<document id="8C37650AC63B619DE008C51B4A9B26CE" ID-DOI="10.5281/zenodo.5342006" ID-ISSN="2345-7600" ID-Zenodo-Dep="5342006" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630192162219" checkinUser="diego" docAuthor="Plant, Adrian R." docDate="2009" docId="5D4B87DA235BFFC698B485E1D257FEE0" docLanguage="en" docName="57rbz255-277.pdf" docOrigin="Raffles Bulletin of Zoology 57 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Chelipoda menglunana Grootaert, Yang &amp; Saigusa 2000" docType="treatment" docVersion="4" lastPageNumber="269" masterDocId="A172FFA22356FFC8985F837FD36EFFB5" masterDocTitle="Diversity Of Chelipoda Macquart, 1823 (Diptera: Empididae: Hemerodromiinae) In Northern Thailand With Discussion Of A Biodiversity ‘ Hot Spot’ At Doi Inthanon" masterLastPageNumber="277" masterPageNumber="255" pageNumber="268" updateTime="1699181974063" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A9A9679D5BDF11CB3923CC49D808AD17">Diversity Of Chelipoda Macquart, 1823 (Diptera: Empididae: Hemerodromiinae) In Northern Thailand With Discussion Of A Biodiversity ‘ Hot Spot’ At Doi Inthanon</mods:title>
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<taxonomicName id="12E24D4F235BFFC598B485E1D142F963" ID-CoL="TV4B" authority="Grootaert, Yang &amp; Saigusa, 2000" authorityName="Grootaert, Yang &amp; Saigusa" authorityYear="2000" class="Insecta" family="Empididae" genus="Chelipoda" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="menglunana">
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<emphasis id="E796EADE235BFFC598B485E1D29EF903" bold="true" box="[235,496,1694,1718]" italics="true" pageId="13" pageNumber="268">Chelipoda menglunana</emphasis>
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(
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)
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<emphasis id="E796EADE235BFFC598308462D22EF887" bold="true" box="[111,320,1821,1842]" italics="true" pageId="13" pageNumber="268">Material examined. –</emphasis>
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, 
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, 
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, 
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coll. 
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; 
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(
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, 
<specimenCode id="85449EB7235BFFC59982840CD17AF83D" box="[477,532,1907,1928]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="268">T526</specimenCode>
)
</materialsCitation>
: Nern Wibaak Ditch, 
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, Malaise trap, 
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</materialsCitation>
: Pan Hin Khan Maak Ditch, 
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, Malaise and pan traps, 
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<specimenCount id="C3E4FD45235BFFC59A6C84B5D1E5F86A" box="[563,651,1994,2015]" pageId="13" pageNumber="268" type="male">11 males</specimenCount>
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, (
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<specimenCode id="85449EB7235BFFC5993C8499D2F4F84E" box="[355,410,2022,2043]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="268">T517</specimenCode>
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</materialsCitation>
; 
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<specimenCount id="C3E4FD45235BFFC599F68499D29BF84E" box="[425,501,2022,2043]" pageId="13" pageNumber="268" type="male">2 males</specimenCount>
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<date id="A15C100C235BFFC59A3A8499D180F84E" box="[613,750,2022,2043]" pageId="13" pageNumber="268" value="2006-08-06" valueMax="2006-08-07" valueMin="2006-08-06">
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<specimenCount id="C3E4FD45235BFFC599558B7CD238F7AD" box="[266,342,2051,2072]" pageId="13" pageNumber="268" type="male">5 males</specimenCount>
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<date id="A15C100C235BFFC599968B7CD13CF7AD" box="[457,594,2051,2072]" pageId="13" pageNumber="268" value="2006-08-07" valueMax="2006-08-08" valueMin="2006-08-07">
<collectingDate id="B118E9E4235BFFC599968B7CD13CF7AD" box="[457,594,2051,2072]" pageId="13" pageNumber="268" value="2006-08-07" valueMax="2006-08-08" valueMin="2006-08-07">7–8 Aug.2006</collectingDate>
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(
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; 
<materialsCitation id="658A3C91235BFFC598308B5FD280F780" box="[111,494,2080,2101]" collectingDate="2006-08-08" collectingDateMax="2006-08-09" collectingDateMin="2006-08-08" collectionCode="QSBG" country="Thailand" pageId="13" pageNumber="268" specimenCode="T520" specimenCount="5" specimenCount-male="5" stateProvince="Loei">
<specimenCount id="C3E4FD45235BFFC598308B5FD3D2F780" box="[111,188,2080,2101]" pageId="13" pageNumber="268" type="male">5 males</specimenCount>
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(
<collectionCode id="B3F3AE09235BFFC5993D8B5FD2C8F780" box="[354,422,2080,2101]" pageId="13" pageNumber="268">QSBG</collectionCode>
, 
<specimenCode id="85449EB7235BFFC599EE8B5FD289F780" box="[433,487,2080,2101]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="268">T520</specimenCode>
)
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; 
<materialsCitation id="658A3C91235BFFC59A5E8B5FD747FF66" box="[513,1065,190,2101]" collectingDate="2006-08-09" collectingDateMax="2006-08-10" collectingDateMin="2006-08-09" collectionCode="QSBG" country="Thailand" pageId="13" pageNumber="268" specimenCode="T521" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Loei">
<specimenCount id="C3E4FD45235BFFC59A5E8B5FD128F780" box="[513,582,2080,2101]" pageId="13" pageNumber="268" type="male">1 male</specimenCount>
, 
<specimenCount id="C3E4FD45235BFFC59A0E8B5FD1DCF780" box="[593,690,2080,2101]" pageId="13" pageNumber="268" type="female">3 females</specimenCount>
, 
<date id="A15C100C235BFFC59AE28B5FD0E4FF66" box="[701,906,190,2101]" pageId="13" pageNumber="268" value="2006-08-09" valueMax="2006-08-10" valueMin="2006-08-09">
<collectingDate id="B118E9E4235BFFC59AE28B5FD0E4FF66" box="[701,906,190,2101]" pageId="13" pageNumber="268" value="2006-08-09" valueMax="2006-08-10" valueMin="2006-08-09">9–10 Aug.2006</collectingDate>
</date>
(
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, 
<specimenCode id="85449EB7235BFFC59BB483C1D74FFF66" box="[1003,1057,190,211]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="268">T521</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="658A3C91235BFFC59C6983C1D0BEFF45" collectingDate="2006-08-10" collectingDateMax="2006-08-11" collectingDateMin="2006-08-10" collectionCode="NMWC" country="Thailand" pageId="13" pageNumber="268" specimenCode="T522" specimenCount="10" specimenCount-female="7" specimenCount-male="3" stateProvince="Loei">
<specimenCount id="C3E4FD45235BFFC59C6983C1D7E8FF66" box="[1078,1158,190,211]" pageId="13" pageNumber="268" type="male">3 males</specimenCount>
, 
<specimenCount id="C3E4FD45235BFFC59CCC83C1D798FF66" box="[1171,1270,190,211]" pageId="13" pageNumber="268" type="female">7 females</specimenCount>
, 
<date id="A15C100C235BFFC59D5C83C1D6C6FF66" box="[1283,1448,190,211]" pageId="13" pageNumber="268" value="2006-08-10" valueMax="2006-08-11" valueMin="2006-08-10">
<collectingDate id="B118E9E4235BFFC59D5C83C1D6C6FF66" box="[1283,1448,190,211]" pageId="13" pageNumber="268" value="2006-08-10" valueMax="2006-08-11" valueMin="2006-08-10">10–11 Aug.2006</collectingDate>
</date>
(
<collectionCode id="B3F3AE09235BFFC59B6E83A4D0EAFF45" box="[817,900,219,240]" country="United Kingdom" httpUri="http://grbio.org/cool/e7dr-nxad" name="National Museum of Wales" pageId="13" pageNumber="268" type="Museum">NMWC</collectionCode>
, 
<specimenCode id="85449EB7235BFFC59BCE83A4D0A7FF45" box="[913,969,219,240]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="268">T522</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="658A3C91235BFFC59B8183A4D009FEB8" collectingDate="2006-08-11" collectingDateMax="2006-08-12" collectingDateMin="2006-08-11" collectionCode="QSBG" country="Thailand" pageId="13" pageNumber="268" specimenCode="T523" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Loei">
<specimenCount id="C3E4FD45235BFFC59B8183A4D748FF45" box="[990,1062,219,240]" pageId="13" pageNumber="268" type="male">1 male</specimenCount>
, 
<specimenCount id="C3E4FD45235BFFC59C6C83A4D7F7FF45" box="[1075,1177,219,240]" pageId="13" pageNumber="268" type="female">2 females</specimenCount>
, 
<date id="A15C100C235BFFC59CF983A4D621FF45" box="[1190,1359,219,240]" pageId="13" pageNumber="268" value="2006-08-11" valueMax="2006-08-12" valueMin="2006-08-11">
<collectingDate id="B118E9E4235BFFC59CF983A4D621FF45" box="[1190,1359,219,240]" pageId="13" pageNumber="268" value="2006-08-11" valueMax="2006-08-12" valueMin="2006-08-11">11–12 Aug.2006</collectingDate>
</date>
(
<collectionCode id="B3F3AE09235BFFC59D3F83A4D6CBFF45" box="[1376,1445,219,240]" pageId="13" pageNumber="268">QSBG</collectionCode>
, 
<specimenCode id="85449EB7235BFFC59B758387D00FFEB8" box="[810,865,248,269]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="268">T523</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="658A3C91235BFFC59B2B8387D679FEB8" box="[884,1303,248,269]" collectingDate="2006-08-12" collectingDateMax="2006-08-19" collectingDateMin="2006-08-12" collectionCode="QSBG" country="Thailand" pageId="13" pageNumber="268" specimenCode="T528" specimenCount="1" specimenCount-female="1" stateProvince="Loei">
<specimenCount id="C3E4FD45235BFFC59B2B8387D0A2FEB8" box="[884,972,248,269]" pageId="13" pageNumber="268" type="female">1 female</specimenCount>
, 
<date id="A15C100C235BFFC59B878387D715FEB8" box="[984,1147,248,269]" pageId="13" pageNumber="268" value="2006-08-12" valueMax="2006-08-19" valueMin="2006-08-12">
<collectingDate id="B118E9E4235BFFC59B878387D715FEB8" box="[984,1147,248,269]" pageId="13" pageNumber="268" value="2006-08-12" valueMax="2006-08-19" valueMin="2006-08-12">12–19 Aug.2006</collectingDate>
</date>
(
<collectionCode id="B3F3AE09235BFFC59CD58387D7A0FEB8" box="[1162,1230,248,269]" pageId="13" pageNumber="268">QSBG</collectionCode>
, 
<specimenCode id="85449EB7235BFFC59C858387D67FFEB8" box="[1242,1297,248,269]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="268">T528</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="658A3C91235BFFC59D7B8387D712FE9C" collectingDate="2006-08-26" collectingDateMax="2006-09-02" collectingDateMin="2006-08-26" collectionCode="NMWC" country="Thailand" pageId="13" pageNumber="268" specimenCode="T534" specimenCount="3" specimenCount-female="3" stateProvince="Loei">
<specimenCount id="C3E4FD45235BFFC59D7B8387D6EBFEB8" box="[1316,1413,248,269]" pageId="13" pageNumber="268" type="female">3 females</specimenCount>
, 
<date id="A15C100C235BFFC59DCE8387D0BDFE9C" pageId="13" pageNumber="268" value="2006-08-26" valueMax="2006-09-02" valueMin="2006-08-26">
<collectingDate id="B118E9E4235BFFC59DCE8387D0BDFE9C" pageId="13" pageNumber="268" value="2006-08-26" valueMax="2006-09-02" valueMin="2006-08-26">26 Aug.–2 Sep.2006</collectingDate>
</date>
(
<collectionCode id="B3F3AE09235BFFC59BBE826BD75CFE9C" box="[993,1074,276,297]" country="United Kingdom" httpUri="http://grbio.org/cool/e7dr-nxad" name="National Museum of Wales" pageId="13" pageNumber="268" type="Museum">NMWC</collectionCode>
, 
<specimenCode id="85449EB7235BFFC59C61826BD71BFE9C" box="[1086,1141,276,297]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="268">T534</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9DF86547235BFFC59B758230D0FEFE13" pageId="13" pageNumber="268" type="diagnosis">
<paragraph id="D55D36CC235BFFC59B758230D0FEFE13" blockId="13.[810,1449,335,422]" pageId="13" pageNumber="268">
<emphasis id="E796EADE235BFFC59B758230D0D3FED2" bold="true" box="[810,957,335,359]" italics="true" pageId="13" pageNumber="268">Diagnosis. –</emphasis>
A yellowish species with black head and distinctively a strongly marked dark subterminal band on the wing.
</paragraph>
</subSubSection>
<subSubSection id="9DF86547235BFFC59B7582B0D791F793" pageId="13" pageNumber="268" type="description">
<paragraph id="D55D36CC235BFFC59B7582B0D662FD13" blockId="13.[810,1449,463,678]" pageId="13" pageNumber="268">
<emphasis id="E796EADE235BFFC59B7582B0D765FE52" bold="true" box="[810,1035,463,487]" pageId="13" pageNumber="268">
<emphasis id="E796EADE235BFFC59B7582B0D0AAFE52" bold="true" box="[810,964,463,487]" italics="true" pageId="13" pageNumber="268">Description. –</emphasis>
Male.
</emphasis>
Body length 
<quantity id="121A9B29235BFFC59CC482B0D79EFE53" box="[1179,1264,463,486]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="268" unit="mm" value="2.5">2.5 mm</quantity>
. Head black with pale dust. All setae yellow or whitish; ocl and vtl strong, upo and lpo becoming progressively weaker ventrally; patch of pale setae behind mouth conspicuous. Basal antennal segments yellow; poped yellowish basally, darker apically, almost 3× long as wide; stylus black, thickened, 2× long as poped. Mouthparts whitish, proboscis black.
</paragraph>
<paragraph id="D55D36CC235BFFC59B7581B0D604FCF3" blockId="13.[810,1449,719,838]" pageId="13" pageNumber="268">Thorax yellow, scutellum and mediotergite brownish. Setae yellowish; two dc (including one level with npl) upper npl, sa and sct all strong; otherwise only minute hairs posteriorly on scutum, pospronotum and lower notopleural area.</paragraph>
<paragraph id="D55D36CC235BFFC59B758010D643FB33" blockId="13.[810,1449,879,1158]" pageId="13" pageNumber="268">
Legs yellow, tarsomeres 4–5 darker, T 
<subScript id="49663489235BFFC59CAC8003D795FC3F" attach="left" box="[1267,1275,892,906]" fontSize="6" pageId="13" pageNumber="268">1</subScript>
narrowly dark brown posteriorly. C 
<subScript id="49663489235BFFC59C5480E3D77DFC1F" attach="left" box="[1035,1043,924,938]" fontSize="6" pageId="13" pageNumber="268">1</subScript>
0.8–0.85× as long as thorax, slightly inflated, 6.1–6.2× as long as wide, widest 0.25 from base. F 
<subScript id="49663489235BFFC59B6680A3D02FFC5F" attach="left" box="[825,833,988,1002]" fontSize="6" pageId="13" pageNumber="268">1</subScript>
slightly longer than C 
<subScript id="49663489235BFFC59C7180A3D758FC5F" attach="left" box="[1070,1078,988,1002]" fontSize="6" pageId="13" pageNumber="268">1</subScript>
, distinctly inflated, 3.7× as long as wide, widest 0.3 from base; femoral formula 7(5–8)/17(15– 19)/13(12–15)/8(8–11) +1–2 (
<tableCitation id="98600377235BFFC59CD58770D78AFB93" box="[1162,1252,1039,1062]" captionStart="Table 1" captionStartId="3.[110,165,602,623]" captionText="Table 1. Femoral formula of Chelipoda species having four rows of setae on front femur. The median number of setae in each series is followed by the range (in parentheses) and standard deviation [in brackets]; n = number of femora sampled. Standard deviation was not calculated for values of n less than 6. Abbreviations: M, male; F, female; u., unknown. Species Sex av spines av denticles pv denticles pv spines basals n Chelipoda menglunana M 7(5–8) [0.90] 17(15–19) [1.12] 13(12–15) [1.07] 8(8–11) [1.01] 1–2 14" pageId="13" pageNumber="268">Table 1</tableCitation>
), spines yellow, denticles black; 
<emphasis id="E796EADE235BFFC59BB6874FD76DFBF2" box="[1001,1027,1072,1095]" italics="true" pageId="13" pageNumber="268">pv</emphasis>
spines usually rather close together basally; basal spine(s) rather weak. Mid and posterior femora and tibiae slender, lacking strong setae.
</paragraph>
<paragraph id="D55D36CC235BFFC59B7587D0D7CAFAB3" blockId="13.[810,1449,1199,1286]" pageId="13" pageNumber="268">
Abdomen brownish yellow; genitalia (
<figureCitation id="4DD92A49235BFFC59C9587D0D676FB73" box="[1226,1304,1199,1222]" captionStart="Figs" captionStartId="10.[133,177,2057,2078]" captionTargetBox="[138,1469,1322,2026]" captionTargetId="figure-508@10.[123,1483,1306,2043]" captionTargetPageId="10" captionText="Figs. 12–13. Male genitalia of Chelipoda species in lateral view: 12. C. nakladam new species; 13. C. menglunana Grootaert, Yang &amp; Saigusa. Abbreviations: lcer, lower lobe of cercus; subep, subepandrial process; ucer, upper lobe of cercus." figureDoi="http://doi.org/10.5281/zenodo.5342012" httpUri="https://zenodo.org/record/5342012/files/figure.png" pageId="13" pageNumber="268">Fig. 13</figureCitation>
) yellow with only apex of phallus and sheath blackish yellow and extreme apex of subepandrial process dark.
</paragraph>
<paragraph id="D55D36CC235BFFC59B758650D776F973" blockId="13.[810,1449,1327,1734]" pageId="13" pageNumber="268">Epan and Hypan fused, with lateral lobes completely fused posteroapically but slightly separated posteroventrally, with scattered short setae. Cercus fused with Epan+Hypan, consisting of two roughly triangular parts connected by a narrow intermediate part along dorsum of Epan+Hypan; posterior part narrowly triangular with rather erect short setulae dorsally; anterior part free apically and bearing 2-3 stout black setae apically. Subepandrial process, erect, narrow with apical hook. Phallus strongly reflexed forward, rather broad but abruptly narrowed distally and terminating just anterior of anterior lobe of cercus; phallic sheath also broad and continuing forward beyond apex of phallus for a considerable distance.</paragraph>
<paragraph id="D55D36CC235BFFC59B758590D620F8D3" blockId="13.[810,1450,1775,1894]" pageId="13" pageNumber="268">Wing veins yellowish basally, darker distally; membrane faintly brownish yellow, a strong black subterminal band commencing at apex of cell dm leaving apex of wing clear. Squamae with pale fringes. Halter whitish yellow.</paragraph>
<paragraph id="D55D36CC235BFFC59B7584F0D623F813" blockId="13.[810,1357,1935,1959]" box="[810,1357,1935,1959]" pageId="13" pageNumber="268">
<emphasis id="E796EADE235BFFC59B7584F0D0EAF812" bold="true" box="[810,900,1935,1959]" pageId="13" pageNumber="268">Female.</emphasis>
Antenna with poped darker than in male.
</paragraph>
<paragraph id="D55D36CC235BFFC59B7584B0D791F793" blockId="13.[810,1449,1999,2086]" pageId="13" pageNumber="268">
Femoral formula (female) 5(5–6)/19(15–21)/17(14– 22)/5(4–6) +1–2, 
<emphasis id="E796EADE235BFFC59BA9848FD77EF7B2" box="[1014,1040,2032,2055]" italics="true" pageId="13" pageNumber="268">pv</emphasis>
spines usually more evenly spaced basally. Cercus moderately long, yellowish
</paragraph>
</subSubSection>
<subSubSection id="9DF865472358FFC698DA83C1D257FEE0" pageId="14" pageNumber="269" type="discussion">
<paragraph id="D55D36CC2358FFC698DA83C1D257FEE0" blockId="14.[133,772,190,341]" pageId="14" pageNumber="269">
<emphasis id="E796EADE2358FFC698DA83C1D263FF63" bold="true" box="[133,269,190,214]" italics="true" pageId="14" pageNumber="269">Remarks. –</emphasis>
Described originally from 
<collectingRegion id="1726F82E2358FFC69A0A83C1D1DFFF60" box="[597,689,190,213]" country="China" name="Yunnan" pageId="14" pageNumber="269">Yunnan</collectingRegion>
, 
<collectingCountry id="ADF5765C2358FFC69A9E83C1D06DFF60" box="[705,771,190,213]" name="China" pageId="14" pageNumber="269">China</collectingCountry>
(
<bibRefCitation id="B1734B3D2358FFC698D283A1D2E6FF40" author="Grootaert, P &amp; Yang, D &amp; Saigusa, T" box="[141,392,222,245]" pageId="14" pageNumber="269" pagination="71 - 80" refId="ref20704" refString="Grootaert, P., Yang, D. &amp; Saigusa, T., 2000. Empididae (Diptera: Empidoidea) from Xishuangbanna, Yunnan (I): Hemerodromiinae. Bulletin de L'Institute Royal des Sciences Naturelles de Belgique, Entomologie, 70: 71 - 80." type="journal article" year="2000">Grootaert et al., 2000</bibRefCitation>
), 
<taxonomicName id="12E24D4F2358FFC699C183A0D126FF43" authorityName="Grootaert, Yang &amp; Saigusa" authorityYear="2000" box="[414,584,223,246]" class="Insecta" family="Empididae" genus="Chelipoda" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="menglunana">
<emphasis id="E796EADE2358FFC699C183A0D126FF43" box="[414,584,223,246]" italics="true" pageId="14" pageNumber="269">C. menglunana</emphasis>
</taxonomicName>
is here reported from 
<collectingRegion id="1726F82E2358FFC6989E8381D234FEA0" box="[193,346,254,277]" country="Thailand" name="Loei" pageId="14" pageNumber="269">Loei Province</collectingRegion>
in north eastern 
<collectingCountry id="ADF5765C2358FFC69A4C8381D11BFEA0" box="[531,629,254,277]" name="Thailand" pageId="14" pageNumber="269">Thailand</collectingCountry>
. Adults were collected for a short period of the wet season in August at around 
<quantity id="121A9B292358FFC698878241D25BFEE0" box="[216,309,318,341]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="269" unit="m" value="1200.0">1,200 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>