Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae) Author Génier, François Author Moretto, Philippe text Zootaxa 2017 4248 1 1 110 journal article 33324 10.5281/zenodo.439444 cf486ccc-d298-4179-8ccf-e29a9c894ab4 1175-5326 439444 71F0AC03-C1FB-40AB-8532-99A638FC91E9 Digitonthophagus lusinganus species group 11. Digitonthophagus aksumensis Génier , new species Distribution: Djibouti , Eritrea , Ethiopia , Saudi Arabia , Somalia , Sudan , Yemen 12. Digitonthophagus dilatatus Génier , new species Distribution: Burkina Faso , Cameroon , Chad , Côte d’Ivoire , Ghana , Nigeria 13. Digitonthophagus fimator Génier , new species Distribution: Benin , Burkina Faso , Cameroon , Central Africa republic, Chad , Côte d'Ivoire , Democratic Republic of the Congo , Ethiopia, Ghana, Guinea, Guinea-Bissau, Nigeria, Senegal, Sierra Leone , Togo , Uganda 14. Digitonthophagus lusinganus (d’Orbigny, 1905) , valid species Distribution: Ethiopia , Kenya , Somalia , South Sudan , Sudan , Tanzania 15. Digitonthophagus ulcerosus Génier , new species Distribution: Ethiopia , Kenya , Somalia , Tanzania