Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood Author Frolov, Andrey V. Author Montreuil, Olivier Author Akhmetova, Lilia A. text Zootaxa 2016 4207 1 1 93 journal article 37549 10.11646/zootaxa.4207.1.1 cef35771-ff95-4a41-9af6-196dea899a64 1175-5326 200444 F2875582-31E2-496F-AEEF-1D657DD86C33 Madecorphnus hanskii Frolov, 2012 ( Figs. 23 A–F) Madecorphnus hanskii Frolov, 2012b : 160 . Type material examined. Holotype ( Figs. 23 A–E), male, “ Madagascar Ambohitantely [ 18°10' S , 47°17' E ] 24.3.2005 , wet forest, I. Hanski-group leg. / Holotypus Madec. hanskii Frolov 2009 ” ( ZIN ) . Diagnosis. Madecorphnus hanskii is similar to M. falciger and M. punctatus in having internal sac of the aedeagus with 3 sclerites: 1 relatively long, spur-like, and 2 comma-shaped or somewhat bifurcate ( Figs. 16 E–F, 22D–E, 23D–E). It can be separated from these species by the shape and relative size of these sclerites and, from the latter, by the shape of parameres and sparse punctation of head and pronotum. Description. Male. Body length 5.3 mm . Color uniformly brown ( Fig. 23 A). Right mandible as long as left, without tooth behind apex. Labrum trapezoidal, with broadly rounded sides, length about 1/6 width (in dorsal view). Frontoclypeus slightly asymmetrical, apically obtuse, with 2 long and 6 shorter setae on the apical margin. Canthus and frontoclypeus almost flat, minutely depressed apicomedially. Head without traces of frontoclypeal suture, finely punctate with minute punctures separated by greater than 5 times their diameter. Pronotum approximately 1.6 times wider than long, widest medially. Disc of pronotum convex, without any depressions, tubercles, or ridges. Punctation on pronotum similar to that on head. Margins with relatively wide border, lateral margins with 4 long setae: 1 seta on basal angle, 1 seta approximately in the middle of lateral margin, and 2 setae on the apical angle. Scutellum triangular, right angled apically, about 1/13 length of elytra. Elytra convex, with distinct humeral humps, widest at basal third, glabrous. First stria distinct and reaching the apex of elytron, other striae indistinct. Disc of elytra with a few relatively large punctures. Epipleura with long, sparse, brown setae. Base of elytron with border. Wings fully developed. Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Protibial apex with robust, spurlike seta and 2 smaller setae basally. Mesothoracic and metathoracic legs similar in shape to each other, metathoracic legs somewhat longer. Longer tibial spur shorter than 2 basal tarsomeres in mesothoracic legs and longer than 2 basal tarsomeres in metathoracic legs. FIGURE 23. Madecorphnus hanskii , holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F). Parameres with teeth laterally, narrowly rounded apically in dorsal view and curved downwards in lateral view ( Fig. 23 B). Internal sac with a long, spur-like sclerite and 2 smaller, somewhat comma-shaped sclerites ( Figs. 23 D– E). Female. Unknown. Distribution and habitat. The type specimen of M. hanskii was collected in wet forest in Ambohitantely special reserve ( Fig. 23 F). The reserve is situated at altitudes of about 1500 m and houses the last remnants of indigenous forest on the central plateau of Madagascar . Madecorphnus hanskii is apparently a forest litter dweller and, due to severe deforestation of the central plateau, its current range may be limited to this small forest remnant.