New cocalodine jumping spiders from Papua New Guinea (Araneae: Salticidae: Cocalodinae)
Author
Maddison, Wayne P.
text
Zootaxa
2009
2021
1
22
journal article
37721
10.5281/zenodo.186069
99d44e7b-46c3-47a2-9695-3ea43c4186a0
1175-5326
186069
Cucudeta gahavisuka
new species
(
Figs 66–67
,
83–86
)
Type
material.
Holotype
male in
UBC
–
SEM
with labels: "
PAPUA
NEW
GUINEA
: Eastern Highlands Province: Mt.
Gahavisuka
Provincial Park.
S 6.01451
E 145.41171
. elev.
2318 m
a.s.l. Leaf litter in forest.
1–2 August 2008
. W. Maddison. WPM#08–028.", "
Cucudeta gahavisuka Maddison
,
Holotype
", and "
UBC
–
SEM AR
00026".
Paratypes
:
3 males
,
9 females
and
11 juveniles
.
Etymology.
Named after the
type
locality.
Diagnosis.
Most specimens are larger than those of the other two
Cucudeta
species, and with pale chevron markings on the dorsum of the opisthosoma (
Figs 66–67
). The male palp is very different, with a small uncoiled embolus and inconspicuous median apophysis (
Fig. 83
). Although in all species of
Cucudeta
the area of the joint between the first patella and tibia is transparent white, in
C. gahavisuka
this pale area is particularly pronounced.
FIGURES 68–71.
Walking poses of
Cucudeta
species. 68–69
C. zabkai
, male. 70–71
C. uzet
, female.
Description.
Male
(
holotype
). Carapace length 1.5; abdomen length 1.5. Chelicerae: at least two promarginal and four retromarginal teeth; one of the retromarginal teeth is long and near the base of the fang, and may be homologous to the apophyses cited under
C. zabkai
and
C. uzet
. Palpus (
Figs 83–84
): embolus short, hidden. The median apophysis is distinct though smaller than in the other two species. There is a flange near the median apophysis that may be the conductor. Tibia of first leg with no ventral macrosetae; first metatarsus with 4 pairs. Carapace relatively flat, PME small and medially displaced. Colour (
Fig. 67
): carapace dark brown to black, paler centrally, with thin golden setae in ocular area. Legs pale to medium brown with first leg darker. Abdomen black with white chevrons dorsally, flanked by a longitudinal white stripe and two spots on each side. The tip of the posterior spinnerets is white.
FIGURES 72–86.
Cucudeta
species. 72–78
C. zabkai
,
new species
. 72–73 Palp of holotype male (72 ventral view,73 retrolateral view). 74 Metatarsus and tarsus of first leg, female. 75–76 female carapace. 77 Epigynum, ventral view. 78 Cleared epigynum, dorsal view. 79–82
C. uzet
,
new species
. 79–80 Palp of holotype male (79 ventral view, 80 retrolateral view). 81 Epigynum, ventral view. 82 Cleared epigynum, dorsal view. 83–86
C. gahavisuka
,
new species
. 83–84 Palp of holotype male (83 ventral view,84 retrolateral view). Figure 85 Epigynum, ventral view. 86 Cleared epigynum, dorsal view. Scale bars 0.1 mm on genitalia and first leg, 1 mm on carapace.
FIGURES 87–92.
Habitats of cocalodines. Microhabitat indicated by arrows. 87 Habitat of
Allococalodes madidus
, on mosses on small tree trunks along stream (Umgé, S 5.3054 E 142.5098); approximate width of field 1.5 m. 88 Habitat of
Yamangalea frewana
, on tree branches and trunks (Mt.
Gahavisuka
, S 6.015 E 145.412); approximate diameter of branch indicated 5 cm. Most were found on larger, vertical tree trunks. 89 Habitat of
Tabuina varirata
, on conifer in clearing (
Varirata
National Park, S 9.436 E 147.364); approximate width of field 0.7 m. 90 Habitat of
Cucudeta zabkai
, in leaf litter at base of
Pandanus
(near Tualapa, S 5.291 E 142.500); approximate width of field 1 m. 91 Habitat of
Cucudeta uzet
, in leaf litter at base of
Pandanus
(Umgé, S 5.304 E 142.512); approximate width of field 1 m. 92 Habitat of
Cucudeta gahavisuka
, in leaf litter on terraces on slope in forest (Mt.
Gahavisuka
, S 6.01451 E 145.41171); approximate width of field 1.5 m.
Female
(
paratype
, UBC–SEM AR00027). Carapace length 1.6; abdomen length 2.1. Chelicerae: at least three promarginal and four retromarginal teeth. Palp without tarsal claw, or if present then highly reduced. Tibia of first leg with no ventral macrosetae; first metatarsus with 4 pairs. Epigynum (
Figs 85–86
) with much shorter ducts than in the other two species, corresponding to the shorter embolus. Carapace relatively flat, PME small and medially displaced. Colour (
Fig. 66
): carapace dark brown to black. Legs pale to dark brown; base of first tibia distinctly paler. Abdomen brown with central paler chevrons.