<document id="3922272CA76D6CC0202075EDB696154C" ID-CLB-Dataset="308253" ID-DOI="10.11646/zootaxa.5588.4.1" ID-GBIF-Dataset="e8d80c74-68cf-4598-9172-65280e118626" ID-ISSN="1175-5326" ID-Zenodo-Dep="14896684" ID-ZooBank="444F5F59-9228-4100-9135-DF824FF9BACA" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1740004642831" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2025" docId="50613239FF866C49FF0AFCE2FBFCFC39" docLanguage="en" docName="zootaxa.5588.4.1.pdf" docOrigin="Zootaxa 5588 (4)" docSource="https://doi.org/10.11646/zootaxa.5588.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Omalium sectum Shavrin, 2025, sp. nov." docType="treatment" docVersion="6" lastPageNumber="510" masterDocId="AC584A41FF8E6C40FF9DFFA0FFBBFFF0" masterDocTitle="Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China" masterLastPageNumber="543" masterPageNumber="501" pageNumber="509" updateTime="1740481210813" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3B20693F9BD8F9AA40D7695154F90C57">Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China</mods:title>
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<mods:namePart id="804E0EA7A2D78FCB14D6B7C14256423C">Shavrin, Alexey V.</mods:namePart>
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<mods:affiliation id="FC2671D7757CDF4116C76845954736C4">Institute of Life Sciences and Technologies, Daugavpils University, Vienibas 13, Daugavpils, LV- 5401, Latvia</mods:affiliation>
<mods:nameIdentifier id="D042724E2A732FB581A191CAE7A5D9E2" type="email">ashavrin@hotmail.com</mods:nameIdentifier>
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<mods:date id="DA3EA87BB089DF00418D65BE8B79221E">2025</mods:date>
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<treatment id="50613239FF866C49FF0AFCE2FBFCFC39" ID-DOI="http://doi.org/10.5281/zenodo.14918408" ID-GBIF-Taxon="254531113" ID-Zenodo-Dep="14918408" LSID="urn:lsid:plazi:treatment:50613239FF866C49FF0AFCE2FBFCFC39" httpUri="http://treatment.plazi.org/id/50613239FF866C49FF0AFCE2FBFCFC39" lastPageId="9" lastPageNumber="510" pageId="8" pageNumber="509" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Omaliinae" scope_tribe="Omaliini">
<subSubSection id="90D2D0A4FF866C48FF0AFCE2FDAFFCAC" box="[151,532,834,861]" pageId="8" pageNumber="509" type="nomenclature">
<paragraph id="D877832FFF866C48FF0AFCE2FDAFFCAC" blockId="8.[151,532,834,896]" box="[151,532,834,861]" pageId="8" pageNumber="509">
<heading id="833F3443FF866C48FF0AFCE2FDAFFCAC" bold="true" box="[151,532,834,861]" fontSize="11" level="1" pageId="8" pageNumber="509" reason="1">
<emphasis id="EABC5F3DFF866C48FF0AFCE2FDAFFCAC" bold="true" box="[151,532,834,861]" pageId="8" pageNumber="509">
3.2.3. 
<taxonomicName id="1FC8F8ACFF866C48FF7CFCE2FE14FCAD" authorityName="Shavrin" authorityYear="2025" box="[225,431,834,861]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="sectum" status="sp. nov.">
<emphasis id="EABC5F3DFF866C48FF7CFCE2FE14FCAD" bold="true" box="[225,431,834,861]" italics="true" pageId="8" pageNumber="509">Omalium sectum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F18FE246FF866C48FE2BFCE2FDAFFCAC" box="[438,532,834,860]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF866C48FF0AFCC6FED6FC70" box="[151,365,870,896]" pageId="8" pageNumber="509" type="description">
<paragraph id="D877832FFF866C48FF0AFCC6FED6FC70" blockId="8.[151,532,834,896]" box="[151,365,870,896]" pageId="8" pageNumber="509">
(
<figureCitation id="40F39FAAFF866C48FF02FCC6FF5CFC70" box="[159,231,870,896]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1097,1121]" captionTargetBox="[151,1436,181,1073]" captionTargetId="figure-16@3.[151,1436,181,1073]" captionTargetPageId="3" captionText="FIGURES 1–3. Habitus of Omalium: 1—O. anguliferum (holotype). 2—O. rutilum (paratype), 3—O. sectum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896686" httpUri="https://zenodo.org/record/14896686/files/figure.png" pageId="8" pageNumber="509">Figs 3</figureCitation>
, 
<figureCitation id="40F39FAAFF866C48FF69FCC6FE87FC70" box="[244,316,870,896]" captionStart="FIGURES 4–12" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 4–12. Aedeagus of Omalium (Figs 4–6—O. anguliferum (paratype), Figs 7–9—O. rutilum (paratype), 10–12—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="8" pageNumber="509">10–12</figureCitation>
, 
<figureCitation id="40F39FAAFF866C48FED5FCC6FEDEFC70" box="[328,357,870,896]" captionStart="FIGURE 30" captionStartId="9.[151,250,1849,1873]" captionTargetBox="[151,1435,993,1826]" captionTargetId="figure-318@9.[151,1435,993,1826]" captionTargetPageId="9" captionText="FIGURE 30. Distribution of Omalium in China: O. anguliferum (circles), O. rutilum (squares), O. sectum (triangle)." figureDoi="http://doi.org/10.5281/zenodo.14896690" httpUri="https://zenodo.org/record/14896690/files/figure.png" pageId="8" pageNumber="509">30</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF866C48FF0AFC08FC66FBA1" pageId="8" pageNumber="509" type="materials_examined">
<paragraph id="D877832FFF866C48FF0AFC08FC66FBA1" blockId="8.[151,1437,935,2041]" pageId="8" pageNumber="509">
<emphasis id="EABC5F3DFF866C48FF0AFC08FE78FC31" bold="true" box="[151,451,935,962]" pageId="8" pageNumber="509">Type material examined.</emphasis>
<materialsCitation id="68A08972FF866C48FE51FC07FC66FBA1" ID-GBIF-Occurrence="5069426360" collectingDate="2005-06-11" collectionCode="cSch" country="China" county="Deqin Co." determinerName="M. Schulke" elevation="2580" latitude="28.457834" location="Diqing Tibet. Aut. Pref." longLatPrecision="12" longitude="98.7725" municipality="Meili Xue Shan" pageId="8" pageNumber="509" specimenCount="1" specimenCount-male="1" stateProvince="YUNNAN" typeStatus="holotype">
<typeStatus id="07733D8DFF866C48FE51FC07FD8FFC31" box="[460,564,935,961]" pageId="8" pageNumber="509" type="holotype">Holotype</typeStatus>
<specimenCount id="CECE48A6FF866C48FDA0FC08FDE9FC32" box="[573,594,936,962]" count="1" pageId="8" pageNumber="509" type="male">♂</specimenCount>
(
<figureCitation id="40F39FAAFF866C48FDFEFC07FD11FC32" box="[611,682,935,962]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1097,1121]" captionTargetBox="[151,1436,181,1073]" captionTargetId="figure-16@3.[151,1436,181,1073]" captionTargetPageId="3" captionText="FIGURES 1–3. Habitus of Omalium: 1—O. anguliferum (holotype). 2—O. rutilum (paratype), 3—O. sectum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896686" httpUri="https://zenodo.org/record/14896686/files/figure.png" pageId="8" pageNumber="509">Fig. 3</figureCitation>
; dissected): 
<emphasis id="EABC5F3DFF866C48FCA4FC07FBA8FC32" bold="true" box="[825,1043,935,962]" pageId="8" pageNumber="509">
<collectingCountry id="A0DFC3BFFF866C48FCA4FC07FC20FC31" box="[825,923,935,961]" name="China" pageId="8" pageNumber="509">CHINA</collectingCountry>
: 
<collectingRegion id="1A0C4DCDFF866C48FC35FC07FBB1FC31" box="[936,1034,935,961]" country="China" name="Yunnan" pageId="8" pageNumber="509">YUNNAN</collectingRegion>
:
</emphasis>
‘CHINA: N-Yunnan [C2005-09] | 
<location id="DD17D5F4FF866C48FF0AFC6BFE1FFC15" LSID="urn:lsid:plazi:treatment:50613239FF866C49FF0AFCE2FBFCFC39:DD17D5F4FF866C48FF0AFC6BFE1FFC15" box="[151,420,971,998]" country="China" county="Deqin Co." latitude="28.457834" longLatPrecision="12" longitude="98.7725" municipality="Meili Xue Shan" name="Diqing Tibet. Aut. Pref." pageId="8" pageNumber="509" stateProvince="YUNNAN">Diqing Tibet. Aut. Pref.</location>
, | 
<collectingCounty id="3116FBA3FF866C48FE26FC6BFD8AFC16" box="[443,561,971,998]" pageId="8" pageNumber="509">Deqin Co.</collectingCounty>
, 
<collectingMunicipality id="38131955FF866C48FDA0FC6BFD4AFC15" box="[573,753,971,998]" pageId="8" pageNumber="509">Meili Xue Shan</collectingMunicipality>
, | E-side, 
<locationDeviation id="C9C59A0DFF866C48FCC2FC6CFC7CFC16" bearing="270.0" box="[863,967,971,998]" distance="14000.0" pageId="8" pageNumber="509">
<quantity id="1F302ECAFF866C48FCC2FC6CFC1DFC15" box="[863,934,971,998]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="509" unit="km" value="14.0">14 km</quantity>
W
</locationDeviation>
<location id="DD17D5F4FF866C48FC53FC6BFBAEFC15" LSID="urn:lsid:plazi:treatment:50613239FF866C49FF0AFCE2FBFCFC39:DD17D5F4FF866C48FC53FC6BFBAEFC15" box="[974,1045,971,997]" country="China" county="Deqin Co." latitude="28.457834" longLatPrecision="12" longitude="98.7725" municipality="Meili Xue Shan" name="Deqin" pageId="8" pageNumber="509" stateProvince="YUNNAN">Deqin</location>
, 
<quantity id="1F302ECAFF866C48FBBCFC6BFBC1FC16" box="[1057,1146,971,998]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="8" pageNumber="509" unit="m" value="2580.0">
<elevation id="53E5641CFF866C48FBBCFC6BFBC1FC16" box="[1057,1146,971,998]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="8" pageNumber="509" unit="m" value="2580.0">2580 m</elevation>
</quantity>
,’ &lt;printed&gt;, ‘ 
<geoCoordinate id="BDFCE5E8FF866C48FA90FC6CFA23FC16" box="[1293,1432,972,998]" degrees="28" direction="north" minutes="27.47" orientation="latitude" pageId="8" pageNumber="509" precision="9" value="28.457834">28°27.47´N</geoCoordinate>
, 
<geoCoordinate id="BDFCE5E8FF866C48FF0AFC4FFEA1FBFA" box="[151,282,1007,1034]" degrees="98" direction="east" minutes="46.35" orientation="longitude" pageId="8" pageNumber="509" precision="9" value="98.7725">98°46.35´E</geoCoordinate>
, creek | valley below glacier, mixed forest, | leaf litter, moss, dead wood, sifted, | 
<date id="AC76A5EFFF866C48FB0CFC4FFAA8FBFA" box="[1169,1299,1007,1034]" pageId="8" pageNumber="509" value="2005-06-11">
<collectingDate id="BC325C07FF866C48FB0CFC4FFAA8FBFA" box="[1169,1299,1007,1034]" pageId="8" pageNumber="509" value="2005-06-11">11.VI.2005</collectingDate>
</date>
, 
<determinerName id="2DAB6FEBFF866C48FA80FC50FA27FBF9" box="[1309,1436,1007,1034]" pageId="8" pageNumber="509">M. Schülke</determinerName>
[C2005-09]’ &lt;printed&gt;, ‘Museum für Naturkunde | Berlin | Sammlung M. Schülke’ &lt;printed&gt;, ‘HOLOTYPE | 
<taxonomicName id="1FC8F8ACFF866C48FF0AFB97FF44FBA1" authorityName="Gravenhorst" authorityYear="1802" box="[151,255,1079,1105]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="genus">
<emphasis id="EABC5F3DFF866C48FF0AFB97FF44FBA1" box="[151,255,1079,1105]" italics="true" pageId="8" pageNumber="509">Omalium</emphasis>
</taxonomicName>
| 
<taxonomicName id="1FC8F8ACFF866C48FE8EFB98FEDAFBA1" authorityName="Shavrin" authorityYear="2025" box="[275,353,1080,1105]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="sectum" status="sp. nov.">
<emphasis id="EABC5F3DFF866C48FE8EFB98FEDAFBA1" box="[275,353,1080,1105]" italics="true" pageId="8" pageNumber="509">sectum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF866C48FEF5FB98FE7AFBA2" bold="true" box="[360,449,1080,1106]" pageId="8" pageNumber="509">
<taxonomicNameLabel id="F18FE246FF866C48FEF5FB98FE7AFBA2" box="[360,449,1080,1106]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2025’ &lt;red, printed&gt; (
<collectionCode id="BED91BEAFF866C48FC08FB97FC6AFBA1" box="[917,977,1079,1105]" pageId="8" pageNumber="509">cSch</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF866C49FF5AFBFBFE2AFD59" lastPageId="9" lastPageNumber="510" pageId="8" pageNumber="509" type="description">
<paragraph id="D877832FFF866C48FF5AFBFBFEF9FB4E" blockId="8.[151,1437,935,2041]" pageId="8" pageNumber="509">
<emphasis id="EABC5F3DFF866C48FF5AFBFBFEE2FB85" bold="true" box="[199,345,1115,1141]" pageId="8" pageNumber="509">Description.</emphasis>
Measurements: HW: 0.62; HL: 0.43; OL: 0.17; TL: 0.12; AL: 1.35; PL: 0.55; PWmax: 0.72; PWmin: 0.67; ESL: 1.01; EW: 1.16; MTbL: 0.35; MTrL: 0.17 (MTrL 1–4: 0.07; MTrL 5: 0.10); AW: 1.02; AedL: 0.70; BL: 3.54.
</paragraph>
<paragraph id="D877832FFF866C48FF5AFB67FBEAFA65" blockId="8.[151,1437,935,2041]" pageId="8" pageNumber="509">
Habitus as in 
<figureCitation id="40F39FAAFF866C48FEC3FB67FE1BFB12" box="[350,416,1223,1250]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1097,1121]" captionTargetBox="[151,1436,181,1073]" captionTargetId="figure-16@3.[151,1436,181,1073]" captionTargetPageId="3" captionText="FIGURES 1–3. Habitus of Omalium: 1—O. anguliferum (holotype). 2—O. rutilum (paratype), 3—O. sectum (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14896686" httpUri="https://zenodo.org/record/14896686/files/figure.png" pageId="8" pageNumber="509">Fig. 3</figureCitation>
. Body reddish-brown, with slightly paler abdomen; antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellow-brown. Punctation of head dense, denser and coarser in middle and on infraorbital portions; neck with moderately fine and sparse punctation; punctation of pronotum fine and sparse, sparser in medioapical, lateral and mediobasal portions; punctation of pronotum distinctly denser than that on pronotum, coarser in middle; abdominal tergites without punctation. Clypeus with fine transverse microsculpture; scutellum with fine isodiametric microreticulation; abdomen with dense isodiametric sculpture.
</paragraph>
<paragraph id="D877832FFF866C48FF5AFA3FFD6BF8B6" blockId="8.[151,1437,935,2041]" pageId="8" pageNumber="509">Head 1.4 times as broad as long, with broad and deep anteriomedian depressions; posteriolateral margins of clypeus stretching toward level of middle length of eyes. Latero-apical margin between anterior margin of eyes and clypeus with small semicircular notch. Dorsal surface with distinct transverse and longitudinal elevations between punctures in middle and irregular elongate elevations on postocular parts. Anteocellar foveae very wide and deep, somewhat elongate, reaching level of posterior third length of eyes. Temples 1.4 times as long as longitudinal length of eye, from posterior margin of eyes gradually narrowed posteriad, each with distinct subacute angle protruded basad. Apical part of neck relatively deeply depressed, with short irregular and narrow longitudinal elevations. Distance between ocelli 1.7 times as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere about three times as long as penultimate segment. Antenna with distinctly elongate antennomeres 2–8 and slightly elongate 9–10; antennomere 3 slightly longer and narrower than 2, 4 distinctly shorter than 3, 5 longer than 4, 6–7 longer and distinctly broader than 5, 8 slightly shorter than 7, 9–10 slightly shorter and distinctly broader than 8, apical antennomere slightly longer than 10.</paragraph>
<paragraph id="D877832FFF866C48FF5AF8F0FC80F809" blockId="8.[151,1437,935,2041]" pageId="8" pageNumber="509">Pronotum 1.3 times as broad as long, 1.1 times as broad as head, from widest anterior portion gradually narrowed posteriad toward widely rounded hind angles. Anterior angles acute and protruded anteriad. Anterior margin rounded, distinctly protruded anteriad. Each laterobasal portion not concave. Surface of disc with two wide and deep longitudinal and oval and deep anteriomedian depressions; surface between all depressions strongly elevated. Middle portion with irregular indistinct elevations.</paragraph>
<paragraph id="D877832FFF876C49FF5AFF37FD14FF09" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="510">Elytra 1.1 times as broad as long, 1.8 times as long as pronotum, from basal portion distinctly broadened posteriad; hind margins straight. Dorsal surface of each elytron with irregular elevations, diagonal in medioapical and median, and longitudinal in apical portions.</paragraph>
<paragraph id="D877832FFF876C49FF5AFEA3FC9FFEED" blockId="9.[151,1437,151,969]" box="[199,804,259,285]" pageId="9" pageNumber="510">Metatarsi slightly less than twice as long as metatibia.</paragraph>
<paragraph id="D877832FFF876C49FF5AFE87FF07FE95" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="510">Abdomen slightly narrower than elytra, with wide intersegmental membranes between abdominal tergites V– VI.</paragraph>
<paragraph id="D877832FFF876C49FF5AFECFFD63FD75" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="510">
Male. Posterior margin of abdominal tergite VIII straight. Posterior margin of abdominal sternite VIII sinuate. Aedeagus with wide basal portion, gradually narrowed toward median lobe; median lobe moderately long, from narrow basal part significantly broadened slightly in front middle, strongly narrowed in preapical part and broadened in apical portion, with widely rounded apex; accessory plates narrow, curved and relatively long, rounded apically; parameres wide and long, reaching apical part of median lobe, from widest preapical parts gradully narrowed apically, each with two moderately short apical and two preapical setae; internal sac narrow and long, without sclerotized structures (
<figureCitation id="40F39FAAFF876C49FE08FDE7FE51FD91" box="[405,490,583,609]" captionStart="FIGURES 4–12" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 4–12. Aedeagus of Omalium (Figs 4–6—O. anguliferum (paratype), Figs 7–9—O. rutilum (paratype), 10–12—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="9" pageNumber="510">Fig. 10</figureCitation>
). Lateral aspect of the aedeagus as in 
<figureCitation id="40F39FAAFF876C49FC04FDE7FC50FD91" box="[921,1003,583,609]" captionStart="FIGURES 4–12" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 4–12. Aedeagus of Omalium (Figs 4–6—O. anguliferum (paratype), Figs 7–9—O. rutilum (paratype), 10–12—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="9" pageNumber="510">Fig. 11</figureCitation>
; apical portion of median lobe (lateral view) with crenulate ventrolateral margin (
<figureCitation id="40F39FAAFF876C49FDEAFDCBFD71FD75" box="[631,714,619,645]" captionStart="FIGURES 4–12" captionStartId="6.[151,264,1630,1654]" captionTargetBox="[159,1428,281,1599]" captionTargetId="figure-60@6.[151,1436,273,1606]" captionTargetPageId="6" captionText="FIGURES 4–12. Aedeagus of Omalium (Figs 4–6—O. anguliferum (paratype), Figs 7–9—O. rutilum (paratype), 10–12—O. sectum (holotype)): 4, 7, 10—aedeagus, parameral view, 5, 8, 11—aedeagus, lateral view, 6, 9, 12—median lobe, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14896688" httpUri="https://zenodo.org/record/14896688/files/figure.png" pageId="9" pageNumber="510">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="D877832FFF876C49FF5AFD2FFE2AFD59" blockId="9.[151,1437,151,969]" box="[199,401,655,681]" pageId="9" pageNumber="510">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF876C49FF5AFD12FA85FD01" pageId="9" pageNumber="510" type="discussion">
<paragraph id="D877832FFF876C49FF5AFD12FA85FD01" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="510">
<emphasis id="EABC5F3DFF876C49FF5AFD12FE0BFD3D" bold="true" box="[199,432,690,717]" pageId="9" pageNumber="510">Comparative notes.</emphasis>
<taxonomicName id="1FC8F8ACFF876C49FE2BFD13FDC8FD3C" authorityName="Shavrin" authorityYear="2025" box="[438,627,691,717]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="sectum" status="sp. nov.">
<emphasis id="EABC5F3DFF876C49FE2BFD13FDC8FD3C" box="[438,627,691,717]" italics="true" pageId="9" pageNumber="510">Omalium sectum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF876C49FDE4FD13FD69FD3D" bold="true" box="[633,722,691,717]" pageId="9" pageNumber="510">
<taxonomicNameLabel id="F18FE246FF876C49FDE4FD13FD69FD3D" box="[633,722,691,717]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the remaining species of the acutangulum group by the shape of the pronotum with acute anterior angles protruded anteriad, and the shape of the median lobe.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF876C49FF5AFD5AFD31FCC9" pageId="9" pageNumber="510" type="distribution">
<paragraph id="D877832FFF876C49FF5AFD5AFD31FCC9" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="510">
<emphasis id="EABC5F3DFF876C49FF5AFD5AFEDAFCE4" bold="true" box="[199,353,762,788]" pageId="9" pageNumber="510">Distribution.</emphasis>
<taxonomicName id="1FC8F8ACFF876C49FEF7FD5BFD91FCE4" authorityName="Shavrin" authorityYear="2025" box="[362,554,763,789]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="sectum" status="sp. nov.">
<emphasis id="EABC5F3DFF876C49FEF7FD5BFD91FCE4" box="[362,554,763,789]" italics="true" pageId="9" pageNumber="510">Omalium sectum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF876C49FDA9FD5BFD2BFCE5" bold="true" box="[564,656,763,789]" pageId="9" pageNumber="510">
<taxonomicNameLabel id="F18FE246FF876C49FDA9FD5BFD2BFCE5" box="[564,656,763,789]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from one locality in Meili Xue Shan and Diancang Shan mountain ranges in 
<collectingRegion id="1A0C4DCDFF876C49FEE9FCBFFE76FCC9" box="[372,461,799,825]" country="China" name="Yunnan" pageId="9" pageNumber="510">Yunnan</collectingRegion>
, 
<collectingCountry id="A0DFC3BFFF876C49FE4AFCBFFDA1FCC9" box="[471,538,799,825]" name="China" pageId="9" pageNumber="510">China</collectingCountry>
(
<figureCitation id="40F39FAAFF876C49FDB7FCBFFDC5FCC9" box="[554,638,799,825]" captionStart="FIGURE 30" captionStartId="9.[151,250,1849,1873]" captionTargetBox="[151,1435,993,1826]" captionTargetId="figure-318@9.[151,1435,993,1826]" captionTargetPageId="9" captionText="FIGURE 30. Distribution of Omalium in China: O. anguliferum (circles), O. rutilum (squares), O. sectum (triangle)." figureDoi="http://doi.org/10.5281/zenodo.14896690" httpUri="https://zenodo.org/record/14896690/files/figure.png" pageId="9" pageNumber="510">Fig. 30</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF876C49FF5AFCE3FF74FC71" pageId="9" pageNumber="510" type="etymology">
<paragraph id="D877832FFF876C49FF5AFCE3FF74FC71" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="510">
<emphasis id="EABC5F3DFF876C49FF5AFCE3FEF5FCAD" bold="true" box="[199,334,835,861]" pageId="9" pageNumber="510">Etymology.</emphasis>
The specific epithet is the Latin 
<emphasis id="EABC5F3DFF876C49FD53FCE3FCA9FCAC" box="[718,786,835,860]" italics="true" pageId="9" pageNumber="510">sectus</emphasis>
, - 
<emphasis id="EABC5F3DFF876C49FCB1FCE3FC81FCAC" box="[812,826,835,860]" italics="true" pageId="9" pageNumber="510">a</emphasis>
, - 
<emphasis id="EABC5F3DFF876C49FCC9FCE3FCCDFCAC" box="[852,886,835,860]" italics="true" pageId="9" pageNumber="510">um</emphasis>
(divided). It alludes to the shape of the median lobe.
</paragraph>
</subSubSection>
<subSubSection id="90D2D0A4FF876C49FF5AFC2AFBFCFC39" pageId="9" pageNumber="510" type="biology_ecology">
<paragraph id="D877832FFF876C49FF5AFC2AFBFCFC39" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="510">
<emphasis id="EABC5F3DFF876C49FF5AFC2AFEF0FC54" bold="true" box="[199,331,906,932]" pageId="9" pageNumber="510">Bionomics.</emphasis>
Specimens were collected at elevation 
<quantity id="1F302ECAFF876C49FC85FC2AFCCAFC55" box="[792,881,906,933]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="9" pageNumber="510" unit="m" value="2580.0">2580 m</quantity>
a.s.l. and was sifted from leaf litter, debris and mosses. It was collected together with 
<taxonomicName id="1FC8F8ACFF876C49FDD5FC0FFD06FC39" authorityName="Shavrin" authorityYear="2025" box="[584,701,943,969]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="rutilum" status="sp. nov.">
<emphasis id="EABC5F3DFF876C49FDD5FC0FFD06FC39" box="[584,701,943,969]" italics="true" pageId="9" pageNumber="510">O. rutilum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF876C49FD59FC0FFCA5FC39" bold="true" box="[708,798,943,969]" pageId="9" pageNumber="510">
<taxonomicNameLabel id="F18FE246FF876C49FD59FC0FFCA5FC39" box="[708,798,943,969]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="1FC8F8ACFF876C49FCC9FC0FFC5DFC39" authorityName="Shavrin" authorityYear="2025" box="[852,998,943,969]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="tibetanum" status="sp. nov.">
<emphasis id="EABC5F3DFF876C49FCC9FC0FFC5DFC39" box="[852,998,943,969]" italics="true" pageId="9" pageNumber="510">O. tibetanum</emphasis>
</taxonomicName>
<emphasis id="EABC5F3DFF876C49FC70FC0FFBFCFC39" bold="true" box="[1005,1095,943,969]" pageId="9" pageNumber="510">
<taxonomicNameLabel id="F18FE246FF876C49FC70FC0FFBFCFC39" box="[1005,1095,943,969]" pageId="9" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>