Dryinidae of the Oriental region (Hymenoptera: Chrysidoidea) Author Xu, Zaifu Author Olmi, Massimo Author He, Junhua text Zootaxa 2013 2013-02-15 3614 1 1 460 http://dx.doi.org/10.11646/zootaxa.3614.1.1 journal article 55852 10.11646/zootaxa.3614.1.1 aea75e24-7003-4c32-b29c-1e0512af5f96 1175-5326 5278372 E75E5224-20F1-431C-A7CB-9EE0D3F25118 37. Dryinus stantoni Ashmead, 1904 ( Plate 85A, B, D ) Dryinus stantoni Ashmead 1904: 134 ; Olmi 1984: 806 ; Guglielmino & Olmi 1997: 227 ; Olmi 1998f: 161 ; He & Xu 2002: 275 ; Guglielmino & Olmi 2006: 50 ; Xu et al. 2011 g (Suppl.): 16 ; Penati & Olmi 2012: 75 . Prodryinus stantoni (Ashmead) : Kieffer & Marshall 1906: 498 ; Kieffer 1914b: 53 . Paradryinus javanus R. Perkins 1912: 11 (synonymized by Olmi 1984 ); type locality: Pekalongan ( Java , Indonesia ). Lestodryinus luzonicus Kieffer 1913a: 189 (synonymized by Olmi 1984 ); type locality: Mt. Makiling ( Philippines ). Lestodryinus stantoni (Ashmead) : Fouts 1922: 629 . Neodryinus stantoni (Ashmead) : Baltazar 1966: 204 . Dryinus szepligettii Kieffer partim : Ponomarenko 1981b: 879 . Dryinus lycormae Yang 1994: 38 (synonymized by Olmi, 1998f ); type locality: Beijing ( China ); Olmi 1998f: 161 . Dryinus gutianshanensis Xu & He 1994c: 266 (synonymized by Olmi 1998f ); type locality: Mt. Gutianshan ( Zhejiang , China ); Olmi 1998f: 161 . Dryinus undatomarginis Xu & He 1998e: 309 ; type locality: Mt. Tianmushan ( Zhejiang , China ); syn. nov. Dryinus lii Xu & He 1998e: 310 (synonymized by He & Xu 2002 ); type locality: Hangzhou ( Zhejiang , China ). Dryinus wuyishanensis He & Xu 2002: 273 ; type locality: Mt. Wuyishan ( Fujian , China ); syn. nov. Description of Female. Fully winged; length 3.1–6.3 mm . Head black, except malar space, mandible and clypeus yellow; antenna testaceous, sometimes brownish, rarely with last segment whitish (in specimen from Thailand , 17°28.826'N 101°21.330'E , antenna brown, except segments 1–2 and 7–10 testaceous-whitish); mesosoma black, except lateral margins and posterior collar of pronotum reddish; occasionally posterior collar of pronotum black (in specimen from China , Liuxihe National Forest Park, pronotum brown, except posterior collar and lateral regions testaceous); metasoma brown-reddish; legs testaceous. Antenna clavate; antennal segments in following proportions: 10:5:26:12:9:8:7:7:5.5:9. Head ( Plate 85A ) flat, dull, granulated; face with few longitudinal keels; frontal line complete; occipital carina complete; POL = 4; OL = 2; OOL = 9; OPL = 1 (posterior ocelli very close the occipital carina, but not touching it); TL = 4; greatest breadth of posterior ocelli shorter than TL (3:4). Pronotum ( Plate 85B ) dull, hairy, granulated, with more or less numerous striae around and on disc; pronotum crossed by two transverse impressions, anterior impression slight and posterior one strong; pronotal tubercle not reaching tegula. Scutum dull, hairy, granulated, occasionally with posterior region reticulate rugose. Notauli usually incomplete, reaching about 0.5–0.7 length of scutum; in specimen from Thailand , 10 Km SW Chiang Mai , notauli complete and posteriorly separated. Scutellum and metanotum as sculptured as scutum. Propodeum rugose, without longitudinal keels on posterior surface. Fore wing with three dark transverse bands; distal part of stigmal vein much longer than proximal part. Protarsal segment 1 slightly longer than segment 4. Enlarged claw ( Plate 85D ) with one strong subdistal tooth and one row of 11 lamellae. Segment 5 of protarsus ( Plate 85D ) with two rows of about 27 lamellae; distal apex with about 14 lamellae. Tibial spurs 1/1/2. Description of Male. Fully winged; length 6.0– 7.5 mm . Head black, except mandible testaceous; antenna brown, except segment 1–2 testaceous; mesosoma black; metasoma black-brownish; legs yellow-brownish, except coxae brown-dark. Antenna filiform. Head dull, granulated; occipital carina complete; temple absent; POL = 6; OOL= 5.5; TL (visible inclining head) = 2.5; greatest breadth of the posterior ocelli shorter than POL (2.5:6). Scutum dull, granulated and rugose. Notauli complete, posteriorly separated; minimum distance between notauli longer than greatest breadth of posterior ocelli (7:2.5). Propodeum reticulate rugose. Fore wing hyaline, without dark transverse bands. Genitalia in fig. 4 in Yang, 1994 . Tibial spurs 1/1/2. Material examined. Type : holotype of D. stantoni : PHILIPPINES : Luzon Island , Manila ( USNM ); lectotype of P. javanus (designated by Olmi, 1982 ): INDONESIA : Java , Pekalongan , F. Muir , reared from Thanatodictya nymph ( BPBM ); holotype of L. luzonicus : PHILIPPINES : Luzon Island , Mt. Makiling ( USNM ); holotype of D. gutianshanensis : CHINA : Zhejiang , Kaihua County , Mt. Gutianshan , 29°03'N 118°02'E , 450 m , 21.VII.1992 , Hong Wu leg. ( ZJUC ); holotype of D. lii : CHINA : Zhejiang , Hangzhou City , 30°02'N 120°01'E , 5.VI.1992 , Xuexin Chen leg. ( ZJUC ); holotype of D. undatomarginis : CHINA : Zhejiang , Mt. Tianmushan , 30°02'N 119°07'E , 10–12.IX.1983 , Xingsheng Wang leg. ( ZJUC ); holotype of D. wuyishanensis : CHINA : Fujian , Mt. Wuyishan , 15.IX.1989 , Jiashe Wang leg. ( ZJUC ). Paralectotype of P. javanus : 1♀ , same locality label as lectotype ( BPBM ). The type specimens of Dryinus lycormae Yang (holotype ♀, ♂ allotype, 18♀♀ paratypes and 14♂ paratypes) were originally deposited in CAU. They were not examined, because apparently lost. They were labelled as follows: Beijing, Chi-kun Yang reared from Lycorma delicatula , 10.V.1975 , holotype ♀; 4.V.1975 , ♂ allotype; 1973–1980, paratypes. Other material. CAMBODIA : Koh Kong Prov. , 20 Km SE Koh Kong , 11°34'N 103°07'E , 50–300 m , V.2005 , Jendek & Sausa leg., 1♀ ( OLL ). CHINA : Fukien (= Fujian ), Shaowu, Mt. Wuyishan , Tachulan , 14.XII.1947 , T.C. Maa leg., 1♀ ( TARI ); Fujian , Mt. Wuyishan , 22–25.VIII.2007 , Jie Zeng leg., 1♀ ( SCAU ); Guangdong , Mt. Nankunshan , 8.VI.2002 , Zaifu Xu leg., 1♀ ; 2.VII–3.IX.2005 , Zaifu Xu leg., 2♀♀ ( ZJUC ); same locality label, 4–6.VI.2011 , Zaifu Xu leg., 1♀ ( SCAU ); same locality label, 17–18.VIII.2011 , Zaifu Xu leg., 1♀ ( SCAU ); Guangdong , Shixing County , Chebaling National Nature Reserve, 19.IV.2002 , Zaifu Xu leg., 1♀ ( ZJUC ); same locality label, 21–31.VIII.2003 , Jujian Chen leg., 3♀♀ ( ZJUC ); Guangdong , Yunan County , Mt. Tongledashan, 12–13.VIII.2003 , Zaifu Xu leg., 1♀ ( ZJUC ); Guangdong , Nanling National Nature Reserve, 10.VII.2005 , Zaifu Xu leg., 1♀ ( ZJUC ); Guangdong , Nanling National Nature Reserve, Protection Station, 16–17.V.2011 , YPT, Zaifu Xu leg., 1♀ ( SCAU ); Guangdong , Nanling National Nature Reserve, Arboretum, 10–12.IX.2011 , YPT, Zaifu Xu leg., 1♀ ( SCAU ); Guangdong , Conghua City, Liuxihe National Forest Park, 29–31.VIII.2005 , Zaifu Xu leg., 1♀ ( ZJUC ); Hainan , Mt. Wuzhishan , 16–18.V.2007 , Bin Xiao leg., 1♀ ( SCAU ); Henan , Runan County , Sanmenzha , 1983, reared by Jingfang Dong from Lycorma delicatula, many ♀♀ and ( ZJUC ); Hong Kong , Taipokau ( BPBM ); Hubei , Lichuan District , East Ridge , Suisapa , 1200–1500 m ( CAS ); Hunan , Shimen, Mt. Hupinghsan, Shimanzigou, 9.VII.2009 , Li Ma leg., 1♀ ; Taiwan , Nantou , Meifeng , 2150 m , 24–26.VI.1981 , K.S. Lin & W.S. Tang leg., 1♀ ( TARI ); Taiwan , Nantou , Wushe , 1150 m , 21–23.VI.1983 , K.S. Lin & S.C. Lin leg., 1♀ ( TARI ); same locality label, 25–28.VI.1985 , K.S. Lin leg., 2♀♀ ( TARI ); Taiwan , Nantou , Tungpu , 1200 m , 20–24.VI.1983 ( TARI ); Taiwan , Nantou , Lushan , 1000 m , 27–31.V.1980 ( TARI ); Taiwan , Pingtung , Shantimen , 28.III–1.IV.1981 ( TARI ); Taiwan , Taichung , Wanfeng Hill , X.1984 , K.S. Lin & K.C. Chou leg., 1♀ ( TARI ); Taiwan , Taitung , Chengkung , 16–17.IX.1982 , K.C. Chou & C.N. Lin leg., 1♀ ( TARI ); Taiwan , Taitung , Tawu , 12–14.II.1981 , L.Y. Chou & T. Lin leg., 1♀ ( TARI ); Taiwan , Taoyuan , Paling , 800 m , 3–5.V.1983 , K.C. Chou & C.C. Pan leg., 1♀ ( TARI ); Taiwan , Klapan ( BPBM ); Taiwan , Arisan , 10.X.1912 , I. Nitobe leg., 3♀♀ ( TARI ); Zhejiang , Mt. Tianmushan , 21–25.VII.2011 , Huayan Chen & Chengyuan Jin leg., 5♀♀ ( SCAU ); Zhejiang , Taishun County , Wuyanling Provincial Nature Reserve , 7.VII–5.VIII.2005 , Jingxian Liu and Jie Zeng leg., 1♀ ( ZJUC ). INDIA : Karnataka , Udupi District , Bhatkal , 27.V.2006 , Z. Kejval leg., 1♀ ( OLL ); Rajasthan , 15 Km N Udaipur , 24°45'N 73°43'E , 700 m , VII.2006 , Z. Kejval leg., 1♀ ( OLL ); Tamil Nadu , Coimbatore ( BMNH ); Tamil Nadu , 3 Km E Manjaler Dam , 15–18.X.1979 , J.S. Noyes leg., 1♀ ( BMNH ); Uttar Pradesh , Dehra Dun , 19.X.1979 , Z. Bouček leg., 1♀ ( BMNH ). INDONESIA : Borneo , Sidas , 8.VI.1960 ( OLL ); Maluku Islands , Halmahera Island, Jailolo District , Kampung Pasir Putih , 00°53'N 127°41'E ( USNM ); Maluku Islands , Halmahera (= Halmaheira ) Island, 1♀ ( MCSNG ); Sulawesi , Bantimurung , 05°01.041'S 119°40.622'E , 75 m , 30.VII–11.VIII.2005 , Malaise trap, 1♀ ( MNHN ). LAOS : Vientiane Prov. , Ban Van Eue ( BPBM ); Vientiane Prov. , Gi Sion village, De Tha Ngone ( BPBM ); Vientiane Prov. , Ban Van Heue , Phou-kow-kuei ( BPBM ); Xaignabouli (= Sayaboury ) Prov ., Sayaboury ( BPBM ). MALAYSIA : Malaya, Pahang District , King George V National Park , Kuala Tahang ( BPBM ); Malaya, Negeri Sembilan , Pasoh Forest Reserve ( AEIC ); Malaya , Negeri Sambilan , Port Dickson , IX.1944 , 1♀ ( BMNH ); Malaya , Selangor , Ulu Gombak Field Studies Centre of the University of Malaya ( DETAM ); Malaya , Selangor , Kuala Lumpur , University of Malaya , Rimba Ilma , 100 m , 14.VI.1990 , forest trail, J. Heraty leg., 1♀ ( DEUCR ); Malaya , Jahar , 15 Km NE Labis , near Pekan Aver Panas , 25–27.IV.1997 ( OLL ); Malaya , Pahang , Tanah Rata , Gunung Jasar , Cameron Highlands , 19–25.VI.1995 ( OLL ); Sabah , Sipitang , Mendolong ( ZIL ); Sabah , Kalabakan , Forest Camp ( BPBM ); Sabah , Sandakan ( AMNH ); Sabah , Singkor ( BPBM ); Sarawak , Ngungun , Ngemah ( PMA ); Sarawak , Gunung Mulu , II.1978 , subcanopy, D. Halloway leg., 1♀ ( BMNH ); same locality label, 17.IX–23.X.1977 , D. Hollis leg., 1♀ ( BMNH ); same locality label, 1♀ ( AMNH ). NEPAL : near Birganj , Lothar , 450', 1–12.IX.1967 ( CNC ). PHILIPPINES : Basilan Island ( USNM ); Biliran Island ( USNM ); Luzon Island , Manila , 1♀ ( MCZ ); Luzon Island , La Trinidad ( BPBM ); Luzon Island , Mt. Polis ( MNCNM ); Luzon Island , Imugan , Nueva Vizcaya ( BPBM ); Luzon Island , Mt. Makiling , 1♀ ( NHMLA ); Luzon Island , Laguna Prov. , Los Baños ( NHMLA , AMNH ); Luzon Island , Laguna Prov. , 4 Km E Los Baños , 14°10'N 121°15'E ( GCC ); Luzon Island , Rizal Mts. , Montalban , Wa –wa Dam ( BPBM ); Luzon Island , Mt. Banahao ( NHMLA ); Mindanao Island, Zamboanga ( BPBM , USNM ); Mindanao Island, Butuan ( USNM ); Mindanao Island, Dapitan ( USNM ); Palawan Island, Mantalingajan , Pinigisan ( ZMK ); Palawan Island, Mantalingajan , Tagembung ( ZMK ). SINGAPORE ( USNM ). SRI LANKA : Central Prov ., Kandy District , Anniewatte , 07°18'N 80°38'E , 1–8.V.1997 , K.V. Krombein , B.B. Norden , A.W. Norden & J.W. Norden leg., 2♀♀ ( USNM ); Central Prov ., Kandy District , Kandy ( BMNH ); Central Prov ., Matale District , Sigiriya , 07°57'N 80°46'E , 21.III.1999 , C.J. Burwell leg. ( QMC ); Central Prov ., Kandy District , IX.1909 , O.S.W. leg., 1♀ ( BMNH ); Eastern Prov ., Ampara District , Lahugala , 15.VI.1976 , K. Krombein leg., 1♀ ( USNM ); Sabaragamuwa Prov. , Rathnapura District , Gilimale Induruwa Jungle , 8.III.1979 , Karunaratne leg., 1♀ ( USNM ); Southern Prov ., Galle District , Kanneliya Jungle ( USNM ). THAILAND : Chantaburi Prov. , Soi Dao , Forest Insect Centre , 13°0'8.6''N 102°17'6.6''E , 197 m , I–II.2005 , 1♀ ( DEUK ); Chiang Mai Prov. , Chiangdao ( BPBM , AMNH ); Chiang Mai Prov. , 10 Km SW Chiang Mai , Doi Inthanon National Park , 1915 m ( PMA ); Chiang Mai Prov. , Maerim , 1–3.III.1996 , 19–20.II.1997 , 5–6.IV.1997 , 17.I.2000, 22–26.I.2000 , 8–10.II.2003 , R.A. Beaver leg. ( NHMLA , AMNH , RABC ); Loei Prov. , Phu Ruea National Park , office, 17°28.826'N 101°21.330'E , 860 m , Pan traps, 9–10.VII.2006 , Patikhom Tamtip leg., 1♀ ( QSBG ); Mae Hong Son Prov. , Soppong , 19°27'N 98°20'E , 1500 m , 7–12.V.1996 ( OLL ); Nakhon Nayok Prov. , Ban Na (= Banna ) ( BPBM ). VIET NAM : Dak Lak Prov. , M'Drak ( BPBM ); Gia Lai Prov. , Pleiku ( BPBM ). Palaearctic : CHINA : Beijing ( Yang , 1994 ). Hosts. Dictyopharidae and Fulgoridae ( Guglielmino & Olmi, 1997 , 2006 ): in Indonesia ( Java ): Thanatodictya sp. (Dictyopharidae) (R. Perkins, 1912 : as P. javanus ); in China ( Beijing , Henan ): Lycorma delicatula (White) (Fulgoridae) ( Yang, 1994 : as D. lycormae ). Natural enemies . In Java ( Indonesia ) this species may be parasitized by Cheiloneurus javanus R. Perkins, 1912 (Encyrtidae) (R. Perkins, 1912 ). Distribution. China ( Zhejiang , Fujian , Beijing , Guangdong , Hainan , Hong Kong , Hunan, Taiwan), Philippines (Luzon, Basilan , Biliran , Mindanao, Palawan ), Indonesia ( Java , Borneo, Maluku , Sulawesi ), Cambodia ( Koh Kong ), India ( Karnataka , Rajasthan , Tamil Nadu , Uttar Pradesh ), Laos ( Vientiane , Sayaboury), Malaysia (Malaya, Sabah , Sarawak ), Nepal (Lothar), Sri Lanka (Kandy, Matale, Ampara, Rathnapura, Galle), Thailand ( Chantaburi , Chiang Mai , Loei , Mae Hong Son , Nakhon Nayok ), Viet Nam ( Dak Lak , Gia Lai ). Remarks. Dryinus undatomarginis Xu & He and D. wuyishanensis He & Xu were synonymized with D. stantoni following the study of the types of both species and comparison with the type and other female specimens of D. stantoni . No significative differences were observed.