A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae) Author Edmonds, W. D. Author Zidek, J. Coprophanaeus bonariensis (Gory) text Insecta Mundi 2010 2010-07-02 2010 129 1 111 journal article 10.5281/zenodo.5352924 1942-1354 5352924 Coprophanaeus ( C .) cerberus ( Harold, 1869 ) Fig. 9 , 95 , 104 , 113 , 124-128 Phanaeus cerberus Harold, 1869: 65 Coprophanaeus cerberus (Harold) (recomb. by Blackwelder 1944: 209 ) Type. Lectotype male (des. by Arnaud, 1982:115), Muséum National d’Histoire Naturelle, Paris (examined by photo). Diagnosis. General – Dorsum black or brown, lacking metallic coloration. Clypeal margin noticeably angulate or explanate lateral to median teeth ( Fig. 124, 127 ). Pronotum weakly granulose anterolaterally, disk smooth except for elongate, anteriorly narrowed depression filled with coalesced coarse granules often mixed with punctures extending from posterior margin toward middle of disk ( Fig. 95 ). Basal pronotal fossae effaced or only weakly indicated. Length 19-25 mm . Male ( Fig. 95 , 104 , 126-128 ) – Middle process of head horn short, length about one-fourth that of horn base. Anteromedian pronotal concavity shallow, bordered posteriorly by weak transverse carina bearing rounded central tubercle; ends of transverse carina thickened, lacking distinct tubercle. Female ( Fig. 124-125 ) – Cephalic carina straight, trituberculate, tubercles equal-sized and closely set, distance between lateral tubercles about one-half of interocular distance. Pronotum lacking anteromedian concavity, with simple transverse carina at upper edge of smooth, declivitous area reaching anterior margin; declivity bounded on each side by shallow depression extending toward eye. Specimens examined – 66. Distribution. Brazilian Atlantic Coast Forest province of Paranaian subregion ( Fig. 113 ). Collection Records. BRAZIL : Espiritu SantoParque Estadual Pedra Azul , 1500 m (Feb-Mar). Minas Gerais Poco de Caldas (Jan, Dec) ; Poços do Coicos ( Dec ). Rio de Janeiro 17 km E Nova Friburgo , 22 o 23’04”S 42 o 33’30”W , 750 m (Jan) ; Mury , 8 km E. Nova Friburgo , 22 o 21’49”S 42 o 33’07”W , 1150 m (Jan) ; Itatiaia, 700 m (Feb, Nov); Tijuca Forest (Jan). São Paulo – Catareira (Mar) ; Piauiti; 50 km SE Mogi das Cruzes , Estacão Biológica Boraceia ( Serra do Mar ), 800-900 m (Oct, Dec) ; Bocaina (Apr) . Comments. Olsoufieff’s (1924) erroneously synonymized P . perseus Harold with C. cerberus (see comments under C. corythus ). The coarse sculpturing of the posteromedian groove of the pronotum links this species with C . spitzi and C. acrisius (cf. Fig. 95 and 97 ). The form of the female pronotum is unique to the genus and more reminiscent of females of Phanaeus mexicanus group than other Coprophanaeus . Sculpturing of head and pronotum is weakest of species in the jasius group.