<document id="195C215D40AF7956030F49871AF59672" ID-CLB-Dataset="308815" ID-DOI="10.1111/zsc.12663" ID-GBIF-Dataset="1f91cf0c-277a-4239-827e-dd7f452ae64c" ID-Zenodo-Dep="15089748" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="carolina" IM.metadata_approvedBy="valdenar" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="carolina" checkinTime="1742993903521" checkinUser="valdenar" docAuthor="Riccardi, Paula Raile &amp; Hartop, Emily" docDate="2024" docId="183987D1FF8EF0775828C1F2FA2EF96E" docLanguage="en" docName="ZoolScr.53.614-631.pdf.imf" docOrigin="Zoologica Scripta 53 (5)" docSource="https://doi.org/10.1111/zsc.12663" docStyle="DocumentStyle:C474B9ED07F03274EF3A059EADEEC434.1:ZoolScr.2017-.journal_article.closed" docStyleId="C474B9ED07F03274EF3A059EADEEC434" docStyleName="ZoolScr.2017-.journal_article.closed" docStyleVersion="1" docTitle="Aphanotrigonum Duda 1932" docType="treatment" docVersion="6" lastPageNumber="621" masterDocId="E400FFA9FF89F0705852C736FF8CFF8E" masterDocTitle="Large-scale integrative taxonomy of Swedish grass flies (Diptera, Chloropidae) reveals hitherto unknown complexity of a dark taxon" masterLastPageNumber="631" masterPageNumber="614" pageNumber="621" updateTime="1770418576608" updateUser="carolina">
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<mods:title id="E90B2918010CFE15A91FE0F734C28B52">Large-scale integrative taxonomy of Swedish grass flies (Diptera, Chloropidae) reveals hitherto unknown complexity of a dark taxon</mods:title>
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<subSubSection id="D88A654CFF8EF0775828C1F2FD6CF96F" box="[122,736,1728,1766]" pageId="7" pageNumber="621" type="nomenclature">
<paragraph id="902F36C7FF8EF0775828C1F2FD6CF96F" blockId="7.[122,736,1728,1766]" box="[122,736,1728,1766]" pageId="7" pageNumber="621">
4.2.2 | Genus 
<taxonomicName id="57904D44FF8EF0775908C1F2FD6CF96F" ID-CoL="YBF" authority="Duda, 1932" authorityName="Duda" authorityYear="1932" box="[346,736,1732,1761]" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="genus">
<emphasis id="A2E4EAD5FF8EF0775908C1F2FDB3F96F" box="[346,575,1732,1761]" italics="true" pageId="7" pageNumber="621">Aphanotrigonum</emphasis>
Duda, 1932
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D88A654CFF8EF0775828C03BFA2EF96E" pageId="7" pageNumber="621" type="discussion">
<paragraph id="902F36C7FF8EF0775828C03BFA22FBA4" blockId="7.[122,770,1805,1968]" lastBlockId="7.[818,1466,1008,1760]" pageId="7" pageNumber="621">
The five species of 
<taxonomicName id="57904D44FF8EF077593DC03BFDA0F8AB" authorityName="Duda" authorityYear="1932" box="[367,556,1805,1829]" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="genus">
<emphasis id="A2E4EAD5FF8EF077593DC03BFDA0F8AB" box="[367,556,1805,1829]" italics="true" pageId="7" pageNumber="621">Aphanotrigonum</emphasis>
</taxonomicName>
identified in our dataset include one species complex and 
<taxonomicName id="57904D44FF8EF0775ADDC006FE2DF8E4" authority="Riccardi (Riccardi, 2024)" authorityName="Riccardi (Riccardi" authorityYear="2024" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="species" species="palpalis">
<emphasis id="A2E4EAD5FF8EF0775ADDC006FC8DF8C6" box="[655,769,1840,1864]" italics="true" pageId="7" pageNumber="621">A.palpalis</emphasis>
Riccardi (
<bibRefCitation id="F4014B36FF8EF07758BFC064FE1AF8E4" author="Riccardi" box="[237,406,1874,1898]" firstAuthor="Riccardi" pageId="7" pageNumber="621" pagination="29 - 33" refId="ref12233" refString="Riccardi, P. R. (2024). Two new species of grass flies (Diptera, Chloropidae) from Sweden. Journal of Insect Biodiversity, 48 (1), 29-33. https://doi.org/10.12976/jib/2024.48.1.4" type="journal article" year="2024">Riccardi, 2024</bibRefCitation>
)
</taxonomicName>
. According to 
<bibRefCitation id="F4014B36FF8EF0775A0CC064FEDDF803" author="Nartshuk and Andersson's" firstAuthor="Nartshuk" pageId="7" pageNumber="621" pagination="1 - 234" refId="ref11975" refString="Nartshuk, E. P., &amp; Andersson, H. (2013). The frit flies (Chloropidae, Diptera) of Fennoscandia and Denmark. In The frit flies (Chloropidae, Diptera) of Fennoscandia and Denmark (pp. 1-234). Brill." type="book chapter" year="2013">Nartshuk and Andersson's (2013)</bibRefCitation>
definition, three species in our dataset were identified as 
<taxonomicName id="57904D44FF8EF0775922C0AEFD8BF83E" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[368,519,1944,1968]" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="species" species="trilineatum">
<emphasis id="A2E4EAD5FF8EF0775922C0AEFD8BF83E" box="[368,519,1944,1968]" italics="true" pageId="7" pageNumber="621">A.trilineatum</emphasis>
</taxonomicName>
, leading us to assume that the current definition of 
<taxonomicName id="57904D44FF8EF0775C2BC4C6FA9CFB86" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[1145,1296,1008,1032]" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="species" species="trilineatum">
<emphasis id="A2E4EAD5FF8EF0775C2BC4C6FA9CFB86" box="[1145,1296,1008,1032]" italics="true" pageId="7" pageNumber="621">A.trilineatum</emphasis>
</taxonomicName>
corresponds to a species complex that has been previously unrecognised.
</paragraph>
<paragraph id="902F36C7FF8EF0775B07C303FAD7F981" blockId="7.[818,1466,1008,1760]" pageId="7" pageNumber="621">
The species 
<taxonomicName id="57904D44FF8EF0775C5FC303FC24FBFE" authority="Dely-Draskovits" authorityName="Dely-Draskovits" authorityYear="1981" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="species" species="beschovskii">
<emphasis id="A2E4EAD5FF8EF0775C5FC303FAD3FBC3" box="[1037,1375,1077,1101]" italics="true" pageId="7" pageNumber="621">Aphanotrigonum beschovskii</emphasis>
Dely-Draskovits
</taxonomicName>
is currently considered a junior synonym of 
<taxonomicName id="57904D44FF8EF0775B60C34DFBBAFB1C" authority="(Meigen)" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[818,1078,1146,1170]" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="species" species="trilineatum">
<emphasis id="A2E4EAD5FF8EF0775B60C34DFC46FB1C" box="[818,970,1146,1170]" italics="true" pageId="7" pageNumber="621">A. trilineatum</emphasis>
(Meigen)
</taxonomicName>
(
<bibRefCitation id="F4014B36FF8EF0775C17C34CFACEFB1C" author="Bystrowski, C. &amp; Richter, V. A. &amp; Beuk, P. &amp; Pape, T. &amp; Nartshuk, E. P." box="[1093,1346,1146,1170]" etAl="et al." firstAuthor="Bystrowski" journalOrPublisher="Fauna Europaea" pageId="7" pageNumber="621" publicationUrl="https://fauna-eu.org" refId="ref10704" refString="Bystrowski, C., Richter, V. A., Beuk, P., Pape, T., &amp; Nartshuk, E. P. (2023). Fauna Europaea: Chloropidae. Fauna Europaea, https://fauna-eu.org" title="Fauna Europaea: Chloropidae" type="url" year="2023">Bystrowski et al., 2023</bibRefCitation>
; 
<bibRefCitation id="F4014B36FF8EF0775D02C34CFBAAFB3B" author="Nartshuk &amp; Andersson" firstAuthor="Nartshuk" pageId="7" pageNumber="621" pagination="1 - 234" refId="ref11975" refString="Nartshuk, E. P., &amp; Andersson, H. (2013). The frit flies (Chloropidae, Diptera) of Fennoscandia and Denmark. In The frit flies (Chloropidae, Diptera) of Fennoscandia and Denmark (pp. 1-234). Brill." type="book chapter" year="2013">Nartshuk &amp; Andersson, 2013</bibRefCitation>
). 
<bibRefCitation id="F4014B36FF8EF0775C1FC3ABFAD0FB3B" author="Dely-Draskovits" box="[1101,1372,1181,1205]" firstAuthor="Dely-Draskovits" pageId="7" pageNumber="621" pagination="115 - 138" refId="ref10827" refString="Dely-Draskovits, A. (1981). Revision der palaearktischen Arten der Gattung Aphanotrigonum Duda, 1932, und Aphanotrigonella Nartshuk, 1964 (Diptera: Chloropidae). Acta Zoologica Academiae Scientiarum Hungaricae, 27 (1 - 2), 115-138." type="journal article" year="1981">Dely-Draskovits (1981)</bibRefCitation>
distinguished 
<taxonomicName id="57904D44FF8EF0775BC5C3F6FBA7FB56" authorityName="Dely-Draskovits" authorityYear="1981" box="[919,1067,1216,1240]" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="species" species="beschovskii">
<emphasis id="A2E4EAD5FF8EF0775BC5C3F6FBA7FB56" box="[919,1067,1216,1240]" italics="true" pageId="7" pageNumber="621">A. beschovskii</emphasis>
</taxonomicName>
by the base of surstylus bearing short setae, unlike 
<taxonomicName id="57904D44FF8EF0775C72C3D5FC74FA93" authority="sensu Nartshuk and Andersson (2013)" authorityName="sensu Nartshuk and Andersson" authorityYear="2013" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="species" species="trilineatum">
<emphasis id="A2E4EAD5FF8EF0775C72C3D5FB34FB74" box="[1056,1208,1250,1274]" italics="true" pageId="7" pageNumber="621">A. trilineatum</emphasis>
sensu 
<bibRefCitation id="F4014B36FF8EF0775D44C3D4FC74FA93" author="Nartshuk and Andersson" firstAuthor="Nartshuk" pageId="7" pageNumber="621" pagination="1 - 234" refId="ref11975" refString="Nartshuk, E. P., &amp; Andersson, H. (2013). The frit flies (Chloropidae, Diptera) of Fennoscandia and Denmark. In The frit flies (Chloropidae, Diptera) of Fennoscandia and Denmark (pp. 1-234). Brill." type="book chapter" year="2013">Nartshuk and Andersson (2013)</bibRefCitation>
</taxonomicName>
. The longitudinal lines in the scutum are distinctive features of 
<taxonomicName id="57904D44FF8EF0775C74C21EFB31FACE" authorityName="sensu Nartshuk and Andersson" authorityYear="2013" box="[1062,1213,1320,1344]" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="species" species="trilineatum">
<emphasis id="A2E4EAD5FF8EF0775C74C21EFB31FACE" box="[1062,1213,1320,1344]" italics="true" pageId="7" pageNumber="621">A. trilineatum</emphasis>
</taxonomicName>
according to 
<bibRefCitation id="F4014B36FF8EF0775D02C21EFBA3FAEC" author="Nartshuk and Andersson" firstAuthor="Nartshuk" pageId="7" pageNumber="621" pagination="1 - 234" refId="ref11975" refString="Nartshuk, E. P., &amp; Andersson, H. (2013). The frit flies (Chloropidae, Diptera) of Fennoscandia and Denmark. In The frit flies (Chloropidae, Diptera) of Fennoscandia and Denmark (pp. 1-234). Brill." type="book chapter" year="2013">Nartshuk and Andersson (2013)</bibRefCitation>
. However, our data show that this feature is shared by four species (Figure 5M1, N1, O1, P1). The morphology of the male terminalia and clusters at a 3% threshold support four species, three of which identified as 
<taxonomicName id="57904D44FF8EF0775B2CC2E3FB9AFA63" authorityName="sensu Nartshuk and Andersson" authorityYear="2013" box="[894,1046,1493,1517]" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="species" species="trilineatum">
<emphasis id="A2E4EAD5FF8EF0775B2CC2E3FB9AFA63" box="[894,1046,1493,1517]" italics="true" pageId="7" pageNumber="621">A. trilineatum</emphasis>
</taxonomicName>
. The diagnostic features of the species assigned to the cluster 147 are outlined in Table 2.
</paragraph>
<paragraph id="902F36C7FF8EF0775B07C12CFA2EF96E" blockId="7.[818,1466,1008,1760]" pageId="7" pageNumber="621">
Our results support the erection of 
<taxonomicName id="57904D44FF8EF0775D52C12CFC24F9DB" authorityName="Dely-Draskovits" authorityYear="1981" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="species" species="beschovskii">
<emphasis id="A2E4EAD5FF8EF0775D52C12CFC24F9DB" italics="true" pageId="7" pageNumber="621">Aphanotrigonum beschovskii</emphasis>
</taxonomicName>
as a valid species. The species identified here as 
<taxonomicName id="57904D44FF8EF0775B09C156FC63F9F6" authorityName="sensu Nartshuk and Andersson" authorityYear="2013" box="[859,1007,1632,1656]" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="species" species="trilineatum">
<emphasis id="A2E4EAD5FF8EF0775B09C156FC63F9F6" box="[859,1007,1632,1656]" italics="true" pageId="7" pageNumber="621">A. trilineatum</emphasis>
</taxonomicName>
var. 2 corresponds to 
<bibRefCitation id="F4014B36FF8EF0775D45C156FB9CF914" author="Nartshuk and Andersson's" firstAuthor="Nartshuk" pageId="7" pageNumber="621" pagination="1 - 234" refId="ref11975" refString="Nartshuk, E. P., &amp; Andersson, H. (2013). The frit flies (Chloropidae, Diptera) of Fennoscandia and Denmark. In The frit flies (Chloropidae, Diptera) of Fennoscandia and Denmark (pp. 1-234). Brill." type="book chapter" year="2013">Nartshuk and Andersson's (2013)</bibRefCitation>
figures 142–148 of 
<taxonomicName id="57904D44FF8EF0775D4DC1B5FA3FF914" authorityName="sensu Nartshuk and Andersson" authorityYear="2013" box="[1311,1459,1666,1690]" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="species" species="trilineatum">
<emphasis id="A2E4EAD5FF8EF0775D4DC1B5FA3FF914" box="[1311,1459,1666,1690]" italics="true" pageId="7" pageNumber="621">A.trilineatum</emphasis>
</taxonomicName>
. Therefore, 
<taxonomicName id="57904D44FF8EF0775BFBC193FBB0F933" authorityName="sensu Nartshuk and Andersson" authorityYear="2013" box="[937,1084,1701,1725]" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="species" species="trilineatum">
<emphasis id="A2E4EAD5FF8EF0775BFBC193FBB0F933" box="[937,1084,1701,1725]" italics="true" pageId="7" pageNumber="621">A.trilineatum</emphasis>
</taxonomicName>
var. 1 should be described as a new species after consultation of the 
<taxonomicName id="57904D44FF8EF0775CDDC1FEFAAFF96E" authorityName="sensu Nartshuk and Andersson" authorityYear="2013" box="[1167,1315,1736,1760]" class="Insecta" family="Chloropidae" genus="Aphanotrigonum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="621" phylum="Arthropoda" rank="species" species="trilineatum">
<emphasis id="A2E4EAD5FF8EF0775CDDC1FEFAAFF96E" box="[1167,1315,1736,1760]" italics="true" pageId="7" pageNumber="621">A.trilineatum</emphasis>
</taxonomicName>
type series.
</paragraph>
</subSubSection>
</treatment>
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