An Annotated Catalog of the Leiodidae (Coleoptera) of the Nearctic Region (Continental North America North of Mexico)
Author
Peck, Stewart B.
Author
Newton, Alfred F.
text
The Coleopterists Bulletin
2017
2017-06-27
71
2
211
258
http://dx.doi.org/10.1649/0010-065x-71.2.211
journal article
10.1649/0010-065X-71.2.211
1938-4394
5460664
Genus
ANISOTOMA
Panzer, 1797
Anisotoma
Panzer, 1797: 8
.
Type
species:
Tritoma glabra
Fabricius, 1787
(designated by
Hatch, 1929b: 50
; “
A. picea
(Illig)
” designated by
Thomson 1859: 58
).
Note.
Authority given as Knoch in
Kugelann (1794)
(Neave 1939–49); two original species (“
picea
Kug.
”, “
glabra
Kug.
”).
North American Revision.
Wheeler (1979)
and Wheeler and Miller (2005).
Biology.
The species generally eat plasmodia or fruiting bodies of Myxomycetes slime molds (
e.g
.,
Wheeler 1979
; Lawrence and Newton 1980;
Newton 1984
;
Wheeler 1984a
, b).
Distribution.
Holarctic,
Nepal
,
China
,
Taiwan
, N Neotropical (to
Panama
) (57 species). New (unpublished) state and province records are included in the distributions below based on SBP voucher specimens deposited in CMNC.
=Pentatoma
Schneider, 1792: 339 (preoccupied, not Olivier, 1789).
Type
species:
Sphaeridium humerale
Olivier, 1790
(cited as
Fabricius, 1792
) (designated by
Hatch 1929b: 50
).
=
Anisotoma
Illiger, 1798: 69
(attributed to Knoch).
Type
species: none designated? (13 original species, including “
piceum
Panzer
” and “
glabrum
Panzer”).
=
Leiodes
;
Schmidt 1841: 132
(misidentification of
Leiodes
Latreille, 1797
).
=
Eucyrta
Portevin, 1927: 82
(preoccupied, not Felder 1874);
Wheeler 1979: 267
(synonymy).
Type
species:
Eucyrta didymata
Portevin, 1927
(designated by
Hatch, 1929b: 57
).
A. amica
Brown, 1937b: 201
;
Hatch 1957: 31
;
Wheeler 1979: 293
.
Holotype
male in CNCI. Type locality: Mt. Layall,
Quebec
. North American distribution:
Canada
:
AB, BC, MB, NF, ON, QC.
USA
:
AK, AZ, CO, NM, WA. Biology: Fungal associates (
Stephenson
et al.
1994
).
A. basalis
(
LeConte, 1853: 285
)
(
Leiodes
)
;
Horn 1880: 299
(
Liodes
);
Brown 1937b: 200
;
Wheeler 1979: 292
; Majka and Langor 2008: 382.
Lectotype
female in MCZC (designated by
Wheeler 1979: 292
). Type locality: Pennsylvania or New Jersey. North American distribution:
Canada
:
MB, NB, NF, NS, ON, QC.
USA
:
AL, AR, CT, GA, IL, IN, KY, LA, MA, MD, ME, MI, MN, MS, NC, NH, NJ, NY, OK, PA, RI, SC, TX, VA, VT, WI. Larva:
Wheeler (1990a
, b). Biology: Fungal associates (Wheeler and Hoebeke 1990;
Stephenson
et al.
1994
).
=
Leiodes dichroa
LeConte, 1853: 285
:
Wheeler 1979: 292
(synonymy).
Lectotype
male in MCZC (designated by
Wheeler 1979: 292
). Type locality: Ann Arbor,
Michigan
.
A. bifoveata
Wheeler, 1979: 273
.
Holotype
male in MCZC. Type locality:
Conkles Hollow
,
Hocking County
, Ohio.
North American
distribution:
Canada
:
ON.
USA:
AL, AR, CT, GA, IL, IN, KY, LA, MI, MO, MS, NC, NY, PA, SC, TX, VA, WV.
A. blanchardi
(
Horn, 1880: 295
)
(
Liodes
)
;
Brown 1937b: 196
;
Wheeler 1979: 273
; Majka and Langor 2008: 383.
Lectotype
male in MCZC (designated by
Wheeler 1979: 273
). Type locality: Tewksbury, Massachusetts. North American distribution:
Canada
:
MB, NB, NS, ON, QC.
USA
:
AL, FL, GA, IL, KY, LA, MA, MD, ME, MI, MN, NC, NH, OH, PA, SC, TN, TX, VA, VT. Larva:
Kilian (2005a)
. Biology: Fungal associates (
Stephenson
et al.
1994
).
A. confusa
(
Horn, 1880: 299
)
(
Liodes
)
;
Brown 1937b: 197
;
Hatch 1957: 31
;
Wheeler 1979: 288
.
Holotype
male in MCZC. Type locality: “Western Nevada” (probably vicinity of Lake Tahoe, Reno, or Carson City). North American distribution:
Canada
:
AB, BC.
USA
:
AK, CA, OR, NV, WA.
=
A. interstrialis
Hatch, 1936: 38
;
Brown 1937b: 197
;
Wheeler 1979: 288
(synonymy).
Holotype
in USNM. Type locality: Seattle,
Washington
.
A. discolor
(F. E.
Melsheimer, 1844: 103
)
(
Leiodes
)
;
LeConte 1853: 285
;
Horn 1880: 298
;
Brown 1937b: 202
;
Wheeler 1979: 295
; Majka and Langor 2008: 384.
Lectotype
female in MCZC (designated by
Wheeler 1979: 295
). Type locality: “Pennsylvania.” North American distribution:
Canada
:
BC, MB, NB, NS, ON, QC.
USA
:
AL, AR, CT, DC, FL, GA, IA, IL, IN, KY, LA, MA, MD, ME, MI, MN, MO, MS, NC, NH, NJ, NY, OH, OK, PA, RI, SC, TN, TX, VA, WV. Larva:
Kilian (2003)
.
=
Anistoma
[sic]
piceum
F. V.
Melsheimer, 1806: 12
(attributed to Knoch;
nomen nudum
); F. E.
Melsheimer 1844: 103
(as synonym of
Leiodes discolor
F. E. Melsheimer
);
Wheeler 1979: 288
(synonymy).
A. errans
Brown, 1937b: 202
;
Hatch 1957: 32
;
Wheeler 1979: 277
; Majka and Langor 2008: 384.
Holotype
male in CNCI. Type locality: Biscotasing,
Ontario
. North American distribution:
Canada
:
AB, BC, NB, NF, NS, ON, QC, SK, YT.
USA
:
AK, CA, ID, ME, MI, NH, OR, RI, VT, WA, WY. Biology: Fungal associates (
Stephenson
et al.
1994
). Larva:
Newton (1991)
.
A. expolita
Brown, 1937b: 199
(replacement name for
Leiodes polita
LeConte, 1853
);
Wheeler 1979: 300
. North American distribution:
USA
:
AR, IL, IN, KY, PA, SC.
=
Leiodes polita
LeConte, 1853: 285
(preoccupied, not
Silpha polita
Marsham, 1802
, not now congeneric but replaced before 1961 and replacement name in use (
ICZN 1999
: Art. 59.3));
Horn 1880: 297
(
Liodes
).
Holotype
male in MCZC. Type locality: “
South Carolina
.”
A. geminata
(
Horn, 1880: 299
)
(
Liodes
)
;
Brown 1937b: 196
;
Wheeler 1979: 288
; Majka and Langor 2008: 385.
Lectotype
male in MCZC (designated by
Wheeler 1979: 288
). Type locality: Tyngsboro, Massachussets. North American distribution:
Canada
:
BC, NB, NF, NS, ON, QC.
USA
:
CT, GA, IL, IN, KY, MA, ME, MI, NC, NH, NJ, NY, OH, PA, RI, SC, TN, TX, VA, WI. Biology: Fungal associates (
Stephenson
et al.
1994
).
A. globososa
Hatch, 1929b: 56
(replacement name for
A. globosa
(
LeConte, 1850
))
;
Hatch 1957: 31
;
Brown 1937b: 197
;
Wheeler 1979
a: 298; Majka and Langor 2008: 386. North American distribution:
Canada
:
AB, BC, MB, NF, ON, QC, SK.
USA
:
AK, CO, CT, ID, MA, ME, MI, NH, SD, TX, WA, WI. Biology: Fungal associates (
Stephenson
et al.
1994
).
=
Cyrtusa globosa
LeConte, 1850: 222
(preoccupied in
Anisotoma
, not
Tetratoma globosa
Herbst, 1791
; not now congeneric, but replaced before 1961 and replacement name in use (
ICZN 1999
: Art. 59.3));
LeConte 1853: 285
(
Leiodes
);
Horn 1880: 297
(
Liodes
).
Lectotype
male in MCZC (designated by
Wheeler 1979: 298
). Type locality: Sault Ste Marie,
Michigan
.
A. humeralis
(
Olivier, 1790: 15
)
(
Sphaeridium
)
;
Hatch 1929b: 51
;
Wheeler 1979: 290
.
Type
in MNHN?
Type
locality: “environs de Paris”. North American distribution:
Canada
:
BC.
USA
:
IN. Extra-limital distribution: Europe; a Holarctic species. Larva:
Ratajczak (1995)
and
Kilian (1998)
.
A. inops
Brown, 1937b: 198
;
Wheeler 1979: 299
; Majka and Langor 2008: 386;
Bouchard 2001: 130
.
Holotype
male in CNCI. Type locality: Arnprior,
Ontario
. North American distribution:
Canada
:
NB, NF, NS, ON, QC.
USA
:
ME, NC, NH, NY.
A. nevadensis
Brown, 1937b: 199
(replacement name for
Liodes globosa
var.
bicolor
Horn, 1880
);
Hatch 1957: 31
;
Wheeler 1979: 300
. North American distribution:
USA
:
CA, NV, OR, WA.
=
Liodes globosa
var.
bicolor
Horn, 1880: 297
(preoccupied by
Anisotoma bicolor
Schmidt, 1841
).
Lectotype
male in MCZC (designated by
Wheeler 1979: 300
). Type locality: “
Nevada
” [probably vicinity of Lake Tahoe, Reno, or Carson City].
A. obsoleta
(
Horn, 1880: 298
)
(
Liodes
)
;
Brown 1937b: 201
; Majka and Langor 2008: 386 (revalidation of name).
Lectotype
male in MCZC (designated by
Wheeler 1979: 276
). Type locality: none given, range stated as “New Hampshire to Virginia and Canada.” North American distribution:
Canada
:
BC, MB, NB, NF, NS, ON, QC.
USA
:
AL, AR, CT, FL, GA, IA, IL, KY, LA, MA, ME, MI, MN, MS, NC, NH, NY, OH, PA, SC, TN, VA, VT, WI, WV. Biology: Fungal associates
Stephenson
et al.
1994
). Larva:
Kilian (2005b)
(as
A
.
horni
Wheeler
).
=
A. horni
Wheeler, 1979: 276
(unnecessary replacement name for
Liodes obsoleta
Horn, 1880
, incorrectly thought to be preoccupied by
Pallodes obsoletus
F. E.
Melsheimer, 1844
(now in
Anogdus
LeConte
)). Biology:
Bouchard (2001)
.
A. reticulonota
Wheeler, 1979: 298
.
Holotype
male in MCZC. Type locality:
White Mountains
,
Apache County
,
Arizona
.
North American
distribution:
USA
:
AZ, NM.