The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group
Author
Harzhauser, Mathias
0000-0002-4471-6655
mathias.harzhauser@nhm-wien.ac.at
Author
Landau, Bernard M.
0000-0002-4471-6655
mathias.harzhauser@nhm-wien.ac.at
text
Zootaxa
2021
2021-08-23
5025
1
1
75
http://dx.doi.org/10.11646/zootaxa.5025.1.1
journal article
10.11646/zootaxa.5025.1.1
1175-5326
5258835
35E6C8FA-4078-4C53-9B74-F9618D5E6E60
Sulcomitrella januskiewiczi
(
Friedberg, 1938
)
nov. comb.
Figs 7B
1
–B
2
*
Columbella Januskiewiczi
Friedb.
—
Friedberg 1938: 129
, text-fig. 40.
Type material.
Lectotype
(designated herein): ZNG PAN 2528,
Zborów
(
Ukraine
), stored in the
Geological Museum
of the
Institute of Geological Sciences
,
Polish Academy of Sciences
(
Kraków
), illustrated in
Friedberg (1938
, text-fig. 40),
Figs 7B
1
–B
2
.
Description.
Small shell with high, weakly cyrtoconoid spire and narrowly incised suture, Protoconch unknown. Teleoconch of six whorls. Spire whorls rapidly increasing in height, weakly convex, with periphery slightly below mid-whorl on penultimate whorl. Last whorl high, weakly convex, slowly contracting. Columellar callus forming narrow rim terminating at siphonal canal, with indistinct denticles. Outer lip weakly thickened with prominent denticles within. Anal canal narrowly incised, V-shaped. Siphonal canal very short, wide, shallow. Sculpture consisting of weak spiral grooves, being most prominent below adapical suture and on base and fasciole.
Shell measurements and ratios.
SL: 7.0 mm, MD:
4.5 mm
, AA: 48°, SL/MD: 2.5, AL/AW: 3.2, LWH/AH: 1.4.
FIGURE 7.
Species described by
Friedberg (1911
,
1938
).
A
1
–A
2
.
“
Columbella
”
paucicostata
(
Friedberg, 1911
), ZNG PAN 1257, holotype, Dryszczów (Ukraine).
B
1
–B
2
.
Sulcomitrella januskiewiczi
(
Friedberg, 1938
)
, ZNG PAN 2528, lectotype, Zborów (Ukraine).
C
2
–C
2
.
Mitrella convexa
Friedberg, 1911
, re-illustrated from
Friedberg (1911
, pl. 2, fig. 2).
Discussion.
Two specimens are labelled as
type
specimens of
Columbella januskiewiczi
in the collection of the Geological Museum of the Institute of Geological Sciences, Polish Academy of Sciences (Kraków). These differ from the illustration in
Friedberg (1938
, text-fig. 40) in their much narrower aperture, less convex whorls and slightly smaller size but agree in the spiral sculpture. The pupoid text figure in
Friedberg (1938)
is reminiscent of
Aesopus
Gould, 1860
(
type
species
Aesopus japonicus
Gould, 1860
; present-day,
Japan
) as defined by
deMaintenon (2019: 161)
. The most characteristic feature of
Aesopus
is the descending suture of the last whorl, which was already emphasized by
Gould (1860: 383)
. This feature is also developed in the Paratethyan species to some degree. The narrow aperture and the weakly incised siphonal canal, however, are atypical for
Aesopus
.
We tentatively place
Columbella januszkiewiczi
in
Sulcomitrella
mainly because of the prominent spiral sculpture but note that the narrow aperture and high last whorl are atypical for
Sulcomitrella
. Consequently, the otherwise comparable extant
Sulcomitrella leylae
Monsecour & Monsecour, 2018
and
Sulcomitrella monodonta
(
Habe, 1958
)
differ in their higher spires and wider apertures.
Sulcomitrella sceauxensis
Landau, Ceulemans & Van Dingenen, 2019
from the Tortonian of
France
, differs in its much broader outline and prominent spiral cords on base and fasciole.
Palaeoenvironment.
Unknown.
Distribution in Central Paratethys.
Badenian (middle Miocene): Carpathian Foredeep: Zborów (
Ukraine
) (
Friedberg 1938
).
Genus
Mitrella
Risso, 1826
Type
species.
Mitrella flaminea
Risso, 1826
[=
Mitrella scripta
(
Linnaeus, 1758
)
]; subsequent designation by
Moerch 1859: 258
, Present-day, Mediterranean Sea.
Discussion.
Radwin (1978: 337)
characterised
Mitrella
as small to moderately large fusiform, with high, acute spire and flat-sided whorls, moderately broad aperture, denticulate and straight columella, denticulate outer lip, very short to moderately long siphonal canal. The genus
Mitrella
Risso, 1826
, as used here, is interpreted rather widely and is unlikely to be monophyletic. Preliminary molecular results reject the genus, as presently used, being monophyletic (deMaintenon and Strong, unpublished results; Bouchet personal communication,
August 2020
).