<document ID-DOI="http://dx.doi.org/10.3897/zookeys.421.7517" ID-PMC="PMC4109472" ID-Pensoft-Pub="1313-2970-421-91" ID-Pensoft-UUID="FFA88030FFBAFF84A232FFF3FF807607" ID-PubMed="25061381" ID-Zenodo-Dep="578387" ID-ZooBank="4DB3DA2D21B14D269544B4008028D304" ModsDocID="1313-2970-421-91" checkinTime="1451245675936" checkinUser="pensoft" docAuthor="Schmidt, B. Christian &amp; Anweiler, Gary G." docDate="2014" docId="2BFF205E42115E918BAA96370CE26F06" docLanguage="en" docName="ZooKeys 421: 91-113" docOrigin="ZooKeys 421" docPubDate="2014-06-27" docSource="http://dx.doi.org/10.3897/zookeys.421.7517" docTitle="Raphia frater subsp. elbea Smith 2014, stat. n." docType="treatment" docVersion="3" id="FFA88030FFBAFF84A232FFF3FF807607" lastPageNumber="106" masterDocId="FFA88030FFBAFF84A232FFF3FF807607" masterDocTitle="Taxonomy and biogeography of the Nearctic Raphia Huebner (Lepidoptera, Noctuidae, Raphiinae)" masterLastPageNumber="113" masterPageNumber="91" pageNumber="105" updateTime="1668158807944" updateUser="ExternalLinkService">
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<mods:title id="FBBD4A1EE6A434364EDBDD848F5E8298">Taxonomy and biogeography of the Nearctic Raphia Huebner (Lepidoptera, Noctuidae, Raphiinae)</mods:title>
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<mods:namePart id="5DC455508AB4690B18FF7062032E362D">Schmidt, B. Christian</mods:namePart>
<mods:affiliation id="45663A0D79762801432644CA40964239">Canadian Food Inspection Agency, Canadian National Collection of Insects, Arachnids and Nematodes, K. W. Neatby Bldg., 960 Carling Ave., Ottawa, ON, Canada K 1 A 0 C 6</mods:affiliation>
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<mods:namePart id="4053D5C04476A57415DA8B062D42B04D">Anweiler, Gary G.</mods:namePart>
<mods:affiliation id="962CCE0FF91693C33971408FE3E568B8">E. H. Strickland Entomological Museum, 218 Earth Sciences Building, Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada, T 6 G 2 E 9</mods:affiliation>
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<mods:date id="CFE70FCA9348DF3179FF4CDB2B9083C8">2014</mods:date>
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<subSection id="BBD1F15E9F001DAFBF7BE2C8CC290D9D" lastPageId="15" lastPageNumber="106" pageId="14" pageNumber="105" type="systematics">
<treatment id="2BFF205E42115E918BAA96370CE26F06" ID-GBIF-Taxon="182234978" LSID="urn:lsid:plazi:treatment:2BFF205E42115E918BAA96370CE26F06" httpUri="http://treatment.plazi.org/id/2BFF205E42115E918BAA96370CE26F06" lastPageId="15" lastPageNumber="106" pageId="14" pageNumber="105">
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<paragraph id="1D93F718E64080516DE0735B50ED83D1" pageId="14" pageNumber="105">
<taxonomicName id="E18424D69FA95381020F5F74F7AC113A" LSID="2BFF205E-4211-5E91-8BAA-96370CE26F06" authority="Smith" authorityName="Smith" authorityYear="2014" baseAuthorityName="Schmidt &amp; Anweiler" baseAuthorityYear="2014" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raphia frater subsp. elbea" order="Lepidoptera" pageId="14" pageNumber="105" phylum="Arthropoda" rank="subSpecies" species="frater" status="stat. n." subSpecies="elbea">Raphia frater elbea Smith</taxonomicName>
<taxonomicNameLabel id="38DE277E257EAD371DE8F55B3B8DC556" pageId="14" pageNumber="105">stat. n.</taxonomicNameLabel>
<figureCitation id="7C4D16CC85625C03EF366AC3D3817953" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographic distribution and phenotypic variation of Raphia frater subspecies. Circles indicate specimens examined during this study: green - subsp. piazzi; pink - subsp. abrupta; black - subsp. frater; blue - subsp. coloradensis; yellow - subsp. cinderella. Multi-coloured circles indicate transitional populations and / or phenotypically intermediate specimens between respective subspecies. a Raphia frater piazzi (Zavallo Co., TX) b Raphia frater abrupta (Oktibeha Co., MS) c Raphia frater abrupta (Cottle Co., TX) d Raphia frater abrupta (Cottle Co., TX) e Raphia frater abrupta (Montgomery Co., MD) f, g Raphia frater frater (Edmunston, NB) h Raphia frater abrupta - frater intermediate (Anne Arundel Co., MD) i Raphia frater abrupta - frater - coloradensis intermediate from highly variable population in Cherry Co., NE j Raphia frater coloradensis (Alamosa Co., CO) k Raphia frater coloradensis (Milk River valley, AB) l Raphia frater coloradensis (Sanpete Co., UT) m Raphia frater coloradensis (Elko Co., NV) n Raphia frater cinderella (Ventura Co., CA) o, p Raphia frater coloradensis - frater intermediates (Chelan Co., WA) q Raphia frater elbea (Cochise Co., AZ) r Raphia frater elbea (San Juan Co., UT) s Raphia frater elbea (Santa Cruz Co., AZ). All specimens are males." httpUri="https://binary.pensoft.net/fig/30255" pageId="14" pageNumber="105">Figs 1q-s</figureCitation>
<figureCitation id="DB9C70934180716E89E9037DAD852CA3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of Raphia frater subspecies (circles) relative to range of Populus species (coloured shading) in western North America. Half-circles represent transitional populations and / or phenotypically intermediate specimens. Ranges for Populus trichocarpa + Populus balsamifera and Populus angustifolia + Populus deltoides are combined, with both Populus balsamifera and Populus deltoides occuring in Alberta - Montana (upper right). Populus ranges adapted from USGS (2013)." httpUri="https://binary.pensoft.net/fig/30257" pageId="14" pageNumber="105">, 3</figureCitation>
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<subSubSection id="D167E248104A9652F3EBB2FC6C7845C5" pageId="14" pageNumber="105" type="reference_group">
<paragraph id="3D17EF966E3B697005D7ECF06C8E57EA" pageId="14" pageNumber="105">
<taxonomicName id="66CAC9750A636B2AB36707DC995D47C7" authorityName="Schmidt &amp; Anweiler" authorityYear="2014" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="elbea" status="stat. n.">Raphia elbea</taxonomicName>
Smith, 1908
</paragraph>
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<subSubSection id="98066315E006D41EB6DDAE616D13A7F9" pageId="14" pageNumber="105" type="type material">
<paragraph id="F7977734FDFAF53782EB706B23AFFB23" pageId="14" pageNumber="105">Type material.</paragraph>
<paragraph id="F36037A192E039127F03C0AAE75C573E" pageId="14" pageNumber="105">
A male lectotype was designated by 
<bibRefCitation id="66FB0FF2F36B3FD47A000E027D631AE5" author="Todd, EL" journalOrPublisher="United States Department of Agriculture Technical Bulletin" pageId="18" pageNumber="109" pagination="1 - 228" refId="B45" refString="Todd, EL, 1982. The noctuid type material of J. B. Smith (Lepidoptera). United States Department of Agriculture Technical Bulletin 1645: 1 - 228" title="The noctuid type material of J. B. Smith (Lepidoptera)." volume="1645" year="1982">Todd (1982)</bibRefCitation>
[AMNH]. Type locality: Deming, [Luna Co.,], New Mexico [USA].
</paragraph>
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<subSubSection id="DA733BC8F631D7C2EF9A33DBC5380287" lastPageId="15" lastPageNumber="106" pageId="14" pageNumber="105" type="diagnosis and description">
<paragraph id="A86EF0142EDBC64035212F752DA5F0C4" pageId="14" pageNumber="105">Diagnosis and description.</paragraph>
<paragraph id="978E7E3C48A76E02E25FE3BB1E172A63" pageId="15" pageNumber="106">
<taxonomicName id="9A214B34F547D2CBF26F0ABA227FFA03" authorityName="Smith" authorityYear="2014" baseAuthorityName="Schmidt &amp; Anweiler" baseAuthorityYear="2014" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="15" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="frater" subSpecies="elbea">
<emphasis id="76F7271CFE8560A0D3A7B82734839BA9" italics="true" pageId="15" pageNumber="106">
<pageBreakToken id="A47544860ABABBD889FCC01AE6713A3E" pageId="15" pageNumber="106" start="start">Raphia</pageBreakToken>
frater elbea
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is most similar to the pale yellowish-ochre forms of 
<taxonomicName id="09EA4B4ED87DB939610C91F200C2269F" authorityName="Putnam-Cramer" authorityYear="2014" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="15" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="frater" subSpecies="coloradensis">
<emphasis id="F1667DFC829222634DF9B85917F0E0C4" italics="true" pageId="15" pageNumber="106">Raphia frater coloradensis</emphasis>
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, but differ from that subspecies in having both the costal and reniform dark patches more prominent; when present, the black medio-anal patch is also darker and more elongate; additionally, 
<taxonomicName id="61C7E1346FC75301FC026BE4F7F1B011" authorityName="Smith" authorityYear="2014" baseAuthorityName="Schmidt &amp; Anweiler" baseAuthorityYear="2014" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="15" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="frater" subSpecies="elbea">
<emphasis id="04959BBEE4D978D299B3F005461BCB39" italics="true" pageId="15" pageNumber="106">Raphia frater elbea</emphasis>
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appears to exhibit a unique, divergent mtDNA haplotype group.
</paragraph>
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<subSubSection id="F18E6C7A44B077ACDF157566F69108DC" pageId="15" pageNumber="106" type="biology and distribution">
<paragraph id="95450EAF5669E2AE7D40A8A8C0DEFFDA" pageId="15" pageNumber="106">Biology and distribution.</paragraph>
<paragraph id="3B1207338A871432BE598B25E7C52B44" pageId="15" pageNumber="106">
This subspecies occurs from southeastern Utah and western New Mexico southward through Arizona into northern Mexico. In southeastern Arizona it occurs in riparian areas in association with the larval host, 
<taxonomicName id="AF7D865640626AC35D39291ED47E6BF5" class="Dicotyledoneae" family="Salicaceae" genus="Populus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Salicales" pageId="15" pageNumber="106" phylum="Angiospermae" rank="species" species="fremonti">
<emphasis id="45B92496509AE62CE3A2355095E81622" italics="true" pageId="15" pageNumber="106">Populus fremonti</emphasis>
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. Flight records are from February to October, with most being from March to May and August to September, indicating at least two generations annually.
</paragraph>
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