Annotated keys to the genera of African Bombylioidea (Diptera: Bombyliidae; Mythicomyiidae) Author Greathead, D. J. Author Evenhuis, N. L. text African Invertebrates 2001 2001-12-31 42 1820 105 224 journal article 10.5281/zenodo.7909975 2305-2562 7909975 Key to the genera of Bombyliini 1 Head usually distinctly narrower than thorax; laterotergites always with hair and cell r 5 (first posterior) usually closed at a distance from the wing margin ( Figs 81– 83 ); wings usually large, broad at base with anal lobe, alula and squama well developed; hind femora always with well developed bristles on underside; body always broad, bristles usually well developed; male genitalia with narrow or leaf-like gonostyli, aedeagus sheathed by epiphallus which only rarely has clasper-like outgrowths, phallosome projecting from middle of gonocoxae, apodemes usually large ( Fig. 101 ) ..................................................................................................... 2 – Head usually as broad as thorax; laterotergites bare and/or cell r 5 broadly open at wing margin (cf. Figs 84–85 ); wings often small, narrowed at base with at least alula reduced; hind femora often without bristles; body usually more elongate even narrow conical or cylindrical, bristles usually weak, often hair-like, especially those posterior to the wings; often with long robust gonostyli ending in a curved point; phallosome usually with a large blunt epiphallus and originating near base of gonocoxae, apodemes usually small ( Figs 102–103 ) ......................................... 10 2 Head as wide as thorax; hind margin of eye indented; cell r 5 (first posterior) pointed and closed near the wing margin; with strong bristles on head and body, including hind margins of abdominal terga; with gold scaling on head and body with a pattern of short black and white hair and scales ( Fig. 76 ); male genitalia ( Fig. 104 ) with a crest of strong spines at apex of gonocoxae which also bear a tuft of long hairs on the dorsal surface, phallosome long and slender without an epiphallus .................. Eurycarenus Bezzi Figs 49–50. Bombyliidae heads (from Evenhuis 1990 a ). 49. Usia bicolor Macquart. a . lateral view. b . frontal view. 50. Parageron ignoratus (Becker) . a . lateral view. b . frontal view. – Head usually narrower than thorax; hind margin of eye not indented; cell r 5 closed and stalked; hair long and dense; gonocoxae without a crest of strong spines or a tuft of long hairs on the dorsal surface, phallosome with or without a prominent epiphallus .............................................................................................................. 3 3 Cell br (first basal) longer than cell bm (second basal) ( Fig. 79 ), r-m sometimes equal to m-m; vestiture various, often with black hair and/or scale patches; wing pattern various, sometimes with a clear-cut dark infuscation and/or with isolated dark spots; phallosome usually without clasper-like outgrowths ( Fig. 101 ) .............................................................................................................. 4 – Cell br equal in length to bm; r-m usually much shorter than m-m, occasionally almost as long; vestiture fine and silky with a clipped appearance at least on occiput and thorax, hair white to straw-yellow or brown shading to paler on underside; wing infuscation, if present, diffuse and darker along fore border ( Figs 80–81 ); male with phallosome compact often with clasper-like outgrowths ( Fig. 105 ) ................................................................................... Systoechus Loew 4 Abdomen appears truncated, rapidly tapered from tergum 4; terminalia appear similar in both sexes with cerci in female and processes on gonocoxae in males projecting; frons of female narrow, with eyes separated by less than width of ocellar triangle; gonocoxae with robust elongate terminal lobes, gonostyli thick, parallelsided blunt or with a small point and with short hairs only ( Fig. 106 ); female with a tubular ovipositor and vestigial sand-chamber, unlaid eggs large spherical and pigmented .......................................................................... Beckerellus Greathead – Abdomen ovate, tapered beyond tergum 4; terminalia normal; frons of female at least as broad as ocellar triangle; gonocoxae with terminal lobes no longer than wide, gonostyli slender, pointed and crowned with long hairs at base or broad with short upright hairs, female with a sand-chamber guarded by long silky hairs, unlaid eggs ellipsoidal, thin-walled, unpigmented .......................................................... 5 5 Eyes of males in contact for a distance at least equal to length of ocellar triangle; head always distinctly narrower than thorax; palpi unsegmented; legs usually very dark or black and at least bristles black; body cuticle mainly or entirely black; gonocoxae slender, gonostyli elongate slender with a tuft of long black hairs at base, phallosome usually without accessory structures ........................................ 6 – Eyes of males in contact for less than length of ocellar triangle; head broad, sometimes as wide as thorax; palpi with two segments; legs pale but bases of femora and tarsi may be darkened; bristles almost always pale at least on males; body cuticle tending to have paler reddish or yellowish areas on calli; scutellum and sides of anterior abdominal terga; gonocoxae broad, gonostyli tending to be broader and shorter and without long black hairs, phallosome often with accessory structures ( Fig.108 ) ........................................................................................ Parisus Walker