The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level Author Rix, Michael Western Australian Museum, Welshpool DC, Perth, Australia Author Harvey, Mark Western Australian Museum, Perth, Welshpool, Australia text ZooKeys 2010 2010-02-22 36 36 1 321 journal article 22759 10.3897/zookeys.36.306 4db6b327-7482-432e-a5f6-36f91c79fef3 1313–2970 576620 ADCACC88-6C78-4386-8E33-3F98234ECE92 Raveniella peckorum Rix & Harvey , sp. n. urn:lsid:zoobank.org:act: 28D43BF3-89EF-407A-B6B4-31BE17F5F7CE Figs 75D , 76 , 77B , 78 D–F Type material. Holotype male: Modong Nature Reserve , 15 km NE. of Rockingham , Western Australia , Australia , sifting leaf litter, 32°13'10"S , 115°54'09"E , 5.vi.2007 , M. Rix ( WAM T94549). Paratypes : Allotype female, same data as holotype ( WAM T94550); 10 males and 5 females , same data as holotype ( WAM T94408 SEM♁ ) . Other material examined. AUSTRALIA : Western Australia : Albany , Emu Point , sifting leaf litter, 35°00'14"S , 117°55'50"E , 30.IV.2008 , M. Rix , 1♁, 1♀ ( WAM T94366) ; Austin Bay Nature Reserve , E. of Peel Inlet , end of Beacham Road , 32°36'42"S , 115°47'11"E , 12.VI.2007 , M. Rix , 1♁, 4♀ ( WAM T94411) ; Avon National Park , off Keating Road , sifting leaf litter, 31°34'30"S , 116°09'56"E , 20.VI.2007 , M. Rix , 1♁, 1♀ ( WAM T94436) ; Beela Road , 7.5 km ENE. of Brunswick Junction , Tullgren funnel , sifted leaf litter, 33°14'14"S , 115°55'00"E , 25.VII.2003 , M. Harvey , 1♀ ( WAM T76276) ; same data, 2♀ ( WAM T76277) ; same data, 2♀ ( WAM T76275) ; Boddington Bauxite Mine , 32°57'S , 116°27'E , VII.2003 , G. Oraby , 1♀ ( WAM T76204) ; same data, 1♀ ( WAM T76213) ; same data except X.2007 , 1♀ ( WAM T76207) ; same data, 1♀ ( WAM T76203) ; Bold Park , Perth , site BP1, wet pitfall , 31°57'11"S , 115°45'50"E , 24.IX.-18.XI.1993 , J. Waldock et al., 1♀ ( WAM T76301) ; same data except 20.V.-20.VII.1993 , M. Harvey , J. Waldock , 1♀ ( WAM T76302) ; Bold Park , Perth , site BP3, wet pitfall , 31°56'30"S , 115°46'27"E , 20.VII.-24.IX.1993 , J. Waldock , 1♀ ( WAM T76306) ; Bold Park , Perth , site BP4, wet pitfall , 31°56'29"S , 115°46'01"E , 24.IX.-18.XI.1993 , J. Waldock et al., 1♀ ( WAM T76305) ; Bold Park , Perth, W. of Challenger Parade , sifting leaf litter, 31°57'06"S , 115°45'26"E , 28.V.2007 , M. Rix , 1♁, 4♀ ( WAM T94374) ; same data except 14.IX.2008 , 5♀ ( WAM T94363) ; same data except 15.V.2007 , 2♁ ( WAM T94371) ; same data except 12.V.2007 , 1♁ ( WAM T94410) ; Bibra Lake Bushland, Perth, E. of North Lake Road , sifting leaf litter, 32°05'16"S , 115°49'12"E , 17.V.2007 , M. Rix , 1♀ ( WAM T94387) ; Brockman National Park , 16 km S. of Pemberton , leaf litter and mould under karri trees, 8. XII.1976 , J. Kethley , 1♀ ( FMC ) ; Buller Nature Reserve , 9.5 km SW. of Waroona , sift- ing leaf litter, 32°52'04"S , 115°49'43"E , 22.VI.2007 , M. Rix , 2♁, 1♀ ( WAM T94431) ; Cup Road , 40 km SE. of Manjimup , 6–28.VII.1980 , S. & J. Peck , 6♁, 9♀ ( WAM T76232) ; same data except 15.VII.1980 , 3♁, 13♀ ( AMNH ) ; Cardup Nature Reserve , 10 km SSW. of Armadale , sifting leaf litter, 32°14'45"S , 115°59'31"E , 5. VI.2007 , M. Rix , 2♁, 12♀ ( WAM T94417) ; same data except site CR2, wet pitfall , 32°14'40"S , 115°59'15"E , 16.IV.-17.VI.1996 , J. Waldock , P. West , A. Longbottom , 1♁ ( WAM T76200) ; Conspicuous Point , litter under peppermint trees, 14.XII.1976 , J. Kethley , 1♀ ( FMC ) ; Dog Pool , on Shannon River , leaf litter, 32°46'S , 116°22'E , 27.IV.-1.V.1990 , M. Harvey , J. Waldock , 1♀ ( WAM T76260) ; Hartfield Park , Forrestfield , Perth, E. of Tonkin Highway , sifting leaf litter, 31°59'53"S , 115°59'49"E , 23.V.2007 , M. Rix , 2♁, 2♀ ( WAM T94399) ; Huntly Mine , 5 km E. of Banksiadale Dam , bark trapped, 32°39'S , 116°05'E , IV.2005 , A. Peck , 1♀ ( WAM T63388) ; same data except 2.VI.-14.VII.2005 , 1♀ ( WAM T63858) ; same data, 1♀ ( WAM T63904) ; same data, 1♀ ( WAM T63861) ; same data, 1♁ ( WAM T63871) ; same data except 13.VII.-24.VIII.2005 , 1♀ ( WAM T64282) ; Jandakot Airport , Perth , site JK1, wet pitfall , 32°05'36"S , 115°52'39"E , 16.VII.-1.IX.1994 , J. Waldock , A. Longbottom , 4♁, 1♀ ( WAM T76293) ; Jandakot Reserve , Perth , off Lakes Way, W. of Jandakot Airport , sifting leaf litter, 32°05'40"S , 115°51'24"E , 6.VI.2007 , M. Rix , 2♁, 1♀ ( WAM T94420) ; Kings Park , Perth , sifting leaf litter, 31°57'19"S , 115°50'00"E , 7. VI.2007 , M. Rix , 4♁, 10♀ ( WAM T94370) ; same data except Berlese litter, 12. VI.1987 , N. Platnick , R. Raven , 27♁, 25♀ ( AMNH ) ; Koondoola Regional Bushland , Perth, S. of Marangaroo Drive , sifting leaf litter, 31°50'01"S , 115°52'13"E , 22.V.2007 , M. Rix , 1♀ ( WAM T94401) ; Leeuwin-Naturaliste National Park , Augusta , 34°20'S , 115°09'E , 22.VII.1980 , S. & J. Peck , 1♁, 1♀ ( WAM T76238) ; same data, 4♁, 12♀ ( AMNH ) ; Leeuwin-Naturaliste National Park , N. of Caves Road , sifting leaf litter, 33°38'39"S , 115°02'44"E , 21.VI.2007 , M. Rix , 3♀ ( WAM T94382) ; Leeuwin-Naturaliste National Park , off Sugarload Road , sifting leaf litter, 33°33'29"S , 115°01'25"E , 25.IV.2008 , M. Rix , 1♀ ( WAM T94379) ; Leeuwin-Naturaliste National Park , Yallingup , marri wood litter, 21.VII.1980 , S. & J. Peck , 6♁, 27♀ ( AMNH ) ; Lightning Swamp Bushland , Perth, N. of Malaga Drive , sifting leaf litter, 31°52'11"S , 115°54'07"E , 22.V.2007 , M. Rix , 1♁, 2♀ ( WAM T94390) ; Mammoth Cave Reserve , karri forest dung traps , 34°04'S , 115°02'E , 20–22.VII.1980 , S. & J. Peck , 2♀ ( WAM T76267) ; same data, 2♀ ( AMNH ) ; 54 km SE. of Manjimup , jarrah forest litter, 34°36'S , 116°34'E , 22.VI–4.VII.1980 , S. & J. Peck , 5♀ ( WAM T76231) ; same data, 1♁, 3♀ ( AMNH ) ; Marangaroo Reserve , Perth , N. of Horton Crescent , sifting leaf litter, 31°49'52"S , 115°50'12"E , 21.V.2007 , M. Rix , 7♁, 4♀ ( WAM T94402) ; S. of Mordalup Road, N. of Lake Muir , wet pitfall , 34°19'01"S , 116°31'49"E , 15.X.1999 1.XI.2000 , P. Van Heurck et al. CALM survey, 1♁ ( WAM T94175) ; Mount Cooke , leaf litter, 32°25'S , 116°18'E , 19.IX.1991 , M. Harvey , J. Waldock , 2♀ ( WAM T76222) ; same data except 27.IV.1992 , 8♁, 4♀ ( WAM T76223) ; same data, 5♀ ( WAM T76268) ; same data except 31.VII.1991 , 3♁, 9♀ ( WAM T76224) ; same data except 1.X.1990 , 1♀ ( WAM T76230) ; same data except pitfall trap, 16.VI.-31. VII.1991 , 1♁, 1♀ ( WAM T76227) ; same data except sifting leaf litter, 20.VII.2002 , M. Rix , M. Harvey , 3♁, 4♀ ( WAM T76212) ; Mount Frankland , 29 km N. of Walpole , unburned litter in burned area, J. Kethley , 1♀ ( FMC ) ; Mount Henry , Perth , E. of Kwinana Freeway , sifting leaf litter, 32°01'57"S , 115°51'39"E , 20.V.2007 , M. Rix , 1♁ ( WAM T94384) ; Mount Lindesay , site 208, 34°50'S , 117°19'E , VIII.1995 , S. Barrett , 1♁ ( WAM T76216) ; Neerabup National Park , W. of Wanneroo Road , sifting leaf litter, 31°38'58"S , 115°43'44"E , 2.IX.2006 , M. Rix , 3♀ ( WAM T94385) ; Perth Airport , site PA6, wet pitfall , 31°58'05"S , 115°58'05"E , 28.VII–23.IX.1993 , J. Waldock et al., 1♁ ( WAM T76299) ; Perth Airport , site PA8, wet pitfall , 31°58'36"S , 115°58'28"E , 28.VII.-23.IX.1993 , J. Waldock et al., 1♁, 1♀ ( WAM T76298) ; Perth Airport , south-eastern sector, N. of Tonkin Highway , sifting leaf litter, 31°58'36"S , 115°58'27"E , 29.V.2008 , M. Rix , J. Wojcieszek , 4♁, 4♀ ( WAM T94422) ; Pinnaroo Valley Memorial Park , Perth , wet pitfall , 31°48'S , 115°46'E , 1.VIII.1993 , D. Leary , 1♁ ( WAM T76274) ; Porongurup National Park , site 226, wet pitfall , 34°40'56"S , 117°52'30"E , 13.VI.1996 , S. Barrett , 1♁ ( WAM T76195) ; Porongurup National Park , Ravine Road , dry moss on seepage area, 34°41'S , 117°53'E , 24.XII.1976 , J. Kethley , 1♀ ( FMC ) ; Porongurup National Park , southern slope of Twin Peaks , 34°41'S , 117°53'E , 2.I.1977 , J. Kethley , 5♀ ( FMC ) ; same data, 1♀ ( FMC ) ; Port Kennedy Scientific Park , S. of Port Kennedy Drive , sifting leaf litter, 32°22'41"S , 115°45'02"E , 12.VI.2007 , M. Rix , 2♁, 3♀ ( WAM T94415) ; near Quinninup , 29 km SSE. of Manjimup , karri litter, 34°29'S , 116°16'E , 13–16.VII.1980 , S. & J. Peck , 2♀ ( WAM T76262) ; Rottnest Island , near Lake Timperley , sifting leaf litter, 32°00'23"S , 115°31'11"E , 13.VI.2007 , M. Rix , 9♁, 14♀ ( WAM T94377) ; Shenton Park Bushland , Perth, S. of Lemnos Street , sifting leaf litter, 31°57'37"S , 115°48'01"E , 10.V.2007 , M. Rix , 1♁ ( WAM T94380) ; Stirling Range National Park , base of Pyungoorup Peak , sifting leaf litter, 34°21'54"S , 118°19'44"E , 5.VIII.2008 , M. Rix , M. Harvey , 9♁, 10♀ ( WAM T94362) ; same data except wet pitfall , 27.IV.-4.IX.1996 , M. Harvey et al., 2♁, 1♀ ( WAM T76282) ; Stirling Range National Park , Bluff Knoll , site 230, pitfall , 34°23'S , 118°15'E , 7.IX.1995 , S. Barrett , 1♁ ( WAM T76190) ; Stirling Range National Park , Bluff Knoll , site 213, pitfall , 34°23'37"S , 118°15'10"E , 19.VI.1996 , S. Barrett , 1♁ ( WAM T76189) ; Stirling Range National Park , Bluff Knoll , 34°22'S , 118°15'E , 23.XII.1976 , J. Kethley , 2♀ ( FMC ) ; same data, 1♀ ( FMC ) ; same data except 31.XII.1976 , 1♀ ( FMC ) ; Stirling Range National Park , Ellen Peak , sifting leaf litter, 34°21'30"S , 118°19'57"E , 6.XI.2007 , M. Rix et al., 1♀ ( WAM T94368) ; Stirling Range National Park , Talyuberlup Peak , 34°24'S , 117°57'E , 29.XII.1976 , J. Kethley , 1♀ ( FMC ) ; Stirling Range National Park , Toolbrunup Peak , wet pitfall , 34°23'32"S , 118°03'32"E , 23.IV.-3.IX.1996 , J. Waldock , B. Main , 1♁ ( WAM T76281) ; Stirling Range National Park , Toolbrunup Peak track, 34°22'S , 118°02'E , 10.VI.1980 , S. & J. Peck , 10♁ ( AMNH ) ; Stirling Range National Park , Toolbrunup Peak , 34°22'S , 118°02'E , 27.XII.1976 , J. Kethley , 1♀ ( FMC ) ; same data, 1♀ ( FMC ) ; Sues Bridge , 50 km SW. of Nannup , marri log litter, 34°18'S , 115°22'E , 26.VII.1980 , S. & J. Peck , 3♀ ( WAM T76233) ; same data, 1♁, 3♀ ( WAM T76234) ; same data, 2♁, 12♀ ( AMNH ) ; Sultana Road Bushland , Perth, W. of Roe Highway , sifting leaf litter, 31°57'55"S , 116°00'11"E , 23.V.2007 , M. Rix , 3♁, 3♀ ( WAM T94398) ; Talbot Road Reserve , Stratton , Perth , sifting leaf litter, 31°52'04"S , 116°03'02"E , 31.V.2007 , M. Rix , 8♁, 8♀ ( WAM T94396) ; same data except marri litter Tullgren funnel, 29.VIII.2006 , J. Waldock , K. Edward , 1♁ ( WAM T94533) ; Talbot Road Reserve , Stratton , Perth , site TR3, wet pitfall , 31°52'24"S , 116°02'52"E , 24.VI.-28.VII.1993 , M. Harvey , J. Waldock , 1♁ ( WAM T94565) ; Thomsons Lake Nature Reserve , Perth , S. of Senecio Lane , sifting leaf litter, 32°08'14"S , 115°49'28"E , 6.VI.2007 , M. Rix , 2♀ ( WAM T94414) ; Torbay Head , lot 40, pitfall , 35°08'S , 117°38'E , 14–27.VI.1987 , B. Main , 1♁ ( WAM T76271) ; Torndirrup National Park , pitfall , 25.V.-9.XI.1983 , P. Dyer , J. Lyon , 11♁, 9♀ ( WAM T76221) ; same data except 6–13.VII.1983 , 1♀ ( WAM T76235) ; Torndirrup National Park , Eclipse Island Road, Tullgren funnel , 35°06'05"S , 117°52'23"E , 15.III.2005 , M. Harvey et al., 3♀ ( WAM T76236) ; Torndirrup National Park , Flinders Peninsula near Isthmus Bay , 35°06'S , 117°58'E , 28.XII.1976 , J. Kethley , 1♀ ( FMC ) ; Trigg Bushland Reserve north, Perth, W. of Duart Road , sifting leaf litter, 31°52'23"S , 115°45'44"E , 15.V.2007 , M. Rix , 1♀ ( WAM T94400) ; Trigg Dune Bush , Perth , site TD1, wet pitfall , 31°52'09"S , 115°45'38"E , 13.VII.-25.IX.1995 , M. Harvey , J. Waldock , 1♀ ( WAM T76217) ; Tuart Forest National Park , near junction of Stirling Road and Higgins Road , sifting leaf litter, 33°33'07"S , 115°30'48"E , 21.VI.2007 , M. Rix , 4♀ ( WAM T94427) ; Two People’s Bay Nature Reserve , 34°59'S , 118°10'E , 14. VI.1980 , S. & J. Peck , 6♀ ( AMNH ) ; Two People’s Bay Nature Reserve , Mount Gardner , 35°00'S , 118°11'E , 1.I.1977 , J. Kethley , 1♀ ( FMC ) ; same data, 1♀ ( FMC ) ; same data, 1♀ ( FMC ) ; Walpole-Nornalup National Park , 28 km NE. of Walpole , 34°58'S , 116°55'E , 17.XII.1976 , J. Kethley , 1♀ ( FMC ) ; Walpole-Nornalup National Park , Collier Road , 19.VI.1980 , S. & J. Peck , 2♁, 1♀ ( AMNH ) ; Walpole-Nornalup National Park , Hilltop Road , 21.VI.1980 , S. & J. Peck , 4♁, 8♀ ( AMNH ) ; Walpole- Nornalup National Park , Shedley Drive , 34°59'S , 116°44'E , 22.VI.1980 , S. & J. Peck , 1♀ ( WAM T76263) ; Walpole-Nornalup National Park , Shedley Drive , 35°00'S , 116°38'E , 22.VI.1980 , S. & J. Peck , 2♀ ( AMNH ) ; Walpole-Nornalup National Park , Tingle Tree , 34°59'S , 116°47'E , 18.VI.-29.VII.1980 , S. & J. Peck , 6♁, 5♀ ( AMNH ) ; same data, 1♁ ( AMNH ) ; Walpole-Nornalup National Park , Valley of the Giants , 35°00'S , 116°43'E , VII.1980 , S. & J. Peck , 2♀ ( WAM T76240) ; same data, 1♁, 6♀ ( WAM T76239) ; Walpole-Nornalup National Park , Valley of the Giants , 34°58'S , 116°53'E , 21.VII.1980 , S. & J. Peck , 3♁, 4♀ ( AMNH ) ; Walpole-Nornalup National Park , Zig Zag Road , 34°58'S , 116°53'E , 20.VI.-4.VII.1980 , S. & J. Peck , 6♁, 18♀ ( AMNH ) ; Warren National Park , near Pemberton , 34°25'S , 115°56'E , 12.VII.1980 , S. & J. Peck , 2♀ ( WAM T17744 ) ; Warren National Park , near Pemberton , 34°30'S , 115°57'E , 5.VII.1980 , S. & J. Peck , 1♁, 1♀ ( AMNH ) ; Wellesley River at Marriott Road , Tullgren funnel , 33°12'40"S , 115°47'02"E , 25.VII.2003 , M. Harvey , 2♀ ( WAM T76269) ; Wellington National Park , off Wellington Dam Road , sifting leaf litter, 33°20'28"S , 115°57'27"E , 22.VI.2007 , M. Rix , 3♀ ( WAM T94433) ; Windy Harbour , 34°50'S , 116°01'E , 8–12.VII.1980 , S. & J. Peck , 1♁ ( WAM T76261) ; same data except 8.VII.1980 , 9♁, 9♀ ( AMNH ) ; Wireless Hill Bushland , Perth, N. of Davy Street , sifting leaf litter, 32°01'57"S , 115°49'34"E , 16.V.2007 , M. Rix , 1♁ ( WAM T94391) ; Woodman Point Regional Park, S. of Perth , sifting leaf litter, 32°08'S , 115°45'E , 6.VIII.2006 , M. Rix , J. Wojcieszek , 1♁, 7♀ ( WAM T94413) ; same data, 1♁ ( WAM T80033 DNA-MPE ) ; same data except 21.V.2005 , M. Rix , M. Harvey , 5♁, 3♀ ( WAM T76279) ; Woodman Point Regional Park, S. of Perth , sifting leaf litter, 32°07'55"S , 115°45'35"E , 30.V.2007 , M. Rix , 8♁, 5♀ ( WAM T94394) ; Woodman Point Regional Park, S. of Perth , site WO4, wet pitfall , 32°07'58"S , 115°45'29"E , 4.V.-6.VII.1995 , J. Waldock , M. Harvey , 3♁ ( WAM T76284) ; same data except 1. IX.-4.XI.1994 , 2♁ ( WAM T76182) ; same data except 28.VI.-1.IX.1994 , J. Waldock , A. Longbottom , 10♁, 2♀ ( WAM T76285) ; same data, 1♁ ( WAM T76173) ; Yalgorup National Park , adjacent to White Hill Road , sifting leaf litter, 32°41'15"S , 115°38'57"E , 27.VI.2007 , M. Rix , 1♁, 2♀ ( WAM T94430) ; Yalgorup National Park , off Preston Beach Road , W. of Lake Newnham , sifting leaf litter, 32°53'53"S , 115°41'22"E , 27. VI.2007 , M. Rix , 2♁, 2♀ ( WAM T94437) . Etymology . The specific epithet is a patronym in honour of Stewart and Jarmila Peck, who collected many specimens of this (and other micropholcommatid) species in 1980, during AMNH-sponsored field work in south-western Western Australia . Diagnosis . Males of Raveniella peckorum can be distinguished from R. hickmani by the longer, distally-curled embolus ( Fig. 77 ), and from R. luteola by the absence of a terminal, spur-like extension on the evaginated tegular ridge ( Fig. 77B ). Females can be distinguished from all other described congeners by the cylindrical receptacula and spherical morphology of the spermathecae ( Fig. 75D ). Description . Holotype male : Total length 1.16. Carapace 0.59 long, 0.45 wide. Abdomen 0.63 long, 0.51 wide. Leg I femur 0.45. Cephalothorax, legs dark tan-yellow; abdomen dark olive-grey. Carapace raised anteriorly, not fused to sternum except around petiole; dorsal surface of pars cephalica slightly convex in lateral view. Eight eyes present on anterior margin of pars cephalica; AME 0.6x diameter of ALE; PME separated by their own diameter. Chelicerae each with large, bulging anterior projection; promargin with three peg teeth. Legs relatively short (leg I femur-carapace ratio 0.76); distal, prolateral macroseta present on tibia I. Abdomen oval, covered with hairlike setae; dorsal scute and lateral sclerotic strips absent. Pedipalpal patella with retrolaterally-directed, hooked lRPA, small, conical distal apophysis, larger, subtriangular terminal apophysis and ornate, ridged cuticular microstructure; tegulum smooth, with curved, marginal ETR; embolus very long (length >> 5× width), extending posterior to patella, curling distally to form a three-quarter revolution; cymbium unmodified with three long posterior macrosetae ( Fig. 77B ). Allotype female : Total length 1.22. Carapace 0.60 long, 0.43 wide. Abdomen 0.74 long, 0.59 wide. Leg I femur 0.45. Cephalothorax, legs tan-yellow; abdomen tan-grey. Carapace raised anteriorly, not fused to sternum except around petiole; dorsal surface of pars cephalica slightly convex in lateral view. Eight eyes present on anterior margin of pars cephalica; AME 0.6x diameter of ALE; PME separated by slightly less than their own diameter. Chelicerae without bulging anterior projections; promargin without peg teeth. Legs relatively short (leg I femur-carapace ratio 0.75); macrosetae absent. Abdomen oval, covered with hair-like setae; dorsal scute and lateral sclerotic strips absent. Pedipalp entire, five-segmented. Epigyne with distinctive external morphology ( Fig. 76D ); receptacula cylindrical, with spherical spermathecae; insemination ducts membranous, with complex, convoluted trajectory and multiply-looped distal portion; fertilisation ducts filiform ( Fig. 75D ). Distribution . This species is found throughout the south-western Australian ‘High Rainfall Province’ (see Hopper and Gioia 2004 ), from Two Peoples Bay Nature Reserve near Albany to the Avon National Park, north of Perth, with outlying populations in the Porongurup and Stirling Range National Parks. It is ubiquitous in the tall eucalypt forests of the Darling Escarpment and south coast, and is also found throughout the southern Swan Coastal Plain (including Rottnest Island), north to Neerabup National Park. Remarks . Raveniella peckorum is one of several species of Raveniella known from south-western Western Australia (M. Rix, unpubl. data). It was included in the molecular phylogenetic analysis of Rix et al. (2008) as “ Textricella sp. WA”.