<document id="AF3D13AE13C09BA6FFBF18017D9E9B62" ID-CLB-Dataset="21174" ID-DOI="10.11646/zootaxa.4629.3.1" ID-GBIF-Dataset="9419bc9b-dbe4-426c-a34e-f0034848ff99" ID-ISSN="1175-5326" ID-Zenodo-Dep="3995508" ID-ZooBank="1D47B452-0A40-4BC5-9BBA-C06DEB298C9A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1598059654494" checkinUser="felipe" docAuthor="Silva, Fernando A. B. &amp; Valois, Marcely" docDate="2019" docId="9D12E5573A56FF9195F1F8D9FB90C668" docLanguage="en" docName="zootaxa.4629.3.1.pdf" docOrigin="Zootaxa 4629 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scybalocanthon batesi Vaz-de-Mello &amp; Silva 2017" docType="treatment" docVersion="11" lastPageNumber="316" masterDocId="612B9D2F3A59FF9E9566F840FFF1C516" masterDocTitle="A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini)" masterLastPageNumber="341" masterPageNumber="301" pageNumber="316" updateTime="1732598001296" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0C880A187F5D679F4DE1822127862873">A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini)</mods:title>
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<mods:namePart id="2F998EA75237B085A347E4C0F2FA737F">Silva, Fernando A. B.</mods:namePart>
<mods:affiliation id="8E9B2AD88217D6071C2A11B2336699D4">Universidade Federal do Pará, Instituto de Ciências Biológicas, setor de Zoologia, Campus Belém. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil. E-mail: fernandoabsilva @ yahoo. com. br</mods:affiliation>
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<mods:namePart id="99369839828237BC2AC3AFBBFB278783">Valois, Marcely</mods:namePart>
<mods:affiliation id="1C19A27F7E695D5DC18C0B8BC0D26645">Programa de Pós-graduação em Zoologia, Universidade Federal do Pará / Museu Paraense Emílio Goeldi. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil.</mods:affiliation>
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<treatment id="9D12E5573A56FF9195F1F8D9FB90C668" ID-DOI="http://doi.org/10.5281/zenodo.4335186" ID-GBIF-Taxon="167293216" ID-Zenodo-Dep="4335186" LSID="urn:lsid:plazi:treatment:9D12E5573A56FF9195F1F8D9FB90C668" httpUri="http://treatment.plazi.org/id/9D12E5573A56FF9195F1F8D9FB90C668" lastPageNumber="316" pageId="15" pageNumber="316" scope_family="Scarabaeidae" scope_order="Coleoptera" scope_subFamily="Scarabaeinae" scope_tribe="Deltochilini">
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<heading id="4E4CE32D3A56FF9195F1F8D9FCF2C5A5" bold="true" box="[151,771,153,180]" fontSize="11" level="1" pageId="15" pageNumber="316" reason="1">
<emphasis id="27CF88533A56FF9195F1F8D9FCF2C5A5" bold="true" box="[151,771,153,180]" pageId="15" pageNumber="316">
<taxonomicName id="D2BB2FC23A56FF9195F1F8D9FE57C5A2" ID-CoL="4W3LD" authorityName="Vaz-de-Mello &amp; Silva" authorityYear="2017" box="[151,422,153,180]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="batesi">
<emphasis id="27CF88533A56FF9195F1F8D9FE57C5A2" bold="true" box="[151,422,153,180]" italics="true" pageId="15" pageNumber="316">Scybalocanthon batesi</emphasis>
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<bibRefCitation id="712A29B03A56FF9194C8F8DAFCF2C5A5" author="Vaz-de-Mello, F. Z. &amp; Silva, F. A. B." box="[430,771,153,180]" pageId="15" pageNumber="316" pagination="142 - 146" refId="ref48256" refString="Vaz-de-Mello, F. Z. &amp; Silva, F. A. B. (2017) A new species of the genus Scybalocanthon (Coleoptera, Scarabaeidae, Scarabaeinae) from the Atlantic rainforest, with an identification key to the species from South America south of the Amazon basin. Zootaxa, 4300 (1), 142 - 146. https: // doi. org / 10.11646 / zootaxa. 4300.1.9" type="journal article" year="2017">Vaz-de-Mello &amp; Silva, 2017</bibRefCitation>
</emphasis>
</heading>
</paragraph>
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<paragraph id="150454413A56FF9195F1F8FDFEC2C5CE" blockId="15.[151,771,153,216]" box="[151,307,189,216]" pageId="15" pageNumber="316">
(
<figureCitation id="8D8048C43A56FF9195F9F8FDFF0EC5C1" box="[159,255,189,215]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="15" pageNumber="316">Figs. 1E</figureCitation>
, 
<figureCitation id="8D8048C43A56FF91946AF8FDFEDAC5CE" box="[268,299,189,216]" captionStart="FIGURE 6" captionStartId="17.[151,250,1784,1809]" captionTargetBox="[179,1376,186,1734]" captionTargetId="figure@17.[174,1414,181,1759]" captionTargetPageId="17" captionText="FIGURE 6. Known distribution of species of Scybalocanthon." figureDoi="http://doi.org/10.5281/zenodo.3995522" httpUri="https://zenodo.org/record/3995522/files/figure.png" pageId="15" pageNumber="316">6F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A56FF9195F1F942FC43C40C" box="[151,946,257,282]" pageId="15" pageNumber="316" type="reference_group">
<paragraph id="150454413A56FF9195F1F942FC43C40C" blockId="15.[151,946,257,282]" box="[151,946,257,282]" pageId="15" pageNumber="316">
<treatmentCitationGroup id="35AB736F3A56FF9195F1F942FC43C40C" box="[151,946,257,282]" pageId="15" pageNumber="316">
<taxonomicName id="D2BB2FC23A56FF9195F1F942FE8DC40C" ID-CoL="4W3LD" authorityName="Vaz-de-Mello &amp; Silva" authorityYear="2017" box="[151,380,258,282]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="batesi">
<emphasis id="27CF88533A56FF9195F1F942FE8DC40C" box="[151,380,258,282]" italics="true" pageId="15" pageNumber="316">Scybalocanthon batesi</emphasis>
</taxonomicName>
<treatmentCitation id="941A72503A56FF9194E4F941FD24C40C" ID-DOI="http://doi.org/10.5281/zenodo.6001662" ID-Zenodo-Dep="6001662" author="Vaz-de-Mello, F. Z. &amp; Silva, F. A. B." box="[386,725,257,282]" httpUri="http://treatment.plazi.org/id/03AF87EAFFC4FFF8FF393B47FE8802E4" page="143" pageId="15" pageNumber="316" year="2017">
<bibRefCitation id="712A29B03A56FF9194E4F941FD24C40C" author="Vaz-de-Mello, F. Z. &amp; Silva, F. A. B." box="[386,725,257,282]" pageId="15" pageNumber="316" pagination="142 - 146" refId="ref48256" refString="Vaz-de-Mello, F. Z. &amp; Silva, F. A. B. (2017) A new species of the genus Scybalocanthon (Coleoptera, Scarabaeidae, Scarabaeinae) from the Atlantic rainforest, with an identification key to the species from South America south of the Amazon basin. Zootaxa, 4300 (1), 142 - 146. https: // doi. org / 10.11646 / zootaxa. 4300.1.9" type="journal article" year="2017">Vaz-de-Mello &amp; Silva, 2017: 143</bibRefCitation>
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(original description)
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</paragraph>
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<subSubSection id="5DA107CA3A56FF9195F1F907FE99C4D8" pageId="15" pageNumber="316" type="diagnosis">
<paragraph id="150454413A56FF9195F1F907FE99C4D8" blockId="15.[151,1437,327,895]" pageId="15" pageNumber="316">
<emphasis id="27CF88533A56FF9195F1F907FEE3C474" bold="true" box="[151,274,327,354]" pageId="15" pageNumber="316">Diagnosis.</emphasis>
<taxonomicName id="D2BB2FC23A56FF91947EF908FDE4C474" authorityName="Vaz-de-Mello &amp; Silva" authorityYear="2017" box="[280,533,328,354]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="batesi">
<emphasis id="27CF88533A56FF91947EF908FDE4C474" box="[280,533,328,354]" italics="true" pageId="15" pageNumber="316">Scybalocanthon batesi</emphasis>
</taxonomicName>
can be easily differentiated from all remaining species in the genus by the elytra having a yellow horizontal band on the central portion occupying approximately one-third of the elytral length (
<figureCitation id="8D8048C43A56FF919009F92CFF49C4BC" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="15" pageNumber="316">Fig. 1E</figureCitation>
). Dark brown or black metepisternae. Anterior and posterior edge of the pronotum with a triangular black spot on middle portion.
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A56FF9195A1F997FA9EC605" pageId="15" pageNumber="316" type="description">
<paragraph id="150454413A56FF9195A1F997FA6CC7B0" blockId="15.[151,1437,327,895]" pageId="15" pageNumber="316">
<emphasis id="27CF88533A56FF9195A1F997FE6CC4E4" bold="true" box="[199,413,471,498]" pageId="15" pageNumber="316">Description. Body</emphasis>
. Oval, lateral edges rounded. Surface opaque, head and elytra microgranulate. 
<emphasis id="27CF88533A56FF91906FF997FABCC4E4" bold="true" box="[1289,1357,471,498]" pageId="15" pageNumber="316">Color</emphasis>
. Head, anterior and posterior edge of pronotum, anterior and posterior portions of elytra, prosternum, internal portion of hypomera, mesoventrite, mesoepisternae, lateral portions of metaventrite, metepisternae, procoxae, trochanters, basal and apical parts of femora, tibiae, and mesotarsomeres and metatarsomeres dark brown or black. Other parts of the pronotum, horizontal band at the middle portion of elytra, pygidium, most of the hypomera, central portion of metaventrite, abdomen, most of the mesocoxa and metacoxa, and femur (except the extremities) yellowish brown.
</paragraph>
<paragraph id="150454413A56FF9195F1FAF0FA9EC605" blockId="15.[151,1437,327,895]" pageId="15" pageNumber="316">
<emphasis id="27CF88533A56FF9195F1FAF0FF1CC7DC" bold="true" box="[151,237,688,714]" pageId="15" pageNumber="316">Length</emphasis>
. 
<quantity id="D243F9A43A56FF91959DFAF0FECEC7DD" box="[251,319,688,715]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="316" unit="mm" value="8.0">8 mm</quantity>
. 
<emphasis id="27CF88533A56FF91942FFAF0FE53C7DC" bold="true" box="[329,418,688,714]" pageId="15" pageNumber="316">Thorax</emphasis>
. Anterior angles of pronotum acute (approximately 85°). Lateral margin irregularly curved outward, forming an obtuse angle at the middle portion. 
<emphasis id="27CF88533A56FF91967CFA94FC97C7F8" bold="true" box="[794,870,724,750]" pageId="15" pageNumber="316">Elytra</emphasis>
. Striae thin, with elliptical punctures. Eighth stria with a thin carina at the anterior portion. Basal third of the eighth stria inconspicuous (except the basal carina).
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A56FF9195A1FB5CFE46C64C" pageId="15" pageNumber="316" type="materials_examined">
<paragraph id="150454413A56FF9195A1FB5CFE46C64C" blockId="15.[151,1437,327,895]" pageId="15" pageNumber="316">
<emphasis id="27CF88533A56FF9195A1FB5CFE84C620" bold="true" box="[199,373,795,822]" pageId="15" pageNumber="316">Type material.</emphasis>
<materialsCitation id="A5D35E1C3A56FF91941AFB5CFBC5C621" ID-GBIF-Occurrence="2847261501" box="[380,1076,796,823]" collectionCode="MNHN" collectorName="H. W. Bates" country="Brazil" location="BAHIA" pageId="15" pageNumber="316" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="holotype">
<emphasis id="27CF88533A56FF91941AFB5CFE01C620" bold="true" box="[380,496,796,822]" pageId="15" pageNumber="316">
<typeStatus id="CA00EAE33A56FF91941AFB5CFE1AC620" box="[380,491,796,822]" pageId="15" pageNumber="316" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="6DAC14D13A56FF919491FB5CFD90C620" box="[503,609,796,822]" name="Brazil" pageId="15" pageNumber="316">BRAZIL</collectingCountry>
: 
<collectingRegion id="D77F9AA33A56FF91970DFB5CFD36C620" box="[619,711,796,822]" country="Brazil" name="Bahia" pageId="15" pageNumber="316">BAHIA</collectingRegion>
, 1892, 
<collectorName id="B84E31973A56FF91967EFB5DFC66C621" box="[792,919,797,823]" pageId="15" pageNumber="316">H.W. Bates</collectorName>
(
<specimenCount id="03BD9FC83A56FF9196CCFB5CFC3BC620" box="[938,970,796,822]" pageId="15" pageNumber="316" type="female">1♀</specimenCount>
<collectionCode id="73AACC843A56FF9196B7FB5DFBDDC621" box="[977,1068,797,823]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="316">MNHN</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="A5D35E1C3A56FF919158FB5CFE42C64C" ID-GBIF-Occurrence="2847261342" collectionCode="CEMT" country="Brazil" location="MNHN" pageId="15" pageNumber="316" specimenCount="2" specimenCount-female="2" stateProvince="MNHN" typeStatus="paratype">
<emphasis id="27CF88533A56FF919158FB5CFB4CC620" bold="true" box="[1086,1213,796,822]" pageId="15" pageNumber="316">
<typeStatus id="CA00EAE33A56FF919158FB5CFB49C620" box="[1086,1208,796,822]" pageId="15" pageNumber="316" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="6DAC14D13A56FF9191A2FB5CFADFC620" box="[1220,1326,796,822]" name="Brazil" pageId="15" pageNumber="316">BRAZIL</collectingCountry>
: [no data] (
<specimenCount id="03BD9FC83A56FF9195C5FB00FF35C64C" box="[163,196,832,858]" pageId="15" pageNumber="316" type="female">1♀</specimenCount>
<collectingRegion id="D77F9AA33A56FF9195ADFB01FED2C64D" box="[203,291,833,859]" country="Brazil" name="MNHN" pageId="15" pageNumber="316">MNHN</collectingRegion>
, 
<specimenCount id="03BD9FC83A56FF919448FB00FEA0C64C" box="[302,337,832,858]" pageId="15" pageNumber="316" type="female">1♀</specimenCount>
<collectionCode id="73AACC843A56FF91943EFB00FE5AC64C" box="[344,427,832,858]" pageId="15" pageNumber="316">CEMT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A56FF9195A1FB23FB90C668" box="[199,1121,867,895]" pageId="15" pageNumber="316" type="distribution">
<paragraph id="150454413A56FF9195A1FB23FB90C668" blockId="15.[151,1437,327,895]" box="[199,1121,867,895]" pageId="15" pageNumber="316">
<emphasis id="27CF88533A56FF9195A1FB23FE90C668" bold="true" box="[199,353,867,894]" pageId="15" pageNumber="316">Distribution.</emphasis>
Known from 
<collectingCountry id="6DAC14D13A56FF919498FB24FDB3C668" box="[510,578,868,894]" name="Brazil" pageId="15" pageNumber="316">Brazil</collectingCountry>
(
<collectingRegion id="D77F9AA33A56FF919737FB24FD64C669" box="[593,661,868,895]" country="Brazil" name="Bahia" pageId="15" pageNumber="316">Bahia</collectingRegion>
) (
<figureCitation id="8D8048C43A56FF9197CBFB24FCF3C669" box="[685,770,868,895]" captionStart="FIGURE 6" captionStartId="17.[151,250,1784,1809]" captionTargetBox="[179,1376,186,1734]" captionTargetId="figure@17.[174,1414,181,1759]" captionTargetPageId="17" captionText="FIGURE 6. Known distribution of species of Scybalocanthon." figureDoi="http://doi.org/10.5281/zenodo.3995522" httpUri="https://zenodo.org/record/3995522/files/figure.png" pageId="15" pageNumber="316">Fig. 6F</figureCitation>
). 
<emphasis id="27CF88533A56FF919670FB23FC2DC668" bold="true" box="[790,988,867,894]" pageId="15" pageNumber="316">Endemism areas</emphasis>
: Unknown.
</paragraph>
</subSubSection>
</treatment>
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