An Annotated Catalog of the Leiodidae (Coleoptera) of the Nearctic Region (Continental North America North of Mexico) Author Peck, Stewart B. Author Newton, Alfred F. text The Coleopterists Bulletin 2017 2017-06-27 71 2 211 258 http://dx.doi.org/10.1649/0010-065x-71.2.211 journal article 10.1649/0010-065X-71.2.211 1938-4394 5460664 Genus ANISOTOMA Panzer, 1797 Anisotoma Panzer, 1797: 8 . Type species: Tritoma glabra Fabricius, 1787 (designated by Hatch, 1929b: 50 ; “ A. picea (Illig) ” designated by Thomson 1859: 58 ). Note. Authority given as Knoch in Kugelann (1794) (Neave 1939–49); two original species (“ picea Kug. ”, “ glabra Kug. ”). North American Revision. Wheeler (1979) and Wheeler and Miller (2005). Biology. The species generally eat plasmodia or fruiting bodies of Myxomycetes slime molds ( e.g ., Wheeler 1979 ; Lawrence and Newton 1980; Newton 1984 ; Wheeler 1984a , b). Distribution. Holarctic, Nepal , China , Taiwan , N Neotropical (to Panama ) (57 species). New (unpublished) state and province records are included in the distributions below based on SBP voucher specimens deposited in CMNC. =Pentatoma Schneider, 1792: 339 (preoccupied, not Olivier, 1789). Type species: Sphaeridium humerale Olivier, 1790 (cited as Fabricius, 1792 ) (designated by Hatch 1929b: 50 ). = Anisotoma Illiger, 1798: 69 (attributed to Knoch). Type species: none designated? (13 original species, including “ piceum Panzer ” and “ glabrum Panzer”). = Leiodes ; Schmidt 1841: 132 (misidentification of Leiodes Latreille, 1797 ). = Eucyrta Portevin, 1927: 82 (preoccupied, not Felder 1874); Wheeler 1979: 267 (synonymy). Type species: Eucyrta didymata Portevin, 1927 (designated by Hatch, 1929b: 57 ). A. amica Brown, 1937b: 201 ; Hatch 1957: 31 ; Wheeler 1979: 293 . Holotype male in CNCI. Type locality: Mt. Layall, Quebec . North American distribution: Canada : AB, BC, MB, NF, ON, QC. USA : AK, AZ, CO, NM, WA. Biology: Fungal associates ( Stephenson et al. 1994 ). A. basalis ( LeConte, 1853: 285 ) ( Leiodes ) ; Horn 1880: 299 ( Liodes ); Brown 1937b: 200 ; Wheeler 1979: 292 ; Majka and Langor 2008: 382. Lectotype female in MCZC (designated by Wheeler 1979: 292 ). Type locality: Pennsylvania or New Jersey. North American distribution: Canada : MB, NB, NF, NS, ON, QC. USA : AL, AR, CT, GA, IL, IN, KY, LA, MA, MD, ME, MI, MN, MS, NC, NH, NJ, NY, OK, PA, RI, SC, TX, VA, VT, WI. Larva: Wheeler (1990a , b). Biology: Fungal associates (Wheeler and Hoebeke 1990; Stephenson et al. 1994 ). = Leiodes dichroa LeConte, 1853: 285 : Wheeler 1979: 292 (synonymy). Lectotype male in MCZC (designated by Wheeler 1979: 292 ). Type locality: Ann Arbor, Michigan . A. bifoveata Wheeler, 1979: 273 . Holotype male in MCZC. Type locality: Conkles Hollow , Hocking County , Ohio. North American distribution: Canada : ON. USA: AL, AR, CT, GA, IL, IN, KY, LA, MI, MO, MS, NC, NY, PA, SC, TX, VA, WV. A. blanchardi ( Horn, 1880: 295 ) ( Liodes ) ; Brown 1937b: 196 ; Wheeler 1979: 273 ; Majka and Langor 2008: 383. Lectotype male in MCZC (designated by Wheeler 1979: 273 ). Type locality: Tewksbury, Massachusetts. North American distribution: Canada : MB, NB, NS, ON, QC. USA : AL, FL, GA, IL, KY, LA, MA, MD, ME, MI, MN, NC, NH, OH, PA, SC, TN, TX, VA, VT. Larva: Kilian (2005a) . Biology: Fungal associates ( Stephenson et al. 1994 ). A. confusa ( Horn, 1880: 299 ) ( Liodes ) ; Brown 1937b: 197 ; Hatch 1957: 31 ; Wheeler 1979: 288 . Holotype male in MCZC. Type locality: “Western Nevada” (probably vicinity of Lake Tahoe, Reno, or Carson City). North American distribution: Canada : AB, BC. USA : AK, CA, OR, NV, WA. = A. interstrialis Hatch, 1936: 38 ; Brown 1937b: 197 ; Wheeler 1979: 288 (synonymy). Holotype in USNM. Type locality: Seattle, Washington . A. discolor (F. E. Melsheimer, 1844: 103 ) ( Leiodes ) ; LeConte 1853: 285 ; Horn 1880: 298 ; Brown 1937b: 202 ; Wheeler 1979: 295 ; Majka and Langor 2008: 384. Lectotype female in MCZC (designated by Wheeler 1979: 295 ). Type locality: “Pennsylvania.” North American distribution: Canada : BC, MB, NB, NS, ON, QC. USA : AL, AR, CT, DC, FL, GA, IA, IL, IN, KY, LA, MA, MD, ME, MI, MN, MO, MS, NC, NH, NJ, NY, OH, OK, PA, RI, SC, TN, TX, VA, WV. Larva: Kilian (2003) . = Anistoma [sic] piceum F. V. Melsheimer, 1806: 12 (attributed to Knoch; nomen nudum ); F. E. Melsheimer 1844: 103 (as synonym of Leiodes discolor F. E. Melsheimer ); Wheeler 1979: 288 (synonymy). A. errans Brown, 1937b: 202 ; Hatch 1957: 32 ; Wheeler 1979: 277 ; Majka and Langor 2008: 384. Holotype male in CNCI. Type locality: Biscotasing, Ontario . North American distribution: Canada : AB, BC, NB, NF, NS, ON, QC, SK, YT. USA : AK, CA, ID, ME, MI, NH, OR, RI, VT, WA, WY. Biology: Fungal associates ( Stephenson et al. 1994 ). Larva: Newton (1991) . A. expolita Brown, 1937b: 199 (replacement name for Leiodes polita LeConte, 1853 ); Wheeler 1979: 300 . North American distribution: USA : AR, IL, IN, KY, PA, SC. = Leiodes polita LeConte, 1853: 285 (preoccupied, not Silpha polita Marsham, 1802 , not now congeneric but replaced before 1961 and replacement name in use ( ICZN 1999 : Art. 59.3)); Horn 1880: 297 ( Liodes ). Holotype male in MCZC. Type locality: “ South Carolina .” A. geminata ( Horn, 1880: 299 ) ( Liodes ) ; Brown 1937b: 196 ; Wheeler 1979: 288 ; Majka and Langor 2008: 385. Lectotype male in MCZC (designated by Wheeler 1979: 288 ). Type locality: Tyngsboro, Massachussets. North American distribution: Canada : BC, NB, NF, NS, ON, QC. USA : CT, GA, IL, IN, KY, MA, ME, MI, NC, NH, NJ, NY, OH, PA, RI, SC, TN, TX, VA, WI. Biology: Fungal associates ( Stephenson et al. 1994 ). A. globososa Hatch, 1929b: 56 (replacement name for A. globosa ( LeConte, 1850 )) ; Hatch 1957: 31 ; Brown 1937b: 197 ; Wheeler 1979 a: 298; Majka and Langor 2008: 386. North American distribution: Canada : AB, BC, MB, NF, ON, QC, SK. USA : AK, CO, CT, ID, MA, ME, MI, NH, SD, TX, WA, WI. Biology: Fungal associates ( Stephenson et al. 1994 ). = Cyrtusa globosa LeConte, 1850: 222 (preoccupied in Anisotoma , not Tetratoma globosa Herbst, 1791 ; not now congeneric, but replaced before 1961 and replacement name in use ( ICZN 1999 : Art. 59.3)); LeConte 1853: 285 ( Leiodes ); Horn 1880: 297 ( Liodes ). Lectotype male in MCZC (designated by Wheeler 1979: 298 ). Type locality: Sault Ste Marie, Michigan . A. humeralis ( Olivier, 1790: 15 ) ( Sphaeridium ) ; Hatch 1929b: 51 ; Wheeler 1979: 290 . Type in MNHN? Type locality: “environs de Paris”. North American distribution: Canada : BC. USA : IN. Extra-limital distribution: Europe; a Holarctic species. Larva: Ratajczak (1995) and Kilian (1998) . A. inops Brown, 1937b: 198 ; Wheeler 1979: 299 ; Majka and Langor 2008: 386; Bouchard 2001: 130 . Holotype male in CNCI. Type locality: Arnprior, Ontario . North American distribution: Canada : NB, NF, NS, ON, QC. USA : ME, NC, NH, NY. A. nevadensis Brown, 1937b: 199 (replacement name for Liodes globosa var. bicolor Horn, 1880 ); Hatch 1957: 31 ; Wheeler 1979: 300 . North American distribution: USA : CA, NV, OR, WA. = Liodes globosa var. bicolor Horn, 1880: 297 (preoccupied by Anisotoma bicolor Schmidt, 1841 ). Lectotype male in MCZC (designated by Wheeler 1979: 300 ). Type locality: “ Nevada ” [probably vicinity of Lake Tahoe, Reno, or Carson City]. A. obsoleta ( Horn, 1880: 298 ) ( Liodes ) ; Brown 1937b: 201 ; Majka and Langor 2008: 386 (revalidation of name). Lectotype male in MCZC (designated by Wheeler 1979: 276 ). Type locality: none given, range stated as “New Hampshire to Virginia and Canada.” North American distribution: Canada : BC, MB, NB, NF, NS, ON, QC. USA : AL, AR, CT, FL, GA, IA, IL, KY, LA, MA, ME, MI, MN, MS, NC, NH, NY, OH, PA, SC, TN, VA, VT, WI, WV. Biology: Fungal associates Stephenson et al. 1994 ). Larva: Kilian (2005b) (as A . horni Wheeler ). = A. horni Wheeler, 1979: 276 (unnecessary replacement name for Liodes obsoleta Horn, 1880 , incorrectly thought to be preoccupied by Pallodes obsoletus F. E. Melsheimer, 1844 (now in Anogdus LeConte )). Biology: Bouchard (2001) . A. reticulonota Wheeler, 1979: 298 . Holotype male in MCZC. Type locality: White Mountains , Apache County , Arizona . North American distribution: USA : AZ, NM.