Order Pilosa Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1 100 103 book chapter 0-8018-8221-4 10.5281/zenodo.7316519 Bradypus variegatus subsp. brasiliensis Blainville 1840 Synonyms: Bradypus variegatus subsp. ai Wagler 1831 ; Bradypus variegatus subsp. ustus Lesson 1840 ; Bradypus variegatus subsp. pallidus Wagner 1844 .