The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group
Author
Harzhauser, Mathias
0000-0002-4471-6655
mathias.harzhauser@nhm-wien.ac.at
Author
Landau, Bernard M.
0000-0002-4471-6655
mathias.harzhauser@nhm-wien.ac.at
text
Zootaxa
2021
2021-08-23
5025
1
1
75
http://dx.doi.org/10.11646/zootaxa.5025.1.1
journal article
10.11646/zootaxa.5025.1.1
1175-5326
5258835
35E6C8FA-4078-4C53-9B74-F9618D5E6E60
Orthurella moravica
(
Hoernes & Auinger, 1880
)
nov. comb.
Figs 4K
,
13D
1
–D
2
, E
1
–E
2
, F
1
–F
2
, I
[
Anachis
]
Moravica
nov. form.—Hoernes 1880: 127 [
nomen nudum
].
*
Columbella
(
Anachis
)
Moravica
nov. form.—
Hoernes & Auinger 1880: 101
, pl. 12, figs 1–2.
Columbella
(
Macrurella
)
moravica
Hö. Au.
var.—
Boettger 1906: 20
.
Mitrella
(
Macrurella
)
moravica
(
Hoernes und Auinger 1880
)
—
Kojumdgieva & Strachimirov 1960: 185
, pl. 45, fig. 11.
Columbella
(
Macrurella
)
moravica
Hoernes und Auinger 1880
—
Strausz 1966: 295
, pl. 42, figs 4–5.
Type material.
Lectotype
(designated herein):
NHMW1861
/0011/0519a, SL:
17.1 mm
, MD:
6.9 mm
,
Jaroměřice nad Rokytnou
(the
Czech Republic
), illustrated in
Hoernes & Auinger (1880
, pl. 12, figs 1a–c),
Figs 13E
1
–E
2
.
Paralectotypes
:
NHMW1861
/0011/0519b, SL:
14.9 mm
, MD:
6.8 mm
,
Jaroměřice nad Rokytnou
(the
Czech Republic
)
, illustrated in
Hoernes & Auinger (1880
, pl. 12, figs 2a–c),
Figs 13D
1
–D
2
.
NHMW2020
/0159/0001, SL:
19.7 mm
, MD:
7.4 mm
,
Jaroměřice nad Rokytnou
(the
Czech Republic
)
,
Figs 4K
,
13F
1
–F
2
.
NHMW2020
/0159/0002, SL:
16.1 mm
, MD:
7.1 mm
,
Jaroměřice nad Rokytnou
(the
Czech Republic
)
;
Fig. 13I
.
NHMW1864
/0001/0326,
12 spec.
,
Jaroměřice nad Rokytnou
(the
Czech Republic
)
;
NHMW2020
/0160/0001,
1 spec.
,
Boskovice
(the
Czech Republic
)
.
Description.
Shell medium sized, moderately broad fusiform, with strongly gradate spire and broad sutural shelf. Protoconch large, conical, multispiral of 3.5 convex whorls. Teleoconch of eight whorls. Early spire narrowly conical with moderately convex whorls and narrowly incised suture. Spire whorls becoming subcylindrical and rapidly increasing in width within 6 to 8 teleoconch whorls, resulting in broad sutural shelf bordered by crested shoulder. Last whorl subcylindrical, weakly keeled at periphery, strongly constricted, with high base. Sculpture of delicate spiral threads on entire teleoconch; numerous broad spiral cords with narrow interspaces on abapical portion of base and siphonal canal. Aperture narrow, elongate. Outer lip weakly thickened with about nine small weak denticles within. Anal canal narrowly incised, V-shaped. Siphonal canal very long, narrow, deflected.
Columella
twisted, forming weak columellar fold. Columellar callus narrow, thin, adherent, without denticles. Parietal callus slightly thickened.
Shell measurements and ratios.
SL:
14.9–19.7 mm
, MD:
6.9–7.4 mm
, AA: 35–41°, SL/MD: 2.5–2.7, AL/AW: 4.8–5.0, LWH/AH: 1.2–1.3.
Discussion.
This species is characterised by its strikingly allometric growth. The early spire morphology, however, is identical with
Orthurella convexula
, which develops a similar protoconch. In addition, both species develop an extraordinarily long siphonal canal. Therefore, we place
Columbella
(
Anachis
)
moravica
in
Orthurella
. The broad subhorizontal sutural shelf immediately distinguishes
Orthurella moravica
(
Hoernes & Auinger, 1880
)
from
O. convexula
.
Palaeoenvironment.
Unknown.
Distribution in Central Paratethys.
Badenian (middle Miocene):
Carpathian Foredeep
: Jaroměřice nad Rokytnou, Boskovice (the
Czech Republic
);
Pannonian Basin: Szob
(
Hungary
) (
Strausz 1966
);
Făget Basin
: Coşteiu de Sus (
Romania
) (
Boettger 1906
);
Dacian Basin
: Târnene (
Bulgaria
) (Kojumdgieva
in
Kojumdgieva & Strachimirov 1960
).
Genus
Clinurella
Sacco, 1890
Type
species.
Columbella
(
Clinurella
)
scalaris
Sacco 1890
(in
Sacco 1890b
); subsequent designation by
Pace (1902: 43)
. Early Miocene,
Italy
.
Discussion.
Clinurella
comprise medium sized, moderately broad fusiform shells with high to moderately high spire and convex, strongly constricted last whorl with long, narrow, slightly deflected siphonal canal. A flaring outer lip with prominent denticles and prominent spiral cords on the base and on the exterior part of the outer lip are characteristic.
Sacco (1890b: 44)
established
Clinurella
as a subgenus of
Columbella
and listed
Columbella (Clinurella) scalaris
(Sacco, 1890)
,
Columbella
(
Clinurella
)
albucianensis
Sacco, 1890
,
Columbella borsoni
Bellardi, 1848
and
Columbella
(
Clinurella
)
vialensis
Sacco,
1890
in his new subgenus.
Columbella borsoni
,
C. minima
and
C. vialensis
have relatively short siphonal canals and only moderately flaring apertures, if at all, and are placed herein in
Mitrella
(see also
Landau
et al.
2013: 189
). Therefore, we place only
Clinurella scalaris
,
C. albucianensis
and
C. sophiae
(
Boettger, 1902
)
in
Clinurella
.
Cossmann (1901: 242)
synonymised
Clinurella
with
Atilia
H. Adams & A.
Adams, 1853
(=
Anachis
H. Adams & A.
Adams, 1853
).
Anachis scalarina
(G. B. Sowerby I, 1832)
,
type
species of
Anachis
, and
Anachis suffusa
(G.B. Sowerby I, 1844)
,
type
species of
Atilia
, differ clearly from
Clinurella
in their short siphonal canal and simple outer lips and the synonymy proposed by
Cossmann (1901)
is rejected herein.
Clinurella
is reminiscent of
Orthurella
Sacco, 1890
but is distinguished by its broader shell, thickened and flaring outer lip and prominent denticulation on the columellar edge and the outer lip. In
Orthurella
columellar denticles are absent, any sculpture resulting from the spiral cords over the base showing through the columellar callus.
Clinurella
is known so far only from the Burdigalian and Langhian of the Proto-Mediterranean Sea and the Langhian (Badenian) of the Central Paratethys Sea.