68A). Tibia II and metatarsus II apophyses/apophyseal swellings present or absent ........................................................................................................................................... 15
14 Embolus short (BulbW/EmbL 0.80), not apically twisted (
Fig. 59B,C
) .............................................................................................................................
H. modesta
—— Embolus longer (BulbW/EmbL 0.60), twisted apically (
Fig. 62B,C
) ....................................................................................................................................
H. meridiana
15 Dorsal femur I and/or II with spines ............................................................................................ 16
—— Dorsal femur I, II with no spines ................................................................................................. 23
16 Tibia II with weak, proximad, spinose apophyseal swelling (Figs. 38F; 40H,I); metatarsus II lacks apophyseal swelling (
Fig. 40K
). Carapace relatively low (
CH
/CL 0.40) (Fig. 38B) ....................................
H. venenata
(Tasmania)
—— Not as above ................................................................................................................................. 17
17 Tibia II unmodified, with few spines (7–11); metatarsus II with small apophyseal swelling (Figs. 47G; 48H,I). Carapace relatively high (
CH
/CL 0.49) (Fig. 47B) ........................................................................................
H. monaro
—— Not as above ................................................................................................................................. 18
18 Tibia II and metatarsus II with apophyses/apophyseal swellings ................................................ 19
—— Tibia II and metatarsus II lacking apophyses/apophyseal swellings .............................................. 22
19 Embolus with strongly curved shaft (
Fig. 29B
) .........................................................
H. emmalizae
—— Embolus shaft weakly curved–straight (
Fig. 25B
) ...................................................................... 20
20 Male
(and female) coxae I,
II
with thorn-like setae anterobasally (Fig. 21H), and anterolateral sternal angles with tuft of bristle-like hair (
Fig. 20C,H
) ...........................................................................................................
H. cerberea
—— Not as above ................................................................................................................................. 21
21 Tibia II with spines distributed over apophysis and distoventral tibia (
Figs. 23C
;
25H,I
). PLS relatively short (PLSAPW/ L 0.49) (
Fig. 23E
) ...............................................................................................................
H. versuta
(part)
—— Tibia II with spines grouped on apophyseal swelling, few or none on distoventral tibia (
Figs. 34G
;
36H,I
). PLS relatively long (PLSAPW/ L 0.33) (
Fig. 34F
) ................................................................................
H. alpina
(part)
22 Tibia II and metatarsus II ventrally concave (bowed) (
Fig. 43I,K
). Embolus relatively wide (EmbmidW/ L 0.12) (
Fig. 43B
) ....................................
H. marracoonda
—— Tibia II and metatarsus II more or less straight, not bowed (
Fig. 46I, K
). Embolus narrower (EmbmidW/ L 0.08) (
Fig. 46B
) .........................................
H. tambo
(part)
23 Tibia II and metatarsus II with apophyses/apophyseal swellings ................................................ 24
—— Tibia II and metatarsus II lacking apophyses/apophyseal swellings .............................................. 25
24 Tibia II with spines distributed over apophysis and distoventral tibia (
Figs. 23C
;
25H,I
). PLS relatively short (PLSAPW/ L 0.49) (
Fig. 23E
) ...............................................................................................................
H. versuta
(part)
—— Tibia II with spines grouped on apophyseal swelling, few or none distributed onto distoventral tibia (
Figs. 34G
;
36H,I
). PLS longer (PLSAPW/ L 0.33) (
Fig. 34F
) .....................................................................
H. alpina
(part)
25 Embolus relatively broad (EmbmidW/L 0.12–0.16); moderately to strongly twisted distally (
Fig. 86B,C
) .......................................................................................... 26
—— Embolus relatively narrow (EmbmidW/L 0.06–0.09); weakly to moderately twisted distally (
Fig. 74B,C
) ..................................................................................... 29
26 Embolus base strongly offset from tegulum (
Figs. 70B
,
81B
) .................................................... 27
—— Embolus base weakly offset from tegulum (
Figs. 86B
,
89B
) ...................................................... 28
27 Embolus shaft strongly curved proximally, distal third strongly twisted (
Fig. 81B,C
). Tarsi III, IV swollen, “boat-shaped” ..........................................
H. lynabrae
—— Embolus shaft curvature and distal twisting less strongly developed (
Fig. 70B,C
). Tarsi III, IV unmodified ............................................................................
H. infensa
28 Tibia II sinuous ventrally, with ventral spines clustered in proximal half, (
Fig. 89H,I
). Tarsus I spines 8–38 ..........................................................................
H. walkeri
—— Tibia II not sinuous, ventral spines more scattered (
Fig. 86H,I
). Tarsus I spines often more numerous (30–63) .....................................................
H. macquariensis
29 Embolus slender, elongate (EmbL 3.53–4.15) (
Fig. 93B
) .....................................
H. levittgreggae
—— Embolus not as long (EmbL 2.18–3.06) ...................................................................................... 30
30 Labial cuspule number moderate (149–184). Bulb shorter than palpal tibia (BulbL/TibL 0.77) (
Fig. 46A
) ...........................
H. tambo
(part) (Gippsland,
Victoria
)
—— Labial cuspules numerous (246–393) Bulb almost as long as palpal tibia (BulbL/TibL 0.86–0.89) (
Fig. 77A
) .................................................................................... 31
31 Leg spines relatively few: tarsus I with 4–10, metatarsus I with 10– 16, tibia II with 3–7 spines .................................................................................................
H. orana
—— Leg spines more numerous: tarsus I with 23–46, metatarsus I with 31–56, tibia II with 13–25 spines ......................................................................................
H. valida
Generic and Species Descriptions Description
. With characters of
Atracinae
. Medium to large sized
Atracinae
. Carapace longer than wide, caput only
Atrax
O.P.-Cambridge
weakly raised (
CH
/CW 0.35–0.36) and narrow frontally
(CFW/CL 0.55–0.62). Cheliceral paturon relatively less
Atrax
O.P.-Cambridge 1877: 26.
Type
species
Atrax robustus
robust and narrower than in most
Hadronyche
spp.
(Figs. O.P.-Cambridge 1877 by monotypy. Simon, 1891: 302; 4B,D; 5B,E). Cheliceral groove narrow, V-shaped; central 1892: 175, 182, 186; 1903: 967–969. Hogg, 1901: 272; teeth in single short row confined to the basal half of the Rainbow, 1911: 121; 1913: 4; 1914: 252. Rainbow & groove, lateral marginal tooth rows run full length of groove. Pulleine, 1918: 165. Hickman, 1927: 63; 1964: 107. (Fig. 4F). Labium relatively short (LL/LW 0.70–0.75) Musgrave, 1927: 33; 1948: 77. Roewer, 1942: 207. (Fig. 4A). Sternum ovoid to narrow ovoid. Apical segment Bonnet, 1955: 772. Main, 1976: 70; 1981: 839; 1985: of posterior lateral spinnerets relatively long, digitiform 40. Gray, 1978: 121,125. Raven, 1980: 255; 1985: 55, 71; (PLSAPW 0.25–0.28) (Fig. 4G). Male femora I, II with Brignoli, 1983: 122. Gray, 1984: 441; Gray, 1988: 114.
dorsal spines (Fig. 4F). Male palp with patella width less than
Goloboff, 1993:6. Hedin & Bond, 2006: 467. Platnick,
or equal to femur width. Bulb with a long, slender, curved
2010.
Euctimena
.–Rainbow 1914: 248.
Type
species
Euctimena
embolus (EmbmidW/L 0.06–0.08) (
Fig. 6B
). Male tibia II
tibialis
Rainbow 1914
by monotypy. First synonymized with a large mid-ventral conical apophysis, the narrow apex by Musgrave, 1927: 33. Hickman, 1964: 107. Gray, 1978: surmounted by short, peg-like spines (
Fig. 6H,I
); metatarsus 125. Raven, 1980: 255. Main, 1985: 40. Gray, 1988: 114. II sinuous (proximoventrally concave) with a small mid-
Poikilomorpha
.–Rainbow 1914: 264.
Type
species
Poikilo-
ventral apophysis (
Fig. 6J,K
). Spermathecal sacs narrow,
morpha
montana
Rainbow 1914
by monotypy. First elongate (
Fig. 5G
). synonymized by Gray, 1978: 125. Raven, 1980: 255. Main, 1985: 40. Gray, 1988: 114.
Included species
.
Atrax robustus
O.P.-Cambridge,
A.
sutherlandi
n.sp.
,
A. yorkmainorum
n.sp.
Diagnosis
.
Differs from other atracine genera by presence of a large coniform apophysis on male tibia
II
(
Fig. 6H,I
);
Distribution
. Southeastern coast and highlands of
Australia
, and from
Hadronyche
by lower caput height. from the
Hunter River region
,
New South Wales
, to eastern
Victoria
.
Comments
. Burrows often with a weak or incomplete silk lining, typically sited under rocks and logs
Figure 5.
Atrax robustus
, female: (
A
) sternum, labium and maxilla; Figure 4.
Atrax robustus
, male: (
A
) sternum, labium and maxilla; (
B
) (
B
) cephalothorax and chelicerae, lateral; (
C
) leg III, prolateral, cephalothorax and chelicerae, lateral; (
C
) cheliceral groove teeth; patella, tibia, metatarsus; (
D
) spinnerets; (
E
) cephalothorax and (
D
) cephalothorax and chelicerae, dorsal; (
E
) palp, prolateral; (
F
) chelicerae, dorsal; (
F
) cheliceral groove teeth; (
G
) spermathecae. leg II, prolateral; (
G
) spinnerets. Scale lines 1 mm. Scale lines 1 mm.
Figure 6.
Atrax robustus
, male (
A,D–K
, AMS KS3173;
B,C
, AMS KS5170): (
A
) palp, prolateral; (
B
) bulb; (
C
) embolus, distal; (
D,E
) distal patella and tibia I—
D
, ventral,
E
prolateral; (
F,G
) metatarsus I—
F
, ventral,
G
, prolateral; (
H,I
) distal patella and tibia II—
H
, ventral,
I
, prolateral; (
J,K
) metatarsus II—
J
, ventral,
K
, prolateral. Scale lines: 0.3 mm, except
A,B,G
, 1.0 mm.
Atrax robustus
O.P.-Cambridge
Pennant Hills,
21 March 1979
, Thompson;
AMS
KS4045
,Penrith, 25August
1979,
A. Johnson
;
AMS
KS4399
, Potts Point,
5 July 1956
,
P. Berry Smith
;
AMS
KS4015
, Pymble,
14 March 1949
;
AMS
KS7466
, Queenscliff, April
Figures 4–7;
Tables 2
,
34
1981; AMS KS4027, Rhodes,
12 May 1930
, T. White; AMS KS4921,
Riverwood,
12 April 1966
,
R
.
E. Mascord
;
AMS
KS2995
,
Roseville
, 3
June
Atrax robustus
O.P.-Cambridge, 1877: 26.
Hogg, 1901: 273. 1979
,? Hidder;
AMS
KS4869
,
Rydalmere
,
13 March 1974
,
R
.
Hood
;
AMS
Musgrave
, 1927: 33.
Nishikawa
, 1976: 179.
Main
, 1985
:
KS4070
,
Ryde
,
January 1971
;
AMS
KS4007
,
St. Ives
,
12 May 1971
;
AMS
KS4083
,
Scotland Island
, l
April 1974
;
AMS
KS3457
,
Springwood
, 1978
,
41. Gray, 1978: 122. Gray, 1987: 314. Gray, 1988: 114.
A. McLean;AMS KS4922, Stanmore, 1949,January;AMS KS4971,Telopea,
Euctimena tibialis
.–Rainbow, 1914: 249. First synonymized
25 April 1971
,
K.P. Reid
;
AMS
KS2713
, The Oaks
,
Apr 1979
, P. McIntosh;
by Musgrave, 1927: 33. AMS KS4203,The Spit,
4 July 1948
, Page;AMS KS4013, Thornleigh;AMS
Poikilomorpha montana
.–Rainbow, 1914: 265. KS4373, Turramurra, l
April 1970
; AMS KS8664, Wahroonga, 31 January
Atrax montana
.–Main, 1985: 41 (Transferred from 1982,A.S. Martin;AMS KS4008,Waitara,
11 March 1949
;AMS KS10782,
Poikilomorpha
). First synonymized by Gray, 1988; 114. Warrawee,
12April 1983
, Ralleston;AMS KS5146,Waverley,
15 May 1973
, L. Gibson; AMS KS4367, Waverton,
4 March 1971
, Dr C. Browne; AMS
Types
.
Holotype
female:
BMNH 71
(
NMH
).
Type
dry, pinned, labelled
KS5131
,
West Pennant Hills
,
19 May 1973
,
R
.
McDonald
;
AMS
KS6148
, “
New Holland
”, no other data (not seen).
West Ryde
,
1 January 1980
,
W. Bennett
;
AMS
KS4000
, Whale Beach,
21 April 1974
;
AMS
KS4025
,
Willoughby
,
30 July 1973
,
A.C. Jones
;
AMS
KS3185
,
Winston Hills
,
21 July 1973
,
M.Gray
;
AMS
KS3404
,
AMS
KS5720
,
Other material examined.
New South Wales
(males)
: AMS KS2698,
Woonona, near Bulli,
December 1960
, S.H. Roberts;
AMS
KS4662
,
Austinmer,
6 January 1959
, N. Mitchell;
AMS
KS4812
, Avalon Beach, 14
Wyoming
, near
Gosford
,
14 March 1980
;
AMS
KS2670
, New Lambton, near
March 1976
, Porritt;
AMS
KS4837
,Avoca,
11 January 1975
;
AMS
KS4197
,
Newcastle
, 1979,
L. Hallinan
;
AMS
KS1150
,
Newcastle
,
13 March 1963
,
Balgowlah,
7 January 1960
,
Burch;
AMS
KS4946
, Balmoral,
18April 1974
,
B.C. Dyson; AMS KS4125, Wyong, 18 March 9178; AMS KS2923,
Robinson
&
Hoey
;
AMS
KS4902
,
Bankstown
,
14 April 1965
;
AMS
KS5137
,
Benandarah State Forest,
22 March 1979
–
25 April 1979
, C. Horseman;AMS
Baulkham Hills,
18 January 1973
, M. Gray;
AMS
KS4080
, Bayview,2 April
KS5205
, Foxground, near Gerringong,
10 April 1980
,
G. Wishart
; AMS
1974,
S. Thompson
;
AMS
KS2956
,
Beecroft
,
25 April 1979
,
D. Dean
;
AMS
KS5170
, Ryde,
33°49'S
151°06'E
,
20 May 1980
,
A. Brown.
New South
KS3999
, Belrose,
19 January 1967
, Anderson
;
AMS
KS1152
, “The Mists”
,
Wales
(females
):
AMS
KS4076
,
Artarmon
,
Apr 1927
,
J. Stewart
;
AMS
Berambing,
December 1976
, K.J. Street;
AMS
KS8376
,Berowra,December
KS4930
,Ashfield,
20 December 1930
,
E. Hudson
;
AMS
KS1826
,
Austinmer
,
1981,
J. Hingley
;
AMS
KS1852
, Bilgola Plateau,
26 May 1977
,
H. Newton
;
23 November 1959
,
F.E. Havand
;
AMS
KS1377
,
Avalon
,
15 May 1978
, L.
AMS
KS1148
,
Bilpin
,
17 January 1930
,
E.M.Hunt
;
AMS
KS4961
,
Burraneer
Reneman
;
AMS
KS5017
,
Avoca,
20 September 1968
,
N. Tweedale
;
AMS
Bay,
15 December 1972
;
AMS
KS1154
, Camden,
21 February 1974
, G.
KS4825
, Balgowlah,
2 May 1970
,
G. Smith
;
AMS
KS4987
,
Bankstown
, 19
Seymour
;
AMS
KS4914
,
Campsie
,
12 February 1975
,
R
.
N. Peck
;
AMS
April 1949
,
E.A. Cox
;
AMS
KS5140
,
Baulkham Hills
,
18 January 1973
, M.
KS4915
,
Caringbah
,
4 April 1972
,
M. Sinclair
;
AMS
KS4773
,
Cardiff
, 21
Gray
;
AMS
KS2716
,
Bayview
,
20 April 1979
,
B. Makins
;
AMS
KS4358
,
January 1979
;
AMS
KS4194
, Castlecrag,
May 1963
,
I. Webb
;
AMS
KS4484
,
Beecroft,
February 1928
, G.A. Heumann;
AMS
KS4903
, Belmore, 25 April
Charlestown,
1 February 1980
, G. Anderson;
AMS
KS4372
, Chatswood, 16
1924,
M. Codd
;
AMS
KS4803
,
Belrose
,
19 January 1967
,
Anderson
;
AMS
April 1972
,
W. Leonard
;
AMS
KS3990
,
Cheltenham
,
2 April 1974
, S.K.
KS1827
, Bilpin,
12 April 1972
,
R
.
McDonald
;
AMS
KS7564
,
Birmingham
Pearson
;
AMS
KS14383
,
Cherrybrook, l
April 1984
,
M. Backing
;
AMS
Gardens
, near
Newcastle
,
6 December 1978
,
R
.
Mascord
;
AMS
KS1674
,
KS2957
, Chester Hill,
27 May 1979
,
J.Paull
;
AMS
KS4395
,
Clarevale Beach
,
Blackheath,
24April 1948
,
R
.
McKay;
AMS
KS4887
,
Bowral;
AMS
KS8733
,
16 January 1976
,
K.H. Sinfield
;
AMS
KS4378
, Clifton Gardens, 20 April
Bundanoon,
February 1982
, B.Erdman;
AMS
KS4913
,Camden, 21 February
1972,
Dr C.A.Monticone
;
AMS
KS5013
, Cobbitty,
11 March 1959
,
T
.
Down
;
1974,
G. Seymour
;
AMS
KS4408
, Berowra,
15 September 1973
,
J. Disney
;
AMS
KS4066
, Collaroy, near Dee Why Lagoon,
14 December 1958
, A.
AMS
KS3524
, Carlingford,
26 September 1979
, J.Armitage;
AMS
KS3175
,
Kabanoff
;
AMS
KS5072
,
Cromer
,
6 April 1980
,
M. Duncan
;
AMS
KS3036
,
Castle Hill,
Apr 1979
,
F. Pearce
;
AMS
KS4206
, Castlecrag,
17 July 1973
, J.
Denistone,
26 April 1979
, M. Richison;
AMS
KS4001
, Dundas, 1 March
Bunce;
AMS
KS4242
,
Chatswood,
14 November 1928
,
R
. Barnes;
AMS
1976,
B. & A.C. Oldsen
;
AMS
KS7470
,
Dural
,
March 1981
;
AMS
KS2432
,
KS4325
, Cheltenham,
Apr 1972
,
D. Levy
;
AMS
KS4211
,
Clarevale Beach
,
East Maitland
, H
. Miller;
AMS
KS4815
, Eastwood,
30 March 1976
, D.
L. & M.Mortimer
;
AMS
KS4030
, Clifton Gardens,
21 April 1961
, Bradley
;
Griffith
;
AMS
KS7477
,
Elizabeth Bay
,
5 May 1981
, D.
T
.
Cave
;
AMS
KS1153
,
AMS
KS3188
, Collaroy,
February 1928
, E. Osborne;
AMS
KS4400
,
Engadine,
28 November 1972
, S. Couglin;
AMS
KS1132
, Epping, January
Collaroy Plateau,
July 1966
,
B. Tye
;
AMS
KS4892
,
Colo
Vale
,
July 1958
,
1977,
W.J. Bunton
;
AMS
KS1143
, Faulconbridge,
8 January 1973
, Stewart
;
D.H. Wallace
;
AMS
KS4905
,
Coogee
,
November 1926
,
Stokes
;
AMS
AMS
KS5694
,
Fox Valley Reserve
,near
Wahroonga
,
22 June 1980
,
B. Henke
;
KS4233
, Cremorne,
21 June 1927
, F.L. Grutzmacher;
AMS
KS4962
,
AMS
KS4071
,
Frenchs Forest
,
5 June 1972
,
N. Hailstone
;
AMS
KS13409
,
Galston,
2 February 1981
, G. Jackson;
AMS
KS10976
, Galston Gorge, 4
May 1983, G. Jackson;
AMS
KS1156
, Gladesville,
20 February 1970
,
McElwaine
;
AMS
KS13752
,
Gordon,
22 December 1983
,
M. Gray
;
AMS
KS4771
, Gosford,
21 January 1979
,
R
.
E. Mascord
;
AMS
KS4005
,
Grays
Point
,
5 February 1977
,
P. Poppett
;
AMS
KS4382
, Greenwich/Willoughby
;
AMS
KS1160
,
Hartley
Vale
,
8 December 1952
,
E.S. Miller
;
AMS
KS4297
,
Hazelbrook,
8 January 1980
, S. Clark;
AMS
KS5071
, Heathcote, 20 April
1980,
R
.
Witchard
;
AMS
KS2323
,
Helensburgh
,
6 January 1979
,
N. West
;
AMS
KS1157
, Hornsby,
3 March 1976
, Reed
;
AMS
KS4949
, Hurstville, 5
June 1972, A. Osten;
AMS
KS1161
, Kembla Heights,
11 November 1963
,
G. Pearce
;
AMS
KS8678
,
East Beach
,
Kiama area
,
20 January 1982
, D.
Bennedett
;
AMS
KS4377
,
Killara
,
February 1972
,
P.Mayman
;
AMS
KS3200
,
Kurrajong Heights,
30 November 1969
, G. Hunt;
AMS
KS4944
, Lakemba
,
7 January 1948
;
AMS
KS1159
, Lane Cove,
22 April 1930
,? Graham;
AMS
KS1146
, Lawson,
March 1956
,
R
. Schleicher;
AMS
KS8920
, Lindfield, 14
April
1982,
B. Henstridge
;
AMS
KS4078
, McMahons Point,
May 1972
, N.
Mayfield;
AMS
KS6067
,
Manly,
3 November 1980
;
AMS
KS10783
,
Matcham, near Gosford,
27 March 1983
, van Derbruggen;
AMS
KS1162
,
Minnamurra Falls
,
12 May 1964
,
R
. Mascord;
AMS
KS4022
,
Mona
Vale
, 26
February 1973,
T
. Govranik;
AMS
KS4948
, Mortdale,
13 February 1975
;
AMS
KS5314
,
Mosman
,
19 June 1980
,
S. Pfeiffer
;
AMS
KS4947
,
Mount
Colah,
25 January 1979
,M. Gray;
AMS
KS1155
,
Mount Irvine
, 14
November
1944, E.L. Troughton;
AMS KS
5016, Mount St. Thomas, near Wollongong,
17 June 1956
,
L. Evans
;
AMS
KS2146
,
Mount Wilson
,
15 November 1978
–11
December 1978, C. Horseman;
AMS KS
4009, Neutral Bay,
AMS KS
3189,
Newport
,
September 1959
,
A.J. Wright
;
AMS
KS13560
,
Northbridge
, 30
Figure
7.
Collection
records for
Atrax robustus
(squares) and
A
.
December 1983
,
J. Mayer
;
AMS
KS4950
,
Parramatta
,
25 March 1959
, W.G.
yorkmainorum
(circles).
Ashford
;
AMS
KS4951
,
Peakhurst
,
24 April 1945
,
H. Inder
;
AMS
KS2712
,
Cronulla,
13 January 1971
;
AMS
KS4828
,
Dee Why
,
May 1932
,
Rowe
;
Diagnosis
. CL
9.22–12.30 (male).
Differs
from other species
AMS
KS4804
,
Dundas
,
January 1975
,
M. Denham
;
AMS
KS4865
,
Dural
, by having a relatively spinose palpal tibia (4–11) with dorsal
18 January 1971
,
M. Thornley
;
AMS
KS3319
,
Eastwood
,
8 July 1979
, J.
Cullen;
AMS
KS4829
,
Elanora
,
1 March 1967
, S. Marshall;
AMS
KS4989
, spines present.
Differs
from
A. sutherlandi
by having a more
Elizabeth Bay
,
9 January 1958
, D.
T
.
Cave
;
AMS
KS1830
,
Emu Plains
, 30 elongate embolus (BulbW/EmbL 0.30) (
Fig. 6B
), and from
July 1957
, G.D. Sutherland;
AMS
KS4770
,
Enfield
,
5 June 1905
;
AMS
A. yorkmainorum
by larger body size (CL 9.22–12.30) and
KS4546
,
Epping
,
17 February 1980
, J. Brand;
AMS
KS4990
,
Five Dock
, shallower offset of embolus base from tegulum (
Fig. 6B
)
.
May 1928
,
R
.
Williams;
AMS
KS4361
, Forestville,
19 August 1971
;
AMS
KS8868
, Freeman’s Reach, near Windsor,
26 January 1982
,
M. Gray
;
AMS
KS8377
,
French’s Forest
,
24 October 1981
,
R
.
Scott
;
AMS
KS4661
,
Male
(
AMS
KS4016
,
Gordon
,
New South Wales
,
32°38'S
Galston
,
3 February 1980
;
AMS
KS4963
,
Garrie Beach
,
7 October 1965
,
148°39'E
,
F. Hatton
,
10 December 1949
). —
Size
. Carapace
R
.
Mascord
;
AMS
KS4839
,
Gosford
,
25 September 1964
, E.Worrell;
AMS
length 10.20, width 8.70.
Abdomen
length 9.52, width
KS4207
,
Greenwich
,
5 May 1970
, K. Burns;
AMS
KS4966
,
Guildford
7.38. —
Colour
. Basic
colour pattern. —
Carapace
. Longer West
,
27 August 1957
, S.
R
.
Kirkwood
;
AMS
KS1220
,
Hazelbrook
, 23
November 1976, G. Weatherspoon;
AMS
KS1675
,
Hazelbrook
,
February
than wide.
Height
2.86.
Frontal
width 5.14.
Fovea
narrow, 1976
;
AMS
KS4985
,
Heathcote
,
26 June 1975
,
R
.
Witchard
;
AMS
KS4967
, procurved.
Mid-dorsal
cephalic setae reach fovea.
Anterior Helensburgh
,
11 June 1951
, L. Reed;
AMS
KS4225
,
Hornsby
,
7 May 1940
; strial setae numerous. Anterolateral carapace angle with
AMS
KS1208
,
Jamieson Valley
,
22 June 1953
,A.W.Gayley;
AMS
KS4065
, small, weak bristles. —
Eyes
. Central
eye region weakly
Killara
,
8 March 1928
;
AMS
KS5078
,
Kingsgrove
, 1952
;
AMS
KS4993
, raised.
Eye
group width 1.85.
Median
ocular quadrangle
Kingswood
,
May 1962
;
AMS
KS4410
,
Kuring-ai Chase National Park
, 23
January 1980, M. Gray;
AMS
KS1310
,
Kurrajong
,
February 1978
;
AMS
length 0.70; anterior width 0.76, posterior width 1.13.
KS4945
,
Lakemba
,
17 March 1930
;
AMS
KS4853
,
Lane Cove
, 15
March Diameters
: AME 0.28, ALE 0.58, PLE 0.38
,
PME
0.26. 1973;
AMS
KS4968
,
Leichhardt
,
2 September 1913
, D. Hunter;
AMS
—
Chelicerae
. Cheliceral
groove narrow, weakly divergent
.
KS1834
,
Linden
,
3 September 1950
, K. Baames;
AMS
KS5031
,
Lindfield
,
Groove
length 2.92, middle width 0.41.
Cheliceral
teeth: 18
29 September 1962
, M. &
V
.
Gregg
;
AMS
KS4234
,
McMahons Point
,
Apr
1972, J.
Suich
;
AMS
KS6025
,
Manly
,
Manly Hospital
,
3 November 1980
; central, occupying proximal half of groove; 12 prolateral;
AMS
KS4907
,
Matraville South
,
20 October 1952
,
A. Brown
;
AMS 13
retrolateral. —
Labium
. Much
wider than long, apically
KS4238
,
Meadowbank
,
March 1963
,
R
.
H.
Hall
;
AMS
KS1829
,
Megalong
indented.
Length
1.08, width 1.68.
Labiosternal
sigilla
Valley
,
R
.
lbow;
AMS
KS4856
,
Middle Harbour
,
6 September 1933
, broad, entire.
Cuspules
c. 300, number high, in crescentic
Kaufman
;
AMS
KS4896
,
Mittagong
,
30 December 1975
, M. &
V
.
Gregg
; transverse band. —
Sternum
. Ovoid
, long.
Length
5.45,
AMS
KS4360
,
Mona
Vale
,
9 November 1975
,
J. Walker
;
AMS
KS1125
,
Morpeth;
AMS
KS4204
,
Mortdale
,
31 January 1974
, C. Stewart;
AMS
width 3.88.
Posterior
sigilla ovoid, broad.
Strong
bristles
KS4403
,
Mosman
,
3 March 1932
, Patton;
AMS
KS3693
,
Mount Colah
, 29 on posterior margin. —
Palp
. Bulb
about as long as tibia.
July 1979
, J. Fairlie;
AMS
KS4189
,
Mount Fairy
,
25 November 1979
;
AMS
Tegular region
rather small, slightly wider than long.
Base
KS1835
,
Mount Irvine
,
14 November 1944
, E. Troughton;
AMS
KS2960
, of embolus weakly offset from tegulum.
Embolus
shaft
Mount Keira
,
12 May 1979
, Walsh;
AMS
KS4832
,
Mount Kuring-ai
, 5
November
1972;
AMS
KS4808
,
Naremburn
,
26.ii.1961
,
M.Fairburn
;
AMS
long, gently curved, strongly tapered.
Distal
embolus
KS1126
,
Narara
,
20 March 1930
,
B.
R
.
Everingham
;
AMS
KS4405
, weakly twisted, ejaculatory groove wide.
Bulb
length
Narrabeen
,
29 September 1957
,
L.B. Trapp
;
AMS
KS4046
,
Neutral Bay
, 3 4.62, width 1.12.
Embolus
length 3.60, midwidth 0.25.
November 1979
;
AMS
KS797
,
Newport
,
7 December 1977
;
AMS
KS4064
,
Length
of femur 4.35, patella 1.96, tibia 4.71.
Width of Normanhurst
,
6 April 1969
,?
Dymock
;
AMS
KS4085
,
Northbridge
, 26 tibia 1.80.
Spination
: femur 7, patella 5, tibia 10 (dorsal
July 1975
,
D. Dickerson
;
AMS
KS3401
,
Northmead
,
March 1973
, J.
Deviana;
AMS
KS4195
,
North Narrabeen
,
11 June 1952
,
L.J. Lizara
;
AMS
spines present). Some distal femoral bristles and spines
KS4232
,
North Sydney
,
May 1925
,
Dr S.J.W. Moreau
;
AMS
KS4908
, sinuous. —
Legs
.
4123
.
Oatley,
15 February 1963
;
AMS
KS4858
, Palm Beach, l
February 1951
,
H.
R
.
Swaine
;
AMS
KS4909
,
Parramatta
,
21 March 1910
;
AMS
KS1131
,
Leg Femur Patella Tibia Metatarsus
Tarsus Total Peakhurst,
15 November 1969
, D. Fantom;
AMS
KS1837
,
Pearl Beach
, 1 8.43 4.21 6.36 6.69 3.87 29.56
24.iii.1966
, J. Cooks;
AMS
KS1930
,
Pennant Hills
,
6 October 1978
, S. 2 7.75 3.86 6.36 6.28 3.89 28.14 el-Issa;
AMS
KS4045
,
Penrith
,
25 July 1979
, A. Johnson;
AMS
KS4969
,
3
7.14 3.18 5.15 6.03 3.94 25.44
Penshurst
,
R
.
Pollett
;
AMS
KS1844
,
Picton
,
November 1946
, J.K.
McFarlane;
AMS
KS4072
,
Pittwater
, 1927,
A. Musgrave
;
AMS
KS4824
,
4
8.32 3.81 6.80 7.41 4.40 30.74
Point Clare
,
T
.A.
Trewheeler
;
AMS
KS4910
,
Potts Point
,
5 October 1969
,
C.
Catterall
;
AMS
KS4931
,
Punchbowl
,
17 July 1947
, F:
J. Donald
;
AMS
Tibia I
width 1.50.
Femora
I and
II
with dorsal spines.
Leg
KS4809
,
Pymble
,
November 1971
,
A. Ritchie
;
AMS
KS1850
,
Richmond, I
unmodified, tibial and metatarsal spines numerous.
Tibia
12 October 1977
,
Ambulance Station
;
AMS
KS3168
,
Roseville
, 18
January
II
with large, conical, apophysis set just proximal of centre. 1948, H.
Chadwick
;
AMS
KS4964
,
Royal National Park
,
October 1966
,
R
.
Mascord
;
AMS
KS4911
,
Rushcutters Bay
,
24 May 1971
,
S.J. Penlington
; A few ventral tibial spines placed proximally, absent in
AMS
KS4810
,
Ryde
,
28 October 1969
,
J. Garden
;
AMS
KS4084
,
St. Ives
, distal half (except apical ventral spines).
Distal
tibia
II
March 1967
;
AMS
KS4912
,
St. Peters
,
2 February 1959
,
D.J. Gilks
;
AMS
ventrally concave.
Metatarsus
II
sinuous, curved proximally,
KS5240
,
Scotland Island
,
29 February 1980
, J.
Lowry
;
AMS
KS5023
,
ventral spines placed upon and distal to a small, subcentral
Somersby
,
J Brown
;
AMS
KS1839
,
Springwood
,
4 August 1971
;
AMS
KS1135
,
Sydney
,
17 November 1969
;
AMS
KS2715
,
Terry Hills
, 30
April
apophysis.
Anterior
coxal hairs unmodified.
Scopulae
1979
,
R
.
Gleeson
;
AMS
KS4891
,
The Oaks
,
25 February 1956
;
AMS
legs I–
IV
: tarsus, weak; distal metatarsus, absent to weak.
KS4899
,
Thirroul
,
September 1933
, D. King;
AMS
KS796
,
Thornleigh
, 8
Tarsal
claw teeth legs I
,
II
: superior 13, 11; inferior 2,
2. January 1978
;
AMS
KS4844
,
Tuggerah
,
September 1927
,
H.W. Legge
;
Trichobothria
legs I
,
II
: tarsus 20, 19; metatarsus 15, 12;
AMS
KS4376
,
Turramurra
, 1928, H.S.
Wales
;
AMS
KS13791
,
Wahroonga
, tibia p9 r9, p8 r8. —
Leg
spination
.
Leg I
; femur 9 (d6 p3),
2 February 1984
,A.S. Martin;
AMS
KS4845
,
Wamberal
,
4 December 1972
,
J.N. Kgour;
AMS
KS4243
,
Warrawee
,
24 December 1972
,? Rickets;
AMS
patella 13 (p2), tibia 51 (p8), metatarsus 29, tarsus 20.
Leg
II
:
KS4063
,
West Pennant Hills
,
15 May 1973
, S. Zucher;
AMS
KS6069
,
West
femur 12 (d6 p6), patella 6 (p2–3); tibia 18 (p4), metatarsus
Ryde
,
10 November 1980
;
AMS
KS4846
,
West Wallsend
,
30 May 1956
,
J. 22(p1), tarsus 23.
Leg
III
: femur d7 (plus strong bristles),
Donne
;
AMS
KS4847
,
Weston
,
4 November 1969
, I. Waugh;
AMS
KS4357
, patella 5 (p3 r2), tibia 15 (p2 r3), metatarsus 25, tarsus 25.
Willoughby
,
17 August 1969
;
AMS
KS5151
,
Winston Hills
, 9
September
1972,
J. Deviana
;
AMS
KS1848
, Wirrimbirra Sanctuary,
9 July 1969
,
M.N. Leg
IV
: femur 0 (long bristles present), patella r1, tibia 18 (p1 Fackender;
AMS
KS4862
,Woolwich,
May 1927
,
H. Switzer
;
AMS
KS4848
, r4), metatarsus 20, tarsus 21. —
Abdomen
. Posterior lateral Woy Woy,
13 July 1933
,
B. Hynson
;
AMS
KS8295
, Yagoona, February spinnerets, lengths: total 4.68; basal segment 1.52; middle 1981;
AMS
KS1274
, Yerranderie,
20 February 1978
,
H. Kenny.
1.08; apical 2.08. Apical segment width 0.60
.
Table 2
. Male morphological data—
Atrax robustus
(n = 24).
character |
range |
mean |
character |
range |
mean |
character |
ratio |
SD |
CL* |
9.22–12.30 |
10.48 |
Mt1S |
24–32 (p0–1) 27 |
CW/CL |
0.89 |
0.033 |
CW |
8.27–10.59 |
9.28 |
Ta1S |
12–27 |
19 |
CH/CW |
0.35 |
0.025 |
CH |
2.86–4.06 |
3.28 |
Ti2S |
13–27 (p2–5) 18 |
CFW/CL |
0.55 |
0.038 |
CFW |
4.49–7.38 |
5.78 |
STC2teeth |
10–15 |
13 |
CHGW/L |
0.14 |
0.015 |
ChGL |
2.84–3.60 |
3.09 |
BulbL |
4.08–9.06 |
5.79 |
SW/SL |
0.74 |
0.043 |
ChGW |
0.41–0.55 |
0.44 |
EmbL |
3.28–5.26 |
4.25 |
LL/LW |
0.75 |
0.052 |
ChGCT |
10–29 |
20 |
BulbW |
1.12–1.48 |
1.28 |
PLSAPW/L |
0.25 |
0.032 |
LL |
1.08–1.62 |
1.41 |
EmbmidW |
0.20–0.30 |
0.27 |
BulbW/EmbL |
0.30 |
0.055 |
LW |
1.60–2.16 |
1.87 |
PalpTibL |
4.65–6.61 |
5.53 |
EmbmidW/L |
0.06 |
0.012 |
CUSPS* |
261–494 |
340 |
PalpTibW |
1.59–2.08 |
1.83 |
PalpTibW/L |
0.33 |
0.035 |
SL |
5.30–6.56 |
5.81 |
PalpTibS* |
4–11 |
8 |
BulbL/Tib L |
0.97 |
0.068 |
SW |
3.88–5.83 |
4.31 |
PalpPatS* |
1–8 |
5 |
PLSAPW |
0.46–0.64 |
0.53 |
PalpFem* |
5–9 |
6 |
PLSAPL |
1.64–2.40 |
2.03 |
Fe1S |
2–10 |
6 |
Pa1S |
6–13 (p2–4) |
10 |
Female
(
AMS
KS13470
,
Gordon, N.S.W.
,
32°38'S
148°39'E
, 3 (p2 r1), tibia 8 (p2 r2), metatarsus 23 (p9 r2), tarsus 17.
1 November 1983
, C. Horseman). —
Size
. Carapace length Leg
IV
; femur 0, patella r1, tibia 6 (p1 r2), metatarsus 22
12.63, width 10.41. Abdomen length 14.82, width 12.07. (p6 r2), tarsus 25. —
Abdomen
. Posterior lateral spinnerets-
—
Colour
. Basic colour pattern. —
Carapace
. Much longer lengths: total 6.20; basal segment 2.40, middle 1.32, apical than wide. Height 2.72, frontal width 8.28. Cephalic length 2.48. Apical segment width 0.77. —
Genitalia
. Spermathecae
8.50. Fovea narrow, procurved, anterior margin slightly long, slender; gently curved toward midline. Basal separation indented. Mid-dorsal cephalic setae numerous, reach back twice the width of a spermatheca. Length 1.50, width 0.41.
to fovea. Anterior strial setae numerous. Strial and marginal setae numerous, rather short. Anterolateral carapace angle
Distribution
. Sydney, Central
Coast
and
Illawarra
regions with several short, weak bristles. —
Eyes
. Central eye region and west to the Blue Mountains (Fig. 7).
slightly raised. Eye group width 2.60. Diameters:AME 0.26,
ALE 0.60, PLE 0.44,
PME
0.36 Interdistances: AME–AME
Comments
. Specimens from the Hunter River/
Newcastle
0.39, AME–ALE 0.27, ALE–PLE 0.27, ALE–
PME
0.14, region are typically larger than other individuals (male CL
PME
–
PME
1.05. Median ocular quadrangle length 0.90, c.
12 mm
) and the palpal tibia and embolus may be more
anterior width 0.94, posterior width 1.73. —
Chelicerae
. elongate.
Groove narrow, margins diverging distally; groove length
3.70, middle width 0.59. Cheliceral teeth: 29 central,
Mating position in
A. robustus
. Mating was observed for
occupying basal half of groove; 15 prolateral; 14 retrolateral.
A. robustus
(Gray, 1986)
, a species with well-developed
—
Labium
. Wider than long, anterior margin weakly apophyses on tibia and metatarsus
II
. The position adopted
indented. Length 2.18, width 2.64. Cuspules occupying during its mating is almost identical to that recorded for
H.
central two thirds of labium. Labiosternal sigilla entire. —
formidabilis
(
Fig. 1F
: Walker, in Mascord, 1980; Walker,
Sternum
. Ovoid. Length 6.89, width 5.17. Posterior sigilla 1982), another species with large apophyses, the
A. robustus
ovoid: length 1.10, width 0.60. —
Palp
. Spination: patella male, uses his second legs to clasp the basal parts of the
p1 (bristle-like), tibia 5, tarsus 12. Trichobothria: tibia p
11 female
femora
II
, locking them in the cavity between
r10, tarsus 22. Tarsal claw with 10 teeth. —
Legs
. 4123. Legs the tibial and metatarsal apophyses. At the same time he
I,
IV
subequal. braces his first legs against the chelicerae and palps of the
Leg Femur Patella Tibia Metatarsus Tarsus Total rearing female, pushing her up and backwards. The main
1 9.50 5.32 7.32 6.40 3.44 31.98 difference in the
H. formidabilis
mating was the distal
2 8.45 4.88 6.15 5.79 3.28 28.55 crossing of the bracing first legs (
Fig. 1F
)—this was not
3 7.46 3.94 4.80 5.66 3.69 25.55 seen in the
A. robustus
matings. Hickman (1964) described
4 9.00 4.88 6.71 7.18 4.22 31.99 mating in
H. infensa
, a species lacking leg
II
apophyses. The accompanying plate shows the spiders in an almost
Tibia I
width 2.30. Metatarsus I proximal width 1.44. Coxae identical position to that observed for
A. robustus
(although
I,
II
with short, anteromedial bristle-like setae. Tarsal claw Hickman’s notes are somewhat contradictory).
teeth legs I,
II
: superior 11, 11; inferior
III
,
IV
. Trichobothria Snazel & Allison (1989) noted the marked similarities legs I,
II
: tarsus 24, 24; metatarsus 21, 19; tibia p10 r9, p9 between mating in
H. formidabilis
and
Macrothele
, another
r10. —
Leg spination
. Leg I: femur p1, patella p1, tibia 3 (p2), genus in which leg
II
modifications are present or absent.
metatarsus 11, tarsus 15. Leg
II
: femur p2, patella p2, tibia Coyle (1986) observed similar mating behaviour in
Euagrus
5 (p2), metatarsus 14, tarsus 14. Leg
III
: femur 0, patella sp., a diplurid genus with tibia
II
apophyses.
Atrax yorkmainorum
n.sp.
Diagnosis
. CL 7.39–9.22 (male). Usually smaller then
A.
robustus
and differs from it by palp lacking dorsal tibial
Figures 7–10;
Tables 3
,
34
spines, embolus base more strongly offset from tegulum,
and apical part of embolus set at angle to strongly curved
Etymology
. The species is named for Dr Barbara
York
Main embolic shaft (
Fig. 10B,C
). Differs from
A. sutherlandi
by and Professor Bert Main, in recognition of their remarkable its more elongate, slender and curved embolus. achievements in Arachnology and Ecology.
Male
(
holotype
). —
Size
. Carapace length 8.42, width
Types
.
Holotype
male:
AMS
KS1042
,
Batlow
,
New South Wales
,
35°30'S
7.48.
Abdomen
length 8.50, width 6.90. —
Colour
. Basic
148°08'E
, H.
V
.
Smith
,
16 June 1951
.
Paratypes
(
AMS
).
New South Wales
. colour pattern. —
Carapace
. Height
2.69.
Frontal
width
Males
:
AMS
KS4104
,
Bago Forest
, nr.
Batlow
,
35°41'S
148°09'E
,
5 4.57.
Fovea
narrow, procurved.
Mid-dorsal
cephalic setae
November 1979
,
J. Kovacs
;
AMS
KS4607
,
Batlow-Tumut area
,
35°25'S
148°10'E
,
5 March 1980
,
N. Robinson
;
AMS
KS4718
,
Tumbarumba
, few, reach fovea.
Anterior
strial setae present.
Anterolateral
35°45'S
148°00'E
,
25
Mar
AMS
KS1043
,
Batlow
,
35°30'S
148°08'E
.
1980; carapace angle with small, weak bristles. —
Eyes
. Central
AMS
KS12361
,
AMS
KS1050
,
Wondalga
,
35°23'S
148°07'E
,
14
October
eye region weakly raised.
Eye
group width 1.66.
Median
1948,
C.H. Jagoe
;
Females
:
AMS
KS16451
,
Batlow
,
35°30'S
148°08'E
,
ocular quadrangle length 0.57, anterior width 0.69, posterior
16 June 1951
, H.
V
.
Smith
;
AMS
KS5054
,
Bago Forest
,
35°41'S
148°09'E
,
width 1.05. Diameters: AME 0.20, ALE 0.36, PLE 0.35,
8 January 1979
,
J. Kovacs
;
AMS
KS1047
,
Tumut
,
35°18'S
148°13'E
,
4 May 1977
,
B. Buckley.Australian Capital Territory.
Males
:
Uriarra Forest
,
PME
0.19. —
Chelicerae
. Cheliceral
groove narrow, margins
Brindabella Range
,
35°18'S
148°56'E
,
9 July 1983
, M.
R
.
Gray
;
AMS
subparallel, diverging slightly distally.
Groove
length 2.48,
KS13396
,
Tidbinbilla Nature Reserve
,
35°28'S
148°54'E
,
26 October 1983
, middle width 0.26.
Cheliceral
teeth: 16 central, confined
W. Osborne.
Females
:
AMS
KS3542
,
Tidbinbilla Nature Reserve
,
35°28'S
to basal half of groove; 11 prolateral; 12 retrolateral. —
148°54'E
,
November 1978
,
P.Ormay
;
AMS
KS928
,
Gibraltar
Falls
,
35°29'S
148°56'E
,
24 October 1971
,
T
.
McGregor.
Labium
. Much
wider than long, apically weakly indented
.
Length 1.49, width 1.08. Labiosternal sigilla broad, entire.
Other material examined.
New South Wales
(males)
.
AMS
KS13534
,
Cuspules
c. 193, number moderate, in crescentic transverse
Bago Forest
, near
Batlow
,
5 November 1979
,
J. Kovacs
;
AMS
KS1041
, band; frequency moderate. —
Sternum
. Ovoid. Length 4.43, Batlow,
December 1948
,
C.H. Jagoe
;
AMS
KS1042
,
Dora Dora National
width 3.62.
Posterior
sternal sigilla ovoid, broad.
Strong Park
,
Jingellic
near
Albury
,
16 January 1979
,
A.B. Rose
(
AMS
)
.
New South Wales
(females)
:
AMS
KS1041
,
Batlow
,
December 1948
, C.B. bristles on posterior margin. —
Palp
. Tegular area
small, Jagoe;
AMS
KS4608
,
Batlow-Tumut area
,
5 March 1980
,
N. Robinson
; slightly wider than long. Embolus long, slender; shaft tapered
AMS
KS7566
, Cooma,
19 January 1980
,
R
. Mascord;
AMS
KS13545
, and strongly curved.
Distal
embolus very slender, apical
Gundagai
,
25 October 1983
.
Australian Capital Territory
(males)
.
AMS
region weakly twisted and set at angle to the embolic axis;
KS13397
,
Tidbinbilla Nature Reserve
,
26 October 1983
,
W. Osborne
;
AMS
ejaculatory groove narrow.
Embolus
base strongly offset
KS13592
,
Canberra area
,
35°17'S
149°13'E
,
13 Dec 1983
,
T
.
Mitchell
.
Australian Capital Territory
(females)
:
AMS
KS654
,
Honeysuckle
from tegulum.
Bulb
length 4.16, width 1.10.
Embolus
length
Creek
,
24 February 1977
,
M. Noble
;
AMS
KS12032
, Gudgenby National 3.23, midwidth 0.18. Length of femur 3.48, patella 1.57, Park;
AMS
KS929
, near
Piccadilly Circus
,
25 January 1972
,
K. Watson.
tibia 4.61.
Width
of tibia 1.71.
Spination
: femur 3, patella
0, tibia 3 (all bristle-like). Distal femoral bristles and spines sinuous. —
Legs
. 4123. Leg Femur Patella Tibia Metatarsus Tarsus Total entire or narrowed centrally. —
Sternum
. Ovoid, moderately 1 6.40 3.40 4.98 4.73 2.69 22.20 wide. Length 4.59, width 3.84. Posterior sigilla moderately 2 5.98 3.26 4.69 4.61 2.61 21.15 short, ovoid: length 0.75, width 0.37. —
Palp
. Spination: tibia 3 5.30 2.79 4.01 4.59 3.06 19.75 4, tarsus 14. Trichobothria: tibia p6 r7, tarsus 12. Tarsal claw 4 6.19 2.92 5.00 5.44 3.40 22.95 with 7 teeth. —
Legs
. 4123. Legs I, IV subequal.
Figure 9.
Atrax yorkmainorum
, female: (
A
) sternum, labium and Figure 8.
Atrax yorkmainorum
, male: (
A
) sternum, labium and maxilla; (
B
) cephalothorax and chelicerae, lateral; (
C
) cheliceral maxilla; (
B
) cephalothorax and chelicerae, lateral; (
C
) spinnerets; groove teeth; (
D
) spermathecae; (
E
) spinnerets; (
F
) cephalothorax (
D
) cephalothorax and chelicerae, dorsal; (
E
) cheliceral groove and chelicerae, dorsal; (
G
) leg III, prolateral, patella, tibia, teeth; (
F
) palp, prolateral; (
G
) leg II, prolateral. Scale lines 1 mm. metatarsus. Scale lines 1 mm.
Figure 10.
Atrax yorkmainorum
, male (
A,D–K
, AMS KS13592;
B,C
, AMS KS1034): (
A
) palp, prolateral; (
B
) bulb; (
C
) embolus, distal; (
D,E
) distal patella and tibia I—
D
, ventral,
E
, prolateral; (
F,G
) metatarsus I—
F
, ventral,
G
, prolateral; (
H,I
) distal patella and tibia II—
H
, ventral,
I
, prolateral; (
J,K
) metatarsus II—
J
, ventral,
K
, prolateral. Scale lines: 0.3 mm, except
A,F,K
0.2 mm and
B
1.0 mm.
Table 3
. Male morphological data—
Atrax yorkmainorum
(n = 10).
character |
range |
mean |
character |
range |
mean |
character |
ratio |
SD |
CL |
7.83–9.22 |
8.47 |
Mt1S |
12–26 (p0–1) 19 |
CW/CL |
0.92 |
0.027 |
CW |
7.14–8.60 |
7.76 |
Ta1S |
7–13 |
10 |
CH/CW |
0.35 |
0.021 |
CH |
2.53–3.03 |
2.72 |
Ti2S |
13–20 (p1–3) 16 |
CFW/CL |
0.56 |
0.042 |
CFW |
4.14–5.20 |
4.69 |
STC2teeth |
12–15 |
13 |
CHGW/L |
0.11 |
0.016 |
ChGL |
2.33–2.70 |
2.59 |
BulbL |
3.93–4.78 |
4.36 |
SW/SL |
0.77 |
0.032 |
ChGW |
0.22–0.35 |
0.28 |
EmbL |
2.92–3.71 |
3.37 |
LL/LW |
0.72 |
0.023 |
ChGCT |
9–18 |
14 |
BulbW |
1.08–1.22 |
1.12 |
PLSAPW/L |
0.28 |
0.023 |
LL |
1.03–1.30 |
1.16 |
EmbmidW |
0.18–0.25 |
0.20 |
BulbW/EmbL |
0.34 |
0.019 |
LW |
1.41–1.77 |
1.58 |
PalpTibL |
4.32–5.10 |
4.72 |
EmbmidW/L |
0.06 |
0.008 |
CUSP |
167–228 |
199 |
PalpTibW |
1.56–1.87 |
1.69 |
PalpTibW/L |
0.36 |
0.022 |
SL |
4.19–5.06 |
4.56 |
PalpTibS |
1–4 |
3 |
BulbL/TibL |
0.92 |
0.047 |
SW |
3.28–3.83 |
3.52 |
PalpPatS |
0–2 |
<1 |
PLSAPW |
0.36–0.51 |
0.42 |
PalpFemS |
3–4 |
3 |
PLSAPL |
1.15–1.84 |
1.51 |
Fe1S |
5–13 |
8 |
Pa1S |
5–12 (p2–4) |
8 |
Leg Femur Patella Tibia Metatarsus Tarsus Total
Tibia I
width 1.35. Femora I and II with dorsal spines.
1 5.98 3.57 4.56 3.71 2.24 20.06
Leg I unmodified, tibial spines numerous. Tibia II with
2 5.30 3.37 3.88 3.47 2.24 18.26
large, conical apophysis placed centrally. Metatarsus II
3 4.56 2.82 3.15 3.30 2.38 16.21
sinuous, curved proximally with a weak, spined apophysis
4 5.59 3.18 4.28 4.38 2.75 20.18
centrally; ventral spines lacking proximally. Anterior coxal hairs unmodified. Tarsal scopulae legs I–IV: weak. Distal
Tibia I
width 1.51. Metatarsus I proximal width 1.02. Coxal
metatarsal scopulae legs I–IV: absent to weak. Tarsal claw
setae unmodified. Tarsal claw teeth legs I, II: superior 9, 9;
teeth leg I, II: superior 14, 13; inferior 0, 2. Trichobothria legs
inferior 2, 2. Trichobothria legs I, II: tarsus 13, 15; metatarsus
I, II: tarsus 15, 15; metatarsus 14, 14; tibia p8 r8, p8 r8. —
Leg
12, 11; tibia p9 r8, p8 r7. —
Leg spination
. Leg I: femur 0,
spination
. Leg I: femur 7 (d3 p4) (some bristle-like), patella
patella 0, tibia 2, metatarsus 11, tarsus 10. Leg II: femur
5 (p2), tibia 36 (p2), metatarsus 19, tarsus 10. Leg II: femur
0, patella 0, tibia 2 (p1), metatarsus 12, tarsus 8. Leg III:
10 (d4 p6) (plus several bristle-like), patella 5 (p2), tibia 16
femur 0, patella pd3, tibia 6 (p2 r1), metatarsus 20 (p6 rd2).
(p2), metatarsus 16 (p1), tarsus 10. Leg III: femur 0 (long
Leg IV: femur 0, patella 0, tibia 4, metatarsus 14, tarsus
bristles), patella 5 (p4 r1), tibia 14 (p2 r3), metatarsus 23,
12. —
Abdomen
. Posterior lateral spinnerets, lengths: total
tarsus 12. Leg IV: femur 0 (long bristles), patella r1, tibia 13
4.20; basal segment 1.73, middle 1.08, apical 1.39. Apical
(p1 r3), metatarsus 20, tarsus 17. Abdomen. Posterior lateral
segment width 0.45. —
Genitalia
. Spermathecae elongate,
spinnerets, lengths: total 4.41; basal segment 1.63; middle
strongly curved, approximated basally and apically but
1.18; apical 1.60. Apical segment width 0.41.
well separated in middle. Apical region slightly wider than remainder of spermatheca. Length 1.84, width 0.35.
Female
(
paratype
AMS KS16451). —
Size
. Carapace length
8.57, width 7.14. Abdomen length 10.27, width 7.89. —
Distribution
. Southeastern
New South Wales
and Australian
Colour
. Basic colour pattern. —
Carapace
. Much longer than
Capital
Territory (Fig. 7).
wide. Height 3.16; frontal width 5.64. Cephalic length 6.05.
Fovea narrow, procurved. Mid-dorsal cephalic setae few,
alternating large and small, reach back to fovea. Anterior
Atrax sutherlandi
n.sp.
strial setae numerous. Anterolateral carapace angle with weak bristles. —
Eyes
. Eye region slightly raised centrally. Figures 11–14;
Tables 4
,
34
Eye group width 1.65. Diameters:AME 0.20, ALE 0.45, PLE
Etymology
. The species is named for the late Professor
0.35, PME 0.26. Interdistances: AME–AME 0.23, AME–
Struan Sutherland, whose venom research resulted in the
ALE 0.15, ALE–PLE 0.14, PLE–PME 0.07, PME–PME
production of a successful funnel-web spider antivenom.
O.59. Median ocular quadrangle length 0.59, anterior width
0.63, posterior width 1.05. —
Chelicerae
. Cheliceral groove
Types
.
Holotype
male:
AMS
KS1076
,
Bermagui
,
New South Wales
,
36°25'S
narrow, margins subparallel, weakly divergent distally.
150°04'E
,
K Walker.
Paratypes
.
New South Wales
(
AMS
).
Males
:
AMS
Groove
length 2.63, middle width 0.35.
Cheliceral
teeth: 16
KS7464
,
Bega
,
36°40'S
149°54'E
,
K.Walker
;
AMS
KS1595
,
Nadgee Nature
central in basal half of groove; 11 prolateral, 12 retrolateral.
Reserve
;
AMS
KS1074
,
Bermagui
,
36°25'S
150°04'E
,
K. Walker
;
AMS
KS11748
,
Bondi State Forest
, nr.
Bombala
,
37°06'S
149°15'E
, 14
November
—
Labium
. Wider than long, anterior margin indented.
1980,
G. Gowing
et al
.;
AMS
KS1075
, Bermagui,
36°25'S
150°04'E
, K.
Length 1.36, width 1.78. Cuspules spread in shallow v-shape Walker.
Females
:
AMS
KS5127
, Bermagui,
36°25'S
150°04'E
, 18
May
across anterior two-thirds of labium.
Labiosternal
sigilla 1973,
M. Gray
;
AMS
KS4886
,
Bockelo Creek
,
18 May 1972
,
M. Gray
;
AMS
KS5119
,
Bega
,
March 1974
,
Ambulance Station
;
AMS
KS5126
,
Bega
,
Autumn
1973
;
AMS
KS1086
,
Bemboka
,
July 1977
, G. Wishart;
AMS
KS1074
,
Bermagui, K
.
Walker
;
AMS
KS3619
,
Bombala
,
17 January 1930
, A.J. Barrett;
AMS
KS11445
,
Bondi State Forest
, near
Bombala
,
31 March 1981
,
G. Gowing
;
Brown Mountain
,
30 March 1967
,
R
.
J.B.
(
ANIC
)
;
AMS
KS5042
,
Candelo
,
17 May 1973
, M. Gray;
AMS
KS7299
,
Cooma
,
6 April 1981
, A.N. Wallace;
AMS
KS1102
,
Merimbula
,
5 June 1964
;
AMS
KS1595
,
Nadgee Nature Reserve
;
AMS
KS1088
,
Narooma
,
29 January 1959
,
R
Martin
;
AMS
KS3624
,
Narooma
,
15 June 1971
, J. Cobcroft;
AMS
KS1083
,
Nerrigundah
,
1 October 1977
,
C. Cowall.
Victoria
(males)
:
AMS
KS1055
,
Murrindal
,
2 km
N. of,
3 June 1973
,
L. Windsor
;
Buchan
,
9 January 1964
,
Dingey
&
Penshurst
(
MV
)
;
AMS
KS1087
,
Cobargo
,
June 1957
,
C.G.
Bradford
;
Cobargo
,
5 November 1964
(
MV
)
;
Mallacoota
,
26 September 1963
(
MV
)
;
AMS
KS1057
,
Gelantipy
,
6 May 1947
;
AMS
KS1091
,
Mallacoota
,
May 1957
;
Mallacoota
,
January 1963
, C. McArthur;
AMS
KS3397
,
Mallacoota Inlet
,
October 1971
, S. Sutherland;
AMS
KS1055
,
Murrindal
,
3 June 1973
, L. Windsor;
AMS
KS1095
,
Noorinbee
,
17 June 1957
,
N.A.
Wakefield
;
AMS
KS1096
,
Noorinbee
,
25 September 1963
, D.S. Broome;
AMS
KS1108
,
Wingan
,
27 February 1946
.
Diagnosis
. CL 7.22–10.20 (male). Differs from other
Atrax
Figure 11. Collection records for
Atrax sutherlandi
.
spp. by having a relatively shorter and wider embolus, and a shorter palpal tibia (PalpTibW/ L 0.43) (
Fig. 14A–C
;
AMS
KS11848
,
Bondi State Forest
, nr.
Bombala
,
37°06'S
149°15'E
, 14
Table 4
).
November 1980
,
G. Gowing
;
AMS
KS3618
,
Merimbula
,
36°53'S
149°54'E
,
6 September 1976
,
H. Harvey.
Victoria
.
Males
.
K11078
(
MV
),
Noorinbee
,
37°31'S
149°10'E
,
Vic.
,
12 November 1969
,
A. Neboiss
;
K11088
(
MV
),
Male
(
holotype
). —
Size
. Carapace length 7.56, width 6.59.
Cann River
,
37°34'S
149°09'E
,
5 March 1964
.
Females
(
AMS
)
:
AMS
Abdomen
length 6.93, width 5.76. —
Colour
. Basic
colour
KS3397
,
Mallacoota Inlet
,
37°33'S
149°46'E
,
9 October 1971
,
S. Sutherland
; pattern. —
Carapace
. Longer
than wide.
Height
2.55.
Frontal
AMS
KS1092
,
Genoa
,
37°28'S
149°35'E
,
1 November 1964
. width 4.12.
Fovea
narrow, procurved.
Mid-dorsal
cephalic setae reach fovea, anterior strial setae present.
Anterolateral
Other material examined.
New South Wales
(males)
:
AMS
KS8467
, carapace angle with small, very weak bristles. —
Eyes
. Bombala
, 1981
;
AMS
KS5118
,
Bega
,
17 May 1973
,
Ambulance Station
;
Merimbula
,
March 1965
(
MV
)
;
AMS
KS9561
,
Monga
;
AMS
KS5120
,
Central
eye region weakly raised.
Eye
group width 1.58.
Mount Doctor George
, near
Bega
,
17 May 1974
;
AMS
KS6072
,
Mumbulla Median
ocular quadrangle length 0.63, anterior width 0.71,
State Forest
, near
Bega
,
November 1980
,
R
.
Wells
;
AMS
KS1085
, posterior width 1.03.
Diameters
: AME 0.19, ALE 0.44, PLE
Bemboka
,
10 May 1970
, M. Clune;
AMS
KS14232
,
Bermagui
, 8
April
0.32
,
PME
0.25. —
Chelicerae
. Cheliceral
groove narrow, 1984
;
AMS
KS11528
,
Bondi State Forest
, near
Bombala
,
14 November 1980
, G. Gowing
et al
.;
AMS
KS1084
,
Brogo
,
4 December 1962
, K. margins subparallel except distally where they diverge. Walker;
AMS
KS5497
,
Kioloa State Forest
,
4 October 1979
–30 October Groove length 2.27, middle width 0.24. Cheliceral teeth: 1979, C. Horseman;
AMS
KS5121
,
Beckelo Creek
,
18 May 1973
, M. Gray; 23 central, confined to basal half of groove; 13 prolateral;
Figure 13.
Atrax sutherlandi
, female: (
A
) sternum, labium and Figure 12.
Atrax sutherlandi
, male: (
A
) cephalothorax and maxilla; (
B
) cephalothorax and chelicerae, lateral; (
C
) leg III, chelicerae, lateral; (
B
) leg II, prolateral; (
C
) cheliceral groove teeth; prolateral, patella, tibia, metatarsus; (
D
) cephalothorax and (
D
) cephalothorax and chelicerae, dorsal; (
E
) sternum, labium and chelicerae, dorsal; (
E
) spinnerets; (
F
) spermathecae; (
G
) cheliceral maxilla; (
F
) spinnerets; (
G
) palp, prolateral. Scale lines 1 mm. groove teeth. Scale lines 1 mm.
Table 4
. Male morphological data—
Atrax sutherlandi
(n = 17).
character |
range |
mean |
character |
range |
mean |
character |
ratio |
SD |
CL |
7.22–9.64 |
8.23 |
Mt1S |
18–35 |
23 |
CW/CL |
0.87 |
0.039 |
CW |
6.49–8.45 |
7.21 |
Ta1S |
9–24 |
13 |
CH/CW |
0.36 |
0.025 |
CH |
2.13–3.36 |
2.62 |
Ti2S |
10–23 (1–4) |
15 |
CFW/CL |
0.62 |
0.039 |
CFW |
3.81–5.57 |
4.45 |
STC2teeth |
12–14 |
13 |
CHGW/L |
0.12 |
0.018 |
ChGL |
2.15–3.16 |
2.41 |
BulbL |
2.79–4.40 |
3.26 |
SW/SL |
0.70 |
0.042 |
ChGW |
0.19–0.34 |
0.28 |
EmbL |
2.02–3.01 |
2.34 |
LL/LW |
0.70 |
0.056 |
ChGCT |
10–32 |
21 |
BulbW |
0.90–1.21 |
1.02 |
PLSAPW/L |
0.25 |
0.023 |
LL |
0.92–1.39 |
1.07 |
EmbmidW |
0.15–0.22 |
0.19 |
BulbW/EmbL* 0.44 |
0.043 |
LW |
1.36–1.77 |
1.53 |
PalpTibL |
3.24–4.56 |
3.65 |
EmbmidW/L* |
0.08 |
0.012 |
CUSP |
142–295 |
209 |
PalpTibW |
1.34–1.85 |
1.56 |
PalpTibW/L* |
0.43 |
0.035 |
SL |
4.25–5.47 |
4.83 |
PalpTibS |
1–3 |
3 |
BulbL/TibL |
0.89 |
0.077 |
SW |
2.82–3.85 |
3.41 |
PalpPatS |
0–1 |
<1 |
PLSAPW |
0.38–0.66 |
0.44 |
PalpFemS |
0–4 |
2 |
PLSAPL |
1.26–2.07 |
1.53 |
Fe1S |
2–10 |
6 |
Pa1S |
5–16 (p0–3) |
10 |
13 retrolateral. —
Labium
. Much wider than long, apically wide. Height 3.74; frontal width 6.39. Cephalic length 7.11. weakly indented. Length 0.99, width 1.49. Labiosternal Fovea narrow, procurved. Mid-dorsal cephalic setal row with sigilla broad, entire. Cuspules c. 218, number moderate, alternating large and small bristles which extend back to the grouped in broad, central band. —
Sternum
. Ovoid, long. fovea.Anterior strial setae numerous. Anterolateral carapace Length 4.73, width 3.28. Posterior sigilla ovoid. Bristles on angle with weak bristles. —
Eyes
. Eye region weakly raised. posterior margin strong. —
Palp
. Tegular area wider than Eye group width 1.99. Diameters:AME 0.21, ALE 0.41, PLE long. Embolus of moderate length, shaft gently curved and 0.45, PME 0.27. Interdistances: AME–AME 0.27, AME– strongly tapered. Distal embolus slender, weakly twisted, ALE 0.20, ALE–PLE 0.20, PLE–P.M.E 0.12, PME–PME ejaculatory groove wide. Embolus base moderately to 0.81. Median ocular quadrangle length 0.78, anterior width strongly offset from tegulum. Bulb length 2.82, width 0.96. 0.78, posterior width 1.34. —
Chelicerae
. Cheliceral groove Embolus length 2.10, midwidth 0.19. Length of femur 3.32, narrow, margins subparallel, divergent distally. Groove patella 1.47, tibia 3.26. Width of tibia 1.48. Spination: femur length 3.26, middle width 0.47. Cheliceral teeth: 25 central, 3, patella 0, tibia 2. Distal femur with sinuous spines and occupying basal half of groove; 14 prolateral; 14 retrolateral. strong bristles. —
Legs
. 4123. —
Labium
. Much wider than long, anterior margin indented. Length 1.60, width 2.28. Cuspules occupy anterior two-thirds
Leg Femur Patella Tibia Metatarsus Tarsus Total of labium. Labiosternal sigilla entire. —
Sternum
. Long,
1 5.63 3.16 4.23 4.08 2.41 19.51 ovoid. Length 6.12, width 4.01. Posterior sigilla short, ovoid:
2 5.10 2.98 4.10 4.00 2.43 18.60 length 0.48, width 0.27. —
Palp
. Spination: tibia 5, tarsus
3 4.75 2.61 3.41 3.92 2.81 17.50 11. Trichobothria: tibia p8 r8, tarsus 18. Tarsal claws with
4 5.28 2.79 4.61 4.80 3.04 20.52 8 teeth. —
Legs
. 4123.
Tibia I
width 1.26. Femora I and II with dorsal spines. Leg Leg Femur Patella Tibia Metatarsus Tarsus Total
I unmodified, patella and tibial ventral spines numerous. 1 7.48 4.15 5.64 4.84 2.72 24.83 Tibia II with large, conical apophysis placed just proximal 2 6.58 3.88 4.73 4.32 2.65 22.11 of centre. Metatarsus II sinuous, strongly curved proximally, 3 5.81 3.16 3.50 4.35 2.92 19.74 with a small, spined, central apophysis. Anterior coxal hairs 4 7.11 3.84 5.03 5.54 4.01 25.53 unmodified. Tarsal scopulae legs I–IV: weak to moderate.
Distal metatarsal scopulae legs I–IV: absent to weak.
Tibia I
width 1.77. Metatarsus I proximal width 1.13. Tarsal claw teeth legs I, II: superior 13, 13; inferior 3, 2. Tarsal claw teeth legs I, II; superior 10, 11; inferior 3, 2. Trichobothria legs I, II: tarsus 14, 13, metatarsus 14, 14; Trichobothria legs I, II: tarsus 20, 16; metatarsus 20, 13; tibia p9 r9, p9 r8. —
Leg spination
. Leg I: femur d6, patella tibia p10 r9, p9 r9. —
Leg spination
. Leg I: femur 0, patella 11 (p2), tibia 39 (p2), metatarsus 24, tarsus 12. Leg II: femur 0, tibia 2, metatarsus 12, tarsus 15. Leg II: femur p1, patella d7, patella 5 (p2), tibia 16 (p3), metatarsus 16 (p1), tarsus 0, tibia 4 (p2), metatarsus 11, tarsus 12. Leg III: femur 0, 13. Leg III: femur 0 (long bristles), patella 6 (p5 r1), tibia patella pd3 r1, tibia 7 (p2 r2), metatarsus 17 (p6 rd2), tarsus 11 (p4 r3), metatarsus 15, tarsus 11. Leg IV: femur 0 (long 17. Leg IV: femur 0, patella 0, tibia 7 (r2), metatarsus 21 bristles), patella r1, tibia 8 (p1 r2), metatarsus 21, tarsus (p4 rd2), tarsus 18 —
Abdomen
. Posterior lateral spinnerets, 17. —
Abdomen
. Posterior lateral spinnerets, lengths: total lengths: total 5.55; basal segment 2.04, middle 1.16, apical 3.82, basal segment 1.36; middle 0.98; apical 1.48. Apical 2.35.Apical segment width 0.54. —
Genitalia
. Spermathecae segment width 0.41. moderately elongate, gently curved, basally approximated. Length 1.66, width 0.52.
Female
(
paratype
AMS KS3397). —
Size
. Carapace length
10.20, width 8.16. Abdomen length 10.88, width 9.18. —
Distribution
. Southeastern
New South Wales
and north-
Colour
. Basic colour pattern. —
Carapace
. Much longer than eastern
Victoria
(Fig. 11).