<document id="3FDFB6D01C6EF9F1CC77E81D6963744D" ID-CLB-Dataset="22759" ID-DOI="10.3897/zookeys.36.306" ID-GBIF-Dataset="4db6b327-7482-432e-a5f6-36f91c79fef3" ID-ISSN="1313–2970" ID-Zenodo-Dep="576620" ID-ZooBank="ADCACC88-6C78-4386-8E33-3F98234ECE92" IM.materialsCitations_requiresApprovalFor="plazi" IM.metadata_requiresApprovalFor="plazi" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1587967053581" checkinUser="plazi" docAuthor="Rix, Michael &amp; Harvey, Mark" docDate="2010" docId="7E13878EFFAD1B67FF321ACEFC0E8800" docLanguage="en" docName="ZK_article_2154.pdf" docOrigin="ZooKeys 36 (36)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@6ed9075d" docStyleName="zookeys.2008.journal_article" docTitle="Pua Forster 1959" docType="treatment" docVersion="8" lastPageNumber="29" masterDocId="822AFFF6FFB61B7BFFBE1F57FFE98D05" masterDocTitle="The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level" masterLastPageNumber="321" masterPageNumber="1" pageNumber="28" updateTime="1732821582117" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title id="6C0C87E8142C4EE4A6CE61ABFB2108C3">The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level</mods:title>
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<mods:namePart id="01FED46DCE1CF0CC21D6DF9613A834FC">Rix, Michael</mods:namePart>
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<mods:namePart id="698CDD97D0E6CAA2C95B55C220BBFBF2">Harvey, Mark</mods:namePart>
<mods:affiliation id="808152ABB920626D5071D77E355AD617">Western Australian Museum, Perth, Welshpool, Australia</mods:affiliation>
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<treatment id="7E13878EFFAD1B67FF321ACEFC0E8800" ID-DOI="http://doi.org/10.5281/zenodo.3789432" ID-GBIF-Taxon="163749013" ID-Zenodo-Dep="3789432" LSID="urn:lsid:plazi:treatment:7E13878EFFAD1B67FF321ACEFC0E8800" httpUri="http://treatment.plazi.org/id/7E13878EFFAD1B67FF321ACEFC0E8800" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" scope_class="Arachnida" scope_family="Micropholcommatidae" scope_order="Araneae" scope_subFamily="Micropholcommatinae" scope_tribe="Micropholcommatini">
<subSubSection id="BEA06513FFAD1B60FF321ACEFE5D88B6" box="[140,436,1433,1460]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="F6053698FFAD1B60FF321ACEFE5D88B6" blockId="27.[140,436,1433,1460]" box="[140,436,1433,1460]" pageId="27" pageNumber="28">
<heading id="AD4D81F4FFAD1B60FF321ACEFE5D88B6" box="[140,436,1433,1460]" fontSize="11" level="2" pageId="27" pageNumber="28" reason="3">
<emphasis id="C4CEEA8AFFAD1B60FF321ACEFE5D88B6" bold="true" box="[140,436,1433,1460]" pageId="27" pageNumber="28">
Genus 
<taxonomicName id="31BA4D1BFFAD1B60FF611ACDFE5D88B6" authority="Forster, 1959" authorityName="Forster" authorityYear="1959" box="[223,436,1433,1460]" class="Arachnida" family="Micropholcommatidae" genus="Pua" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="C4CEEA8AFFAD1B60FF611ACDFEE788B1" bold="true" box="[223,270,1434,1460]" italics="true" pageId="27" pageNumber="28">Pua</emphasis>
<bibRefCitation id="922B4B69FFAD1B60FEA81ACDFE5D88B6" author="Forster RR" box="[278,436,1433,1460]" pageId="27" pageNumber="28" pagination="269 - 329" refId="ref70149" refString="Forster RR (1959) The spiders of the family Symphytognathidae. Transactions of the Royal Society of New Zealand 86: 269 - 329." type="journal article" year="1959">Forster, 1959</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BEA06513FFAD1B60FF321AB6FEB88B82" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="F6053698FFAD1B60FF321AB6FD3D8B1B" blockId="27.[140,1107,1504,1566]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="D6AA11B6FFAD1B60FF321AB6FD3D8B1B" pageId="27" pageNumber="28">
<treatmentCitation id="771B1089FFAD1B60FF321AB6FE7A88FF" author="Forster RR" box="[140,403,1504,1531]" page="299" pageId="27" pageNumber="28" year="1959">
<taxonomicName id="31BA4D1BFFAD1B60FF321AB6FE7A88FF" authority="Forster, 1959: 299" authorityName="Forster" authorityPageNumber="299" authorityYear="1959" box="[140,403,1504,1531]" class="Arachnida" family="Micropholcommatidae" genus="Pua" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="C4CEEA8AFFAD1B60FF321AB6FF5F88FF" box="[140,182,1505,1530]" italics="true" pageId="27" pageNumber="28">Pua</emphasis>
<bibRefCitation id="922B4B69FFAD1B60FF001AB6FE7A88FF" author="Forster RR" box="[190,403,1504,1531]" pageId="27" pageNumber="28" pagination="269 - 329" refId="ref70149" refString="Forster RR (1959) The spiders of the family Symphytognathidae. Transactions of the Royal Society of New Zealand 86: 269 - 329." type="journal article" year="1959">Forster, 1959: 299</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. 
<typeStatus id="2901883AFFAD1B60FE221AB6FE3A88FE" box="[412,467,1505,1531]" pageId="27" pageNumber="28">Type</typeStatus>
species by original designation 
<taxonomicName id="31BA4D1BFFAD1B60FC861AB6FEC58B1B" authority="Forster, 1959" authorityName="Forster C" authorityYear="1959" class="Arachnida" family="Micropholcommatidae" genus="Pua" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="novaezealandiae">
<emphasis id="C4CEEA8AFFAD1B60FC861AB6FBF588FF" box="[824,1052,1504,1530]" italics="true" pageId="27" pageNumber="28">Pua novaezealandiae</emphasis>
<bibRefCitation id="922B4B69FFAD1B60FB9D1AB6FEC58B1B" author="Forster RR" pageId="27" pageNumber="28" pagination="269 - 329" refId="ref70149" refString="Forster RR (1959) The spiders of the family Symphytognathidae. Transactions of the Royal Society of New Zealand 86: 269 - 329." type="journal article" year="1959">Forster, 1959</bibRefCitation>
</taxonomicName>
. 
<treatmentCitation id="771B1089FFAD1B60FE891954FDF28B1B" author="Brignoli PM" box="[311,539,1539,1566]" page="374" pageId="27" pageNumber="28" year="1983">
<bibRefCitation id="922B4B69FFAD1B60FE891954FDF28B1B" author="Brignoli PM" box="[311,539,1539,1566]" pageId="27" pageNumber="28" refId="ref69575" refString="Brignoli PM (1983) A Catalogue of the Araneae Described Between 1940 and 1981. Manchester University Press, Manchester, 755 pp." type="book" year="1983">Brignoli, 1983: 374</bibRefCitation>
</treatmentCitation>
. 
<bibRefCitation id="922B4B69FFAD1B60FD981954FD398B1B" author="Platnick NI" box="[550,720,1539,1566]" pageId="27" pageNumber="28" refId="ref71855" refString="Platnick NI (2009) The World Spider Catalog, Version 10.0. American Museum of Natural History, New York. http: // research. amnh. org / entomology / spiders / catalog / [accessed 20. X. 2009]" type="url" year="2009">Platnick, 2009</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F6053698FFAD1B60FF32191EFEB88B82" blockId="27.[140,1107,1609,1671]" pageId="27" pageNumber="28">
<emphasis id="C4CEEA8AFFAD1B60FF32191EFEE98B66" bold="true" box="[140,256,1609,1635]" pageId="27" pageNumber="28">Affinities.</emphasis>
The genus 
<taxonomicName id="31BA4D1BFFAD1B60FE32191CFE5F8B61" authorityName="Forster" authorityYear="1959" box="[396,438,1611,1636]" class="Arachnida" family="Micropholcommatidae" genus="Pua" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="C4CEEA8AFFAD1B60FE32191CFE5F8B61" box="[396,438,1611,1636]" italics="true" pageId="27" pageNumber="28">Pua</emphasis>
</taxonomicName>
appears to be the sister-lineage to 
<taxonomicName id="31BA4D1BFFAD1B60FCEA191DFBF88B61" authorityName="Crosby &amp; Bishop" authorityYear="1927" box="[852,1041,1610,1636]" class="Arachnida" family="Micropholcommatidae" genus="Micropholcomma" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="C4CEEA8AFFAD1B60FCEA191DFBF88B61" box="[852,1041,1610,1636]" italics="true" pageId="27" pageNumber="28">Micropholcomma</emphasis>
</taxonomicName>
from 
<collectingCountry id="8EAD7608FFAD1B60FF32193AFF068B82" box="[140,239,1645,1671]" name="Australia" pageId="27" pageNumber="28">Australia</collectingCountry>
(
<figureCitation id="6E812A1DFFAD1B60FF41193AFEAA8B82" box="[255,323,1645,1671]" captionStart="Figure 4" captionStartId="137.[140,212,1307,1329]" captionTargetBox="[161,1087,180,1278]" captionTargetId="figure@137.[140,1107,165,1288]" captionTargetPageId="137" captionText="Figure 4. Optimal cladogram for the Micropholcommatidae under equal and implied weights, and one of six equally parsimonious trees recovered from an equal weights analysis of the ‘new’ matrix (length = 135; see Appendix II). With the exception of the highlighted (*) node ‘Patelliella adusta + Micropholcommatini’, all clades shown were recovered in all iterations of an implied weights analysis (K=1–6); the phylogenetic position of P. adusta changed to ‘P. adusta + Textricellini’ under K-values 1–5. Unambiguous character optimisations are shown for each node, and bold species are newly described, with generic type species highlighted (*)." figureDoi="http://doi.org/10.5281/zenodo.3767693" httpUri="https://zenodo.org/record/3767693/files/figure.png" pageId="27" pageNumber="28">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BEA06513FFAA1B67FF071FF1FC818C4B" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="F6053698FFAA1B67FF071FF1FC818C4B" blockId="28.[140,1108,166,1285]" pageId="28" pageNumber="29">
<emphasis id="C4CEEA8AFFAA1B67FF071FF1FED18DC5" bold="true" box="[185,312,166,192]" pageId="28" pageNumber="29">
<emphasis id="C4CEEA8AFFAA1B67FF071FF1FEDA8DC5" box="[185,307,166,192]" inLineHeading="true" pageId="28" pageNumber="29" reason="1">Diagnosis</emphasis>
.
</emphasis>
<taxonomicName id="31BA4D1BFFAA1B67FEFC1FF0FDCE8DC4" authorityName="Forster C" authorityYear="1959" box="[322,551,167,193]" class="Arachnida" family="Micropholcommatidae" genus="Pua" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="novaezealandiae">
<emphasis id="C4CEEA8AFFAA1B67FEFC1FF0FDCE8DC4" box="[322,551,167,193]" italics="true" pageId="28" pageNumber="29">Pua novaezealandiae</emphasis>
</taxonomicName>
can be distinguished from all other 
<taxonomicName id="31BA4D1BFFAA1B67FC6F1FF0FEEE8DE1" class="Arachnida" family="Null" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="146" phylum="Arthropoda" rank="tribe" tribe="Micropholcommatini">Micropholcommatini</taxonomicName>
by the presence of a short (length &lt;5× width), hooked embolus situated distally on a rotated bulb (
<figureCitation id="6E812A1DFFAA1B67FE0F1FBAFDEA8C02" box="[433,515,237,263]" captionStart="Figure 29" captionStartId="159.[140,212,1136,1158]" captionTargetBox="[140,1108,165,1116]" captionTargetId="figure@159.[140,1108,165,1116]" captionTargetPageId="159" captionText="Figure 29. Scanning electron micrographs of pedipalp of male Pua novaezealandiae Forster from the Makarora Valley, New Zealand (AMNH)." figureDoi="http://doi.org/10.5281/zenodo.3767723" httpUri="https://zenodo.org/record/3767723/files/figure.png" pageId="28" pageNumber="29">Figs 29</figureCitation>
A–C). Other diagnostic characters include the loss of the AME (
<figureCitation id="6E812A1DFFAA1B67FEBA1E47FE838C2E" box="[260,362,272,299]" captionStart="Figure 25" captionStartId="155.[140,212,1465,1487]" captionTargetBox="[140,1108,940,1445]" captionTargetId="figure@155.[140,1109,940,1447]" captionTargetPageId="155" captionText="Figure 25. Pua novaezealandiae Forster from the Makarora Valley, New Zealand (AMNH): A male habitus, antero-lateral view B female cephalothorax, frontal view." figureDoi="http://doi.org/10.5281/zenodo.3767715" httpUri="https://zenodo.org/record/3767715/files/figure.png" pageId="28" pageNumber="29">Fig. 25B</figureCitation>
), the presence of very large, globular spermathecae (Fig. 15F), and the presence of a dorsal scute on the female abdomen (
<figureCitation id="6E812A1DFFAA1B67FD4B1E64FCB28C4B" box="[757,859,307,334]" captionStart="Figure 24" captionStartId="155.[140,212,784,806]" captionTargetBox="[140,1108,165,764]" captionTargetId="figure@155.[140,1109,165,768]" captionTargetPageId="155" captionText="Figure 24. Pua novaezealandiae Forster from the Makarora Valley, New Zealand (AMNH): A male habitus, dorsal view B female habitus, dorsal view C male abdomen, ventral view D female abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3767713" httpUri="https://zenodo.org/record/3767713/files/figure.png" pageId="28" pageNumber="29">Fig. 24B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BEA06513FFAA1B67FF071E01FED189B8" pageId="28" pageNumber="29" type="description">
<paragraph id="F6053698FFAA1B67FF071E01FBD28F24" blockId="28.[140,1108,166,1285]" pageId="28" pageNumber="29">
<emphasis id="C4CEEA8AFFAA1B67FF071E01FEA68C75" bold="true" box="[185,335,342,368]" pageId="28" pageNumber="29">
<emphasis id="C4CEEA8AFFAA1B67FF071E01FEA38C75" box="[185,330,342,368]" inLineHeading="true" pageId="28" pageNumber="29" reason="1">Description</emphasis>
.
</emphasis>
Very small, entelegyne 
<taxonomicName id="31BA4D1BFFAA1B67FDE91E00FD308C74" box="[599,729,343,369]" order="Null" pageId="28" pageNumber="132" rank="superFamily" superFamily="Araneoidea">Araneoidea</taxonomicName>
; total length 0.80 to 1.00. 
<emphasis id="C4CEEA8AFFAA1B67FBBB1E00FF0D8C91" italics="true" pageId="28" pageNumber="29">Cephalothorax</emphasis>
: Carapace without glandular depressions above maxillae (
<figureCitation id="6E812A1DFFAA1B67FC3A1E2DFC1B8C90" box="[900,1010,378,405]" captionStart="Figure 26" captionStartId="156.[140,212,1136,1158]" captionTargetBox="[140,1108,165,1115]" captionTargetId="figure@156.[140,1108,165,1117]" captionTargetPageId="156" captionText="Figure 26. Scanning electron micrographs of Pua novaezealandiae Forster from the Makarora Valley, New Zealand (AMNH): A female eyes and clypeus, frontal view B male sternum, ventro-lateral view C male abdominal cuticle D male corner of carapace above left maxilla E female pars thoracica, lateral view, showing setal tubercles F female chelicerae and labrum." figureDoi="http://doi.org/10.5281/zenodo.3767717" httpUri="https://zenodo.org/record/3767717/files/figure.png" pageId="28" pageNumber="29">Fig. 26D</figureCitation>
); cuticle without glandular pits (
<figureCitation id="6E812A1DFFAA1B67FE221ECAFDEA8CBD" box="[412,515,413,440]" captionStart="Figure 26" captionStartId="156.[140,212,1136,1158]" captionTargetBox="[140,1108,165,1115]" captionTargetId="figure@156.[140,1108,165,1117]" captionTargetPageId="156" captionText="Figure 26. Scanning electron micrographs of Pua novaezealandiae Forster from the Makarora Valley, New Zealand (AMNH): A female eyes and clypeus, frontal view B male sternum, ventro-lateral view C male abdominal cuticle D male corner of carapace above left maxilla E female pars thoracica, lateral view, showing setal tubercles F female chelicerae and labrum." figureDoi="http://doi.org/10.5281/zenodo.3767717" httpUri="https://zenodo.org/record/3767717/files/figure.png" pageId="28" pageNumber="29">Fig. 26E</figureCitation>
); margins fused to sternum via pleural sclerites. Six eyes present on anterior margin of pars cephalica (
<figureCitation id="6E812A1DFFAA1B67FD7B1E97FCC28CDE" box="[709,811,448,475]" captionStart="Figure 25" captionStartId="155.[140,212,1465,1487]" captionTargetBox="[140,1108,940,1445]" captionTargetId="figure@155.[140,1109,940,1447]" captionTargetPageId="155" captionText="Figure 25. Pua novaezealandiae Forster from the Makarora Valley, New Zealand (AMNH): A male habitus, antero-lateral view B female cephalothorax, frontal view." figureDoi="http://doi.org/10.5281/zenodo.3767715" httpUri="https://zenodo.org/record/3767715/files/figure.png" pageId="28" pageNumber="29">Fig. 25B</figureCitation>
); AME absent. Chelicerae without bulging anterior projections in males; promargin with true teeth, a pair of fused setal sockets adjacent to base of fang and single peg tooth in males (
<figureCitation id="6E812A1DFFAA1B67FC771D50FBC78F24" box="[969,1070,519,545]" captionStart="Figure 27" captionStartId="157.[140,212,1263,1285]" captionTargetBox="[140,1108,291,1241]" captionTargetId="figure@157.[140,1108,291,1243]" captionTargetPageId="157" captionText="Figure 27. Scanning electron micrographs of Pua novaezealandiae Forster from the Makarora Valley, New Zealand (AMNH): A female chelicera, frontal view B female cheliceral promargin C female pedipalp, frontal view D tip of female pedipalp, showing fused tibia-tarsus E male chelicera, antero-lateral view F male cheliceral promargin." figureDoi="http://doi.org/10.5281/zenodo.3767719" httpUri="https://zenodo.org/record/3767719/files/figure.png" pageId="28" pageNumber="29">Fig. 27F</figureCitation>
).
</paragraph>
<paragraph id="F6053698FFAA1B67FF071D7CFC058FAB" blockId="28.[140,1108,166,1285]" pageId="28" pageNumber="29">
<emphasis id="C4CEEA8AFFAA1B67FF071D7CFE268F41" box="[185,463,554,580]" italics="true" pageId="28" pageNumber="29">Legs and female pedipalp</emphasis>
: Legs three-clawed (
<figureCitation id="6E812A1DFFAA1B67FD021D7DFCC08F41" box="[700,809,554,580]" captionStart="Figure 28" captionStartId="158.[140,212,940,962]" captionTargetBox="[140,1108,291,920]" captionTargetId="figure@158.[140,1108,291,923]" captionTargetPageId="158" captionText="Figure 28. Scanning electron micrographs of female Pua novaezealandiae Forster from the Makarora Valley, New Zealand (AMNH): A leg I tarsal organ B leg I lyriform organ C leg I tibial trichobothrium D leg IV claws, showing elongate inferior claw." figureDoi="http://doi.org/10.5281/zenodo.3767721" httpUri="https://zenodo.org/record/3767721/files/figure.png" pageId="28" pageNumber="29">Fig. 28D</figureCitation>
), covered with smooth or serrate hair-like setae. Trichobothria present on legs; tibiae each with two (legs I–III) or three (leg IV) trichobothria (
<figureCitation id="6E812A1DFFAA1B67FDBF1D27FD848F8F" box="[513,621,624,650]" captionStart="Figure 28" captionStartId="158.[140,212,940,962]" captionTargetBox="[140,1108,291,920]" captionTargetId="figure@158.[140,1108,291,923]" captionTargetPageId="158" captionText="Figure 28. Scanning electron micrographs of female Pua novaezealandiae Forster from the Makarora Valley, New Zealand (AMNH): A leg I tarsal organ B leg I lyriform organ C leg I tibial trichobothrium D leg IV claws, showing elongate inferior claw." figureDoi="http://doi.org/10.5281/zenodo.3767721" httpUri="https://zenodo.org/record/3767721/files/figure.png" pageId="28" pageNumber="29">Fig. 28C</figureCitation>
); metatarsi without trichobothria. Female pedipalp four-segmented, with fused tibia-tarsus; claw absent (
<figureCitation id="6E812A1DFFAA1B67FCF31DC4FC4B8FAB" box="[845,930,659,686]" captionStart="Figure 27" captionStartId="157.[140,212,1263,1285]" captionTargetBox="[140,1108,291,1241]" captionTargetId="figure@157.[140,1108,291,1243]" captionTargetPageId="157" captionText="Figure 27. Scanning electron micrographs of Pua novaezealandiae Forster from the Makarora Valley, New Zealand (AMNH): A female chelicera, frontal view B female cheliceral promargin C female pedipalp, frontal view D tip of female pedipalp, showing fused tibia-tarsus E male chelicera, antero-lateral view F male cheliceral promargin." figureDoi="http://doi.org/10.5281/zenodo.3767719" httpUri="https://zenodo.org/record/3767719/files/figure.png" pageId="28" pageNumber="29">Figs 27</figureCitation>
C–D).
</paragraph>
<paragraph id="F6053698FFAA1B67FF071DE0FE358EC2" blockId="28.[140,1108,166,1285]" pageId="28" pageNumber="29">
<emphasis id="C4CEEA8AFFAA1B67FF071DE0FEF68FD4" box="[185,287,695,721]" italics="true" pageId="28" pageNumber="29">Abdomen</emphasis>
: Abdomen oval-globose; anterior sclerite present around epigastric region and petiole; large dorsal scute present on males and females (
<figureCitation id="6E812A1DFFAA1B67FCCF1D8DFC2D8FF1" box="[881,964,730,756]" captionStart="Figure 24" captionStartId="155.[140,212,784,806]" captionTargetBox="[140,1108,165,764]" captionTargetId="figure@155.[140,1109,165,768]" captionTargetPageId="155" captionText="Figure 24. Pua novaezealandiae Forster from the Makarora Valley, New Zealand (AMNH): A male habitus, dorsal view B female habitus, dorsal view C male abdomen, ventral view D female abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3767713" httpUri="https://zenodo.org/record/3767713/files/figure.png" pageId="28" pageNumber="29">Figs 24</figureCitation>
A–B); lateral sclerotic strips present on males; posterior sclerotic ring surrounding spinnerets and colulus. Six spinnerets situated posterior to fleshy colulus (
<figureCitation id="6E812A1DFFAA1B67FCA91C77FC838E3F" box="[791,874,800,826]" captionStart="Figure 23" captionStartId="154.[140,212,1453,1475]" captionTargetBox="[140,1108,165,1434]" captionTargetId="figure@154.[140,1108,165,1436]" captionTargetPageId="154" captionText="Figure 23. Scanning electron micrographs of posterior median spinnerets of female (left) and male (right) Micropholcommatini: A–B Micropholcomma bryophilum (Butler) from near Marysville, Victoria (WAM T94453) C–D Pua novaezealandiae Forster from the Makarora Valley, New Zealand (AMNH) E–F Austropholcomma florentine sp. n. from the Florentine Valley, Tasmania (FMC) G–H Tricellina gertschi (Forster &amp; Platnick) from near Chaitén, Chile (AMNH). Note the apparent loss of the anterior CY gland spigot in female Austropholcomma and Tricellina. Arrows denote the anterior, mesal direction." figureDoi="http://doi.org/10.5281/zenodo.3767711" httpUri="https://zenodo.org/record/3767711/files/figure.png" pageId="28" pageNumber="29">Figs 23</figureCitation>
C–D); PMS without AC gland spigots; PLS with complete triad. Anterior tracheal system well-developed, with relatively large lateral atria and multiple radiating tracheae; second tracheal tubes on each side looping mesally and extending through petiole into cephalothorax; posterior tracheal spiracle absent.
</paragraph>
<paragraph id="F6053698FFAA1B67FF071C87FED189B8" blockId="28.[140,1108,166,1285]" pageId="28" pageNumber="29">
<emphasis id="C4CEEA8AFFAA1B67FF071C87FEC98EEF" box="[185,288,976,1002]" italics="true" pageId="28" pageNumber="29">Genitalia</emphasis>
: Male pedipalp (
<figureCitation id="6E812A1DFFAA1B67FE5C1C87FDDD8EEF" box="[482,564,976,1002]" captionStart="Figure 29" captionStartId="159.[140,212,1136,1158]" captionTargetBox="[140,1108,165,1116]" captionTargetId="figure@159.[140,1108,165,1116]" captionTargetPageId="159" captionText="Figure 29. Scanning electron micrographs of pedipalp of male Pua novaezealandiae Forster from the Makarora Valley, New Zealand (AMNH)." figureDoi="http://doi.org/10.5281/zenodo.3767723" httpUri="https://zenodo.org/record/3767723/files/figure.png" pageId="28" pageNumber="29">Fig. 29</figureCitation>
) relatively simple; patella with distally-directed, hooked ligulate retrolateral apophysis; bulb rotated 180 degrees, with embolus inserting distally relative to cymbium; tegulum smooth, with straight, evaginated tegular ridge; embolus exposed, short (length &lt;5× width) and hooked; conductor absent. Female genitalia (Fig. 15F) with pair of separate, very large, globular anterior spermathecae; insemination ducts short, barely visible in dorsal view; fertilisation ducts simple, curved.
</paragraph>
</subSubSection>
<subSubSection id="BEA06513FFAA1B67FF071B91FC0E8800" pageId="28" pageNumber="29" type="distribution">
<paragraph id="F6053698FFAA1B67FF071B91FE1689E4" blockId="28.[140,1108,166,1285]" box="[185,511,1222,1249]" pageId="28" pageNumber="29">
<emphasis id="C4CEEA8AFFAA1B67FF071B91FEB189E5" bold="true" box="[185,344,1222,1248]" pageId="28" pageNumber="29">
<emphasis id="C4CEEA8AFFAA1B67FF071B91FEBA89E5" box="[185,339,1222,1248]" inLineHeading="true" pageId="28" pageNumber="29" reason="1">Distribution</emphasis>
.
</emphasis>
<collectingCountry id="8EAD7608FFAA1B67FEE11B90FE1289E4" box="[351,507,1223,1249]" name="New Zealand" pageId="28" pageNumber="29">New Zealand</collectingCountry>
.
</paragraph>
<paragraph id="F6053698FFAA1B67FF071BBEFC0E8800" blockId="28.[140,1108,166,1285]" box="[185,999,1257,1285]" pageId="28" pageNumber="29">
<emphasis id="C4CEEA8AFFAA1B67FF071BBEFEB68806" bold="true" box="[185,351,1257,1283]" pageId="28" pageNumber="29">
<emphasis id="C4CEEA8AFFAA1B67FF071BBEFEB38806" box="[185,346,1257,1283]" inLineHeading="true" pageId="28" pageNumber="29" reason="1">Composition</emphasis>
.
</emphasis>
Only the 
<typeStatus id="2901883AFFAA1B67FE6D1BBDFDEA8801" box="[467,515,1258,1284]" pageId="28" pageNumber="29">type</typeStatus>
species 
<taxonomicName id="31BA4D1BFFAA1B67FDE11BBCFC0A8800" authority="Forster, 1959" authorityName="Forster C" authorityYear="1959" box="[607,995,1258,1285]" class="Arachnida" family="Micropholcommatidae" genus="Pua" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="novaezealandiae">
<emphasis id="C4CEEA8AFFAA1B67FDE11BBCFCAB8801" box="[607,834,1258,1284]" italics="true" pageId="28" pageNumber="29">Pua novaezealandiae</emphasis>
<bibRefCitation id="922B4B69FFAA1B67FCF71BBDFC0A8800" author="Forster RR" box="[841,995,1258,1285]" pageId="28" pageNumber="29" pagination="269 - 329" refId="ref70149" refString="Forster RR (1959) The spiders of the family Symphytognathidae. Transactions of the Royal Society of New Zealand 86: 269 - 329." type="journal article" year="1959">Forster, 1959</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>