Dryinidae of the Oriental region (Hymenoptera: Chrysidoidea)
Author
Xu, Zaifu
Author
Olmi, Massimo
Author
He, Junhua
text
Zootaxa
2013
2013-02-15
3614
1
1
460
http://dx.doi.org/10.11646/zootaxa.3614.1.1
journal article
55852
10.11646/zootaxa.3614.1.1
aea75e24-7003-4c32-b29c-1e0512af5f96
1175-5326
5278372
E75E5224-20F1-431C-A7CB-9EE0D3F25118
37.
Dryinus stantoni
Ashmead, 1904
(
Plate 85A, B, D
)
Dryinus stantoni
Ashmead 1904: 134
;
Olmi 1984: 806
;
Guglielmino & Olmi 1997: 227
;
Olmi 1998f: 161
;
He & Xu 2002: 275
;
Guglielmino & Olmi 2006: 50
;
Xu
et al.
2011
g (Suppl.): 16
;
Penati & Olmi 2012: 75
.
Prodryinus stantoni
(Ashmead)
:
Kieffer & Marshall 1906: 498
;
Kieffer 1914b: 53
.
Paradryinus javanus
R.
Perkins 1912: 11
(synonymized by
Olmi 1984
);
type
locality: Pekalongan (
Java
,
Indonesia
).
Lestodryinus luzonicus
Kieffer 1913a: 189
(synonymized by
Olmi 1984
);
type
locality: Mt. Makiling (
Philippines
).
Lestodryinus stantoni
(Ashmead)
:
Fouts 1922: 629
.
Neodryinus stantoni
(Ashmead)
:
Baltazar 1966: 204
.
Dryinus szepligettii
Kieffer
partim
:
Ponomarenko 1981b: 879
.
Dryinus lycormae
Yang 1994: 38
(synonymized by
Olmi, 1998f
);
type
locality:
Beijing
(
China
);
Olmi 1998f: 161
.
Dryinus gutianshanensis
Xu & He 1994c: 266
(synonymized by
Olmi 1998f
);
type
locality: Mt. Gutianshan (
Zhejiang
,
China
);
Olmi 1998f: 161
.
Dryinus undatomarginis
Xu & He 1998e: 309
;
type
locality: Mt. Tianmushan (
Zhejiang
,
China
);
syn. nov.
Dryinus lii
Xu & He 1998e: 310
(synonymized by
He & Xu 2002
);
type
locality: Hangzhou (
Zhejiang
,
China
).
Dryinus wuyishanensis
He & Xu 2002: 273
;
type
locality: Mt. Wuyishan (
Fujian
,
China
);
syn. nov.
Description of Female.
Fully winged; length
3.1–6.3 mm
. Head black, except malar space, mandible and clypeus yellow; antenna testaceous, sometimes brownish, rarely with last segment whitish (in specimen from
Thailand
,
17°28.826'N
101°21.330'E
, antenna brown, except segments 1–2 and 7–10 testaceous-whitish); mesosoma black, except lateral margins and posterior collar of pronotum reddish; occasionally posterior collar of pronotum black (in specimen from
China
, Liuxihe National Forest Park, pronotum brown, except posterior collar and lateral regions testaceous); metasoma brown-reddish; legs testaceous. Antenna clavate; antennal segments in following proportions: 10:5:26:12:9:8:7:7:5.5:9. Head (
Plate 85A
) flat, dull, granulated; face with few longitudinal keels; frontal line complete; occipital carina complete; POL = 4; OL = 2; OOL = 9; OPL = 1 (posterior ocelli very close the occipital carina, but not touching it); TL = 4; greatest breadth of posterior ocelli shorter than TL (3:4). Pronotum (
Plate 85B
) dull, hairy, granulated, with more or less numerous striae around and on disc; pronotum crossed by two transverse impressions, anterior impression slight and posterior one strong; pronotal tubercle not reaching tegula. Scutum dull, hairy, granulated, occasionally with posterior region reticulate rugose. Notauli usually incomplete, reaching about 0.5–0.7 length of scutum; in specimen from
Thailand
, 10 Km SW
Chiang Mai
, notauli complete and posteriorly separated. Scutellum and metanotum as sculptured as scutum. Propodeum rugose, without longitudinal keels on posterior surface. Fore wing with three dark transverse bands; distal part of stigmal vein much longer than proximal part. Protarsal segment 1 slightly longer than segment 4. Enlarged claw (
Plate 85D
) with one strong subdistal tooth and one row of 11 lamellae. Segment 5 of protarsus (
Plate 85D
) with two rows of about 27 lamellae; distal apex with about 14 lamellae. Tibial spurs 1/1/2.
Description of Male.
Fully winged; length 6.0–
7.5 mm
. Head black, except mandible testaceous; antenna brown, except segment 1–2 testaceous; mesosoma black; metasoma black-brownish; legs yellow-brownish, except coxae brown-dark. Antenna filiform. Head dull, granulated; occipital carina complete; temple absent; POL = 6; OOL= 5.5; TL (visible inclining head) = 2.5; greatest breadth of the posterior ocelli shorter than POL (2.5:6). Scutum dull, granulated and rugose. Notauli complete, posteriorly separated; minimum distance between notauli longer than greatest breadth of posterior ocelli (7:2.5). Propodeum reticulate rugose. Fore wing hyaline, without dark transverse bands. Genitalia in fig.
4 in
Yang, 1994
. Tibial spurs 1/1/2.
Material examined. Type
:
♀
holotype
of
D. stantoni
:
PHILIPPINES
:
Luzon Island
,
Manila
(
USNM
);
♀
lectotype
of
P. javanus
(designated by
Olmi, 1982
):
INDONESIA
:
Java
,
Pekalongan
,
F. Muir
, reared from
Thanatodictya
nymph (
BPBM
);
♀
holotype of
L. luzonicus
:
PHILIPPINES
:
Luzon Island
,
Mt. Makiling
(
USNM
);
♀
holotype of
D. gutianshanensis
:
CHINA
:
Zhejiang
,
Kaihua County
,
Mt. Gutianshan
,
29°03'N
118°02'E
,
450 m
,
21.VII.1992
,
Hong Wu
leg. (
ZJUC
);
♀
holotype of
D. lii
:
CHINA
:
Zhejiang
,
Hangzhou City
,
30°02'N
120°01'E
,
5.VI.1992
,
Xuexin Chen
leg. (
ZJUC
);
♀
holotype of
D. undatomarginis
:
CHINA
:
Zhejiang
,
Mt. Tianmushan
,
30°02'N
119°07'E
,
10–12.IX.1983
,
Xingsheng Wang
leg. (
ZJUC
);
♀
holotype of
D. wuyishanensis
:
CHINA
:
Fujian
,
Mt. Wuyishan
,
15.IX.1989
,
Jiashe Wang
leg. (
ZJUC
).
Paralectotype
of
P. javanus
:
1♀
, same locality label as
lectotype
(
BPBM
).
The
type
specimens
of
Dryinus lycormae
Yang
(holotype ♀, ♂ allotype, 18♀♀ paratypes and 14♂ paratypes) were originally deposited in CAU. They were not examined, because apparently lost. They were labelled as follows: Beijing, Chi-kun Yang reared from
Lycorma delicatula
,
10.V.1975
, holotype ♀;
4.V.1975
, ♂ allotype; 1973–1980, paratypes.
Other material.
CAMBODIA
:
Koh Kong Prov.
, 20
Km
SE
Koh Kong
,
11°34'N
103°07'E
,
50–300 m
,
V.2005
,
Jendek & Sausa
leg.,
1♀
(
OLL
).
CHINA
: Fukien (=
Fujian
), Shaowu,
Mt. Wuyishan
,
Tachulan
,
14.XII.1947
,
T.C. Maa
leg.,
1♀
(
TARI
);
Fujian
,
Mt. Wuyishan
,
22–25.VIII.2007
,
Jie Zeng
leg.,
1♀
(
SCAU
);
Guangdong
,
Mt. Nankunshan
,
8.VI.2002
,
Zaifu Xu
leg.,
1♀
;
2.VII–3.IX.2005
,
Zaifu Xu
leg.,
2♀♀
(
ZJUC
);
same locality label,
4–6.VI.2011
,
Zaifu Xu
leg.,
1♀
(
SCAU
);
same locality label,
17–18.VIII.2011
,
Zaifu Xu
leg.,
1♀
(
SCAU
);
Guangdong
,
Shixing County
, Chebaling National Nature Reserve,
19.IV.2002
,
Zaifu Xu
leg.,
1♀
(
ZJUC
);
same locality label,
21–31.VIII.2003
,
Jujian Chen
leg.,
3♀♀
(
ZJUC
);
Guangdong
,
Yunan County
, Mt. Tongledashan,
12–13.VIII.2003
,
Zaifu Xu
leg.,
1♀
(
ZJUC
);
Guangdong
, Nanling National Nature Reserve,
10.VII.2005
,
Zaifu Xu
leg.,
1♀
(
ZJUC
);
Guangdong
, Nanling National Nature Reserve, Protection Station,
16–17.V.2011
, YPT,
Zaifu Xu
leg.,
1♀
(
SCAU
);
Guangdong
, Nanling National Nature Reserve, Arboretum,
10–12.IX.2011
, YPT,
Zaifu Xu
leg.,
1♀
(
SCAU
);
Guangdong
, Conghua City, Liuxihe National Forest Park,
29–31.VIII.2005
,
Zaifu Xu
leg.,
1♀
(
ZJUC
);
Hainan
,
Mt. Wuzhishan
,
16–18.V.2007
,
Bin Xiao
leg.,
1♀
(
SCAU
);
Henan
,
Runan County
,
Sanmenzha
, 1983, reared by
Jingfang Dong from
Lycorma
delicatula, many
♀♀
and
♂
(
ZJUC
);
Hong Kong
,
Taipokau
(
BPBM
);
Hubei
,
Lichuan District
,
East Ridge
,
Suisapa
,
1200–1500 m
(
CAS
);
Hunan
, Shimen, Mt. Hupinghsan, Shimanzigou,
9.VII.2009
, Li Ma leg.,
1♀
;
Taiwan
,
Nantou
,
Meifeng
,
2150 m
,
24–26.VI.1981
,
K.S. Lin & W.S. Tang
leg.,
1♀
(
TARI
);
Taiwan
,
Nantou
,
Wushe
,
1150 m
,
21–23.VI.1983
,
K.S. Lin & S.C. Lin
leg.,
1♀
(
TARI
);
same locality label,
25–28.VI.1985
,
K.S. Lin
leg.,
2♀♀
(
TARI
);
Taiwan
,
Nantou
,
Tungpu
,
1200 m
,
20–24.VI.1983
(
TARI
);
Taiwan
,
Nantou
,
Lushan
,
1000 m
,
27–31.V.1980
(
TARI
);
Taiwan
,
Pingtung
,
Shantimen
,
28.III–1.IV.1981
(
TARI
);
Taiwan
,
Taichung
,
Wanfeng Hill
,
X.1984
,
K.S. Lin
&
K.C. Chou
leg.,
1♀
(
TARI
);
Taiwan
,
Taitung
,
Chengkung
,
16–17.IX.1982
,
K.C. Chou & C.N. Lin
leg.,
1♀
(
TARI
);
Taiwan
,
Taitung
,
Tawu
,
12–14.II.1981
,
L.Y. Chou & T. Lin
leg.,
1♀
(
TARI
);
Taiwan
,
Taoyuan
,
Paling
,
800 m
,
3–5.V.1983
,
K.C. Chou
&
C.C. Pan
leg.,
1♀
(
TARI
);
Taiwan
,
Klapan
(
BPBM
);
Taiwan
,
Arisan
,
10.X.1912
,
I. Nitobe
leg.,
3♀♀
(
TARI
);
Zhejiang
,
Mt. Tianmushan
,
21–25.VII.2011
,
Huayan Chen & Chengyuan Jin
leg.,
5♀♀
(
SCAU
);
Zhejiang
,
Taishun County
,
Wuyanling Provincial Nature Reserve
,
7.VII–5.VIII.2005
,
Jingxian Liu
and
Jie Zeng
leg.,
1♀
(
ZJUC
).
INDIA
:
Karnataka
,
Udupi District
,
Bhatkal
,
27.V.2006
,
Z. Kejval
leg.,
1♀
(
OLL
);
Rajasthan
, 15
Km
N
Udaipur
,
24°45'N
73°43'E
,
700 m
,
VII.2006
,
Z. Kejval
leg.,
1♀
(
OLL
);
Tamil Nadu
,
Coimbatore
(
BMNH
);
Tamil Nadu
, 3
Km
E
Manjaler Dam
,
15–18.X.1979
,
J.S. Noyes
leg.,
1♀
(
BMNH
);
Uttar Pradesh
,
Dehra Dun
,
19.X.1979
,
Z. Bouček
leg.,
1♀
(
BMNH
).
INDONESIA
:
Borneo
,
Sidas
,
8.VI.1960
(
OLL
);
Maluku Islands
,
Halmahera
Island,
Jailolo District
,
Kampung Pasir Putih
,
00°53'N
127°41'E
(
USNM
);
Maluku Islands
,
Halmahera
(=
Halmaheira
) Island,
1♀
(
MCSNG
);
Sulawesi
,
Bantimurung
,
05°01.041'S
119°40.622'E
,
75 m
,
30.VII–11.VIII.2005
,
Malaise
trap,
1♀
(
MNHN
).
LAOS
:
Vientiane Prov.
,
Ban Van Eue
(
BPBM
);
Vientiane Prov.
,
Gi Sion
village,
De Tha Ngone
(
BPBM
);
Vientiane Prov.
,
Ban Van Heue
, Phou-kow-kuei (
BPBM
);
Xaignabouli
(=
Sayaboury
)
Prov
.,
Sayaboury
(
BPBM
).
MALAYSIA
: Malaya,
Pahang District
,
King George
V
National Park
,
Kuala Tahang
(
BPBM
);
Malaya,
Negeri Sembilan
,
Pasoh Forest Reserve
(
AEIC
);
Malaya
,
Negeri Sambilan
,
Port Dickson
,
IX.1944
,
1♀
(
BMNH
);
Malaya
,
Selangor
,
Ulu Gombak Field Studies Centre of the University
of
Malaya
(
DETAM
);
Malaya
,
Selangor
,
Kuala Lumpur
,
University
of
Malaya
,
Rimba Ilma
,
100 m
,
14.VI.1990
, forest trail,
J. Heraty
leg.,
1♀
(
DEUCR
);
Malaya
,
Jahar
, 15
Km
NE
Labis
, near
Pekan
Aver Panas
,
25–27.IV.1997
(
OLL
);
Malaya
,
Pahang
,
Tanah Rata
,
Gunung Jasar
,
Cameron Highlands
,
19–25.VI.1995
(
OLL
);
Sabah
,
Sipitang
,
Mendolong
(
ZIL
);
Sabah
,
Kalabakan
,
Forest Camp
(
BPBM
);
Sabah
,
Sandakan
(
AMNH
);
Sabah
,
Singkor
(
BPBM
);
Sarawak
,
Ngungun
,
Ngemah
(
PMA
);
Sarawak
,
Gunung Mulu
,
II.1978
, subcanopy,
D. Halloway
leg.,
1♀
(
BMNH
);
same locality label,
17.IX–23.X.1977
,
D. Hollis
leg.,
1♀
(
BMNH
);
same locality label,
1♀
(
AMNH
).
NEPAL
: near
Birganj
,
Lothar
, 450',
1–12.IX.1967
(
CNC
).
PHILIPPINES
:
Basilan
Island (
USNM
);
Biliran
Island (
USNM
);
Luzon Island
,
Manila
,
1♀
(
MCZ
);
Luzon Island
,
La
Trinidad
(
BPBM
);
Luzon Island
,
Mt. Polis
(
MNCNM
);
Luzon Island
,
Imugan
,
Nueva
Vizcaya
(
BPBM
);
Luzon Island
,
Mt. Makiling
,
1♀
(
NHMLA
);
Luzon Island
,
Laguna Prov.
,
Los Baños
(
NHMLA
,
AMNH
);
Luzon Island
,
Laguna Prov.
, 4
Km
E
Los Baños
,
14°10'N
121°15'E
(
GCC
);
Luzon Island
,
Rizal Mts.
,
Montalban
,
Wa
–wa
Dam
(
BPBM
);
Luzon Island
,
Mt. Banahao
(
NHMLA
);
Mindanao Island,
Zamboanga
(
BPBM
,
USNM
);
Mindanao Island,
Butuan
(
USNM
);
Mindanao Island,
Dapitan
(
USNM
);
Palawan
Island,
Mantalingajan
,
Pinigisan
(
ZMK
);
Palawan
Island,
Mantalingajan
,
Tagembung
(
ZMK
).
SINGAPORE
(
USNM
).
SRI LANKA
: Central
Prov
.,
Kandy District
,
Anniewatte
,
07°18'N
80°38'E
,
1–8.V.1997
,
K.V. Krombein
,
B.B. Norden
,
A.W. Norden & J.W. Norden
leg.,
2♀♀
(
USNM
);
Central
Prov
.,
Kandy District
,
Kandy
(
BMNH
);
Central
Prov
.,
Matale District
,
Sigiriya
,
07°57'N
80°46'E
,
21.III.1999
,
C.J. Burwell
leg. (
QMC
);
Central
Prov
.,
Kandy District
,
IX.1909
, O.S.W. leg.,
1♀
(
BMNH
);
Eastern
Prov
.,
Ampara District
,
Lahugala
,
15.VI.1976
,
K. Krombein
leg.,
1♀
(
USNM
);
Sabaragamuwa Prov.
,
Rathnapura District
,
Gilimale Induruwa Jungle
,
8.III.1979
,
Karunaratne
leg.,
1♀
(
USNM
);
Southern
Prov
.,
Galle District
,
Kanneliya Jungle
(
USNM
).
THAILAND
:
Chantaburi Prov.
,
Soi Dao
,
Forest
Insect
Centre
,
13°0'8.6''N
102°17'6.6''E
,
197 m
,
I–II.2005
,
1♀
(
DEUK
);
Chiang Mai Prov.
,
Chiangdao
(
BPBM
,
AMNH
);
Chiang Mai Prov.
, 10
Km
SW
Chiang Mai
,
Doi Inthanon
National Park
,
1915 m
(
PMA
);
Chiang Mai Prov.
,
Maerim
,
1–3.III.1996
,
19–20.II.1997
,
5–6.IV.1997
, 17.I.2000,
22–26.I.2000
,
8–10.II.2003
,
R.A.
Beaver
leg. (
NHMLA
,
AMNH
,
RABC
);
Loei Prov.
,
Phu Ruea
National Park
, office,
17°28.826'N
101°21.330'E
,
860 m
,
Pan
traps,
9–10.VII.2006
,
Patikhom Tamtip
leg.,
1♀
(
QSBG
);
Mae Hong Son Prov.
,
Soppong
,
19°27'N
98°20'E
,
1500 m
,
7–12.V.1996
(
OLL
);
Nakhon Nayok Prov.
,
Ban Na
(=
Banna
) (
BPBM
).
VIET NAM
:
Dak
Lak Prov.
,
M'Drak
(
BPBM
);
Gia Lai Prov.
, Pleiku (
BPBM
).
Palaearctic
:
CHINA
:
Beijing
(
Yang
, 1994
).
Hosts.
Dictyopharidae
and
Fulgoridae
(
Guglielmino & Olmi, 1997
,
2006
): in
Indonesia
(
Java
):
Thanatodictya
sp. (Dictyopharidae)
(R.
Perkins, 1912
: as
P. javanus
); in
China
(
Beijing
,
Henan
):
Lycorma delicatula
(White) (Fulgoridae)
(
Yang, 1994
: as
D. lycormae
).
Natural enemies
. In
Java
(
Indonesia
) this species may be parasitized by
Cheiloneurus javanus
R.
Perkins, 1912
(Encyrtidae)
(R.
Perkins, 1912
).
Distribution.
China
(
Zhejiang
,
Fujian
,
Beijing
,
Guangdong
,
Hainan
,
Hong Kong
, Hunan, Taiwan),
Philippines
(Luzon,
Basilan
,
Biliran
, Mindanao,
Palawan
),
Indonesia
(
Java
, Borneo,
Maluku
,
Sulawesi
),
Cambodia
(
Koh Kong
),
India
(
Karnataka
,
Rajasthan
,
Tamil Nadu
,
Uttar Pradesh
),
Laos
(
Vientiane
, Sayaboury),
Malaysia
(Malaya,
Sabah
,
Sarawak
),
Nepal
(Lothar),
Sri Lanka
(Kandy, Matale, Ampara, Rathnapura, Galle),
Thailand
(
Chantaburi
,
Chiang Mai
,
Loei
,
Mae Hong Son
,
Nakhon Nayok
),
Viet Nam
(
Dak Lak
,
Gia Lai
).
Remarks.
Dryinus undatomarginis
Xu & He
and
D. wuyishanensis
He & Xu
were synonymized with
D. stantoni
following the study of the
types
of both species and comparison with the
type
and other female specimens of
D. stantoni
. No significative differences were observed.