<document ID-DOI="http://dx.doi.org/10.3897/zookeys.421.7517" ID-PMC="PMC4109472" ID-Pensoft-Pub="1313-2970-421-91" ID-Pensoft-UUID="FFA88030FFBAFF84A232FFF3FF807607" ID-PubMed="25061381" ID-Zenodo-Dep="578387" ID-ZooBank="4DB3DA2D21B14D269544B4008028D304" ModsDocID="1313-2970-421-91" checkinTime="1451245675936" checkinUser="pensoft" docAuthor="Schmidt, B. Christian &amp; Anweiler, Gary G." docDate="2014" docId="A7E6BF4454AD5BEC9C4141EB051867DB" docLanguage="en" docName="ZooKeys 421: 91-113" docOrigin="ZooKeys 421" docPubDate="2014-06-27" docSource="http://dx.doi.org/10.3897/zookeys.421.7517" docTitle="Raphia frater subsp. abrupta Grote 2014, stat. n." docType="treatment" docVersion="3" id="FFA88030FFBAFF84A232FFF3FF807607" lastPageNumber="103" masterDocId="FFA88030FFBAFF84A232FFF3FF807607" masterDocTitle="Taxonomy and biogeography of the Nearctic Raphia Huebner (Lepidoptera, Noctuidae, Raphiinae)" masterLastPageNumber="113" masterPageNumber="91" pageNumber="103" updateTime="1668158807944" updateUser="ExternalLinkService">
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<mods:title id="F42D4C81890B9AB41C4535A4AF02923F">Taxonomy and biogeography of the Nearctic Raphia Huebner (Lepidoptera, Noctuidae, Raphiinae)</mods:title>
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<mods:namePart id="FC00689805D018A212D1C45FF2459006">Schmidt, B. Christian</mods:namePart>
<mods:affiliation id="48BFF36D89618DA45075CC0786103028">Canadian Food Inspection Agency, Canadian National Collection of Insects, Arachnids and Nematodes, K. W. Neatby Bldg., 960 Carling Ave., Ottawa, ON, Canada K 1 A 0 C 6</mods:affiliation>
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<mods:namePart id="3D43334C1E28010668798E377848F7A2">Anweiler, Gary G.</mods:namePart>
<mods:affiliation id="F0D4EC2FEB87B02BA1568EB8ABA97EB4">E. H. Strickland Entomological Museum, 218 Earth Sciences Building, Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada, T 6 G 2 E 9</mods:affiliation>
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<subSubSection id="07AEA73DE9564322778D5B55E41A2B69" pageId="12" pageNumber="103" type="nomenclature">
<paragraph id="16273621916CB735A62A151B4064FC3C" pageId="12" pageNumber="103">
<taxonomicName id="1EC0862C67706F9D41784AED4A78B65D" LSID="A7E6BF44-54AD-5BEC-9C41-41EB051867DB" authority="Grote" authorityName="Grote" authorityYear="2014" baseAuthorityName="Schmidt &amp; Anweiler" baseAuthorityYear="2014" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raphia frater subsp. abrupta" order="Lepidoptera" pageId="12" pageNumber="103" phylum="Arthropoda" rank="subSpecies" species="frater" status="stat. n." subSpecies="abrupta">Raphia frater abrupta Grote</taxonomicName>
<taxonomicNameLabel id="288604F75FD092223DE9301A169B7386" pageId="12" pageNumber="103">stat. n.</taxonomicNameLabel>
<figureCitation id="DE18C6960D78314503988F14EF54E9C4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographic distribution and phenotypic variation of Raphia frater subspecies. Circles indicate specimens examined during this study: green - subsp. piazzi; pink - subsp. abrupta; black - subsp. frater; blue - subsp. coloradensis; yellow - subsp. cinderella. Multi-coloured circles indicate transitional populations and / or phenotypically intermediate specimens between respective subspecies. a Raphia frater piazzi (Zavallo Co., TX) b Raphia frater abrupta (Oktibeha Co., MS) c Raphia frater abrupta (Cottle Co., TX) d Raphia frater abrupta (Cottle Co., TX) e Raphia frater abrupta (Montgomery Co., MD) f, g Raphia frater frater (Edmunston, NB) h Raphia frater abrupta - frater intermediate (Anne Arundel Co., MD) i Raphia frater abrupta - frater - coloradensis intermediate from highly variable population in Cherry Co., NE j Raphia frater coloradensis (Alamosa Co., CO) k Raphia frater coloradensis (Milk River valley, AB) l Raphia frater coloradensis (Sanpete Co., UT) m Raphia frater coloradensis (Elko Co., NV) n Raphia frater cinderella (Ventura Co., CA) o, p Raphia frater coloradensis - frater intermediates (Chelan Co., WA) q Raphia frater elbea (Cochise Co., AZ) r Raphia frater elbea (San Juan Co., UT) s Raphia frater elbea (Santa Cruz Co., AZ). All specimens are males." httpUri="https://binary.pensoft.net/fig/30255" pageId="12" pageNumber="103">Figs 1b-e</figureCitation>
<figureCitation id="4557F2057DA627DAB9D69A57E134E927" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of Raphia frater subspecies (circles) relative to range of Populus larval host plants (coloured shading) in eastern North America. Black circles - subsp. frater; white circles - subsp. abrupta; grey circles - subsp. piazzi; half-circles represent transitional populations and / or phenotypically intermediate specimens. Blue shading - combined ranges of Populus tremuloides and Populus grandidentata; yellow shading - Populus deltoides; range overlap depicted in green. Populus ranges adapted from USGS (2013)." httpUri="https://binary.pensoft.net/fig/30256" pageId="12" pageNumber="103">, 2</figureCitation>
</paragraph>
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<subSubSection id="1C66893ABF63D3EA63A987DA66D1352E" pageId="12" pageNumber="103" type="reference_group">
<paragraph id="B6539A05DFD5BE080298C25A4671EE9E" pageId="12" pageNumber="103">
<taxonomicName id="01BEAA6CD99F813FBC8EEAFE91F69C71" authorityName="Schmidt &amp; Anweiler" authorityYear="2014" baseAuthorityName="Grote" baseAuthorityYear="2014" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="12" pageNumber="103" phylum="Arthropoda" rank="species" species="abrupta" status="stat. n.">Raphia abrupta</taxonomicName>
Grote, 1864
</paragraph>
<paragraph id="92C3090FAD72D137B76F3F276A4600D6" pageId="12" pageNumber="103">
<taxonomicName id="2B2BC6B28509D3AAA27E2E15F99DE70C" authorityName="Schmidt &amp; Anweiler" authorityYear="2014" class="Insecta" family="Noctuidae" genus="Certila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="12" pageNumber="103" phylum="Arthropoda" rank="species" species="flexuosa" status="stat. n.">Certila flexuosa</taxonomicName>
Walker, 1865
</paragraph>
</subSubSection>
<subSubSection id="E7F8F518E821CB7B15E2F54F62723402" pageId="12" pageNumber="103" type="type material">
<paragraph id="2FDF46DCB91AC510E8D44DF2B2C50901" pageId="12" pageNumber="103">Type material.</paragraph>
<paragraph id="BA4B52F7E55A8CCD5F51905345F7B4D4" pageId="12" pageNumber="103">
<taxonomicName id="89A782A9E0D2FFBBF63E527243EF4749" authorityName="Schmidt &amp; Anweiler" authorityYear="2014" baseAuthorityName="Grote" baseAuthorityYear="2014" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="12" pageNumber="103" phylum="Arthropoda" rank="species" species="abrupta">
<emphasis id="8ADC8E08739FB9B8986136A261A18530" italics="true" pageId="12" pageNumber="103">Raphia abrupta</emphasis>
</taxonomicName>
- female holotype # 7675 [ANSP]. Type locality: not given; here restricted to Sycamore Landing, Seneca, Montgomery Co., Maryland. The female type bears no locality or collector label data, and since this is a widespread, geographically variable taxon, we restrict the type locality to Sycamore Landing, Seneca, Montgomery Co., Maryland; a series in USNM from this locality, collected by D. C. Ferguson, is phenotypically more similar to the female type than specimens from the Great Plains; it is also more likley that the holotype originated from the eastern US rather than the Great Plains, which were not well collected in the mid 1800's.
</paragraph>
<paragraph id="7494F1598A144E0251FA5C574C7F6BD6" pageId="12" pageNumber="103">
<taxonomicName id="B59D4E64C59E43F8B64DAA4A3F9A220E" authorityName="Schmidt &amp; Anweiler" authorityYear="2014" class="Insecta" family="Noctuidae" genus="Certila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="12" pageNumber="103" phylum="Arthropoda" rank="species" species="flexuosa">
<emphasis id="429247C039077769AC63BFF662CC5007" italics="true" pageId="12" pageNumber="103">Certila flexuosa</emphasis>
</taxonomicName>
Walker - [BMNH; not examined]. Type locality: North America.
</paragraph>
</subSubSection>
<subSubSection id="C4B7030926EDD3E9FA85EAEBC8F83554" pageId="12" pageNumber="103" type="diagnosis and description">
<paragraph id="BB5955523AB52DACABFC2C9442C65F76" pageId="12" pageNumber="103">Diagnosis and description.</paragraph>
<paragraph id="67FBE26801C6A9E51B126450AFD8249D" pageId="12" pageNumber="103">
<taxonomicName id="290FFBE56019EAA0408017BC623A898F" authorityName="Grote" authorityYear="2014" baseAuthorityName="Schmidt &amp; Anweiler" baseAuthorityYear="2014" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="12" pageNumber="103" phylum="Arthropoda" rank="subSpecies" species="frater" subSpecies="abrupta">
<emphasis id="77B0DBB10D5FB16B9CF619F162120EDB" italics="true" pageId="12" pageNumber="103">Raphia frater abrupta</emphasis>
</taxonomicName>
replaces 
<taxonomicName id="668E88F0A6CCC945606B9671E71DC3CA" authority="frater" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="12" pageNumber="103" phylum="Arthropoda" rank="subSpecies" species="frater" subSpecies="frater">
<emphasis id="D4C2CD01132484CC041822810A3AFF03" italics="true" pageId="12" pageNumber="103">Raphia frater frater</emphasis>
</taxonomicName>
from the central Great Plains eastward to the mid-Atlantic seaboard, and southward to eastern Texas and Florida. It is on average smaller with a more evenly-coloured forewing, a more linear, angulate antemedial band and a fuscous hindwing. Average forewing length is 13.7 mm (
<emphasis id="C5A78DEA03967B87E4152879BA099086" italics="true" pageId="12" pageNumber="103">n</emphasis>
= 9) in males, 15.2 mm in females (
<emphasis id="5798B6D5D768DA914C8196EC39E7E824" italics="true" pageId="12" pageNumber="103">n</emphasis>
= 9). The thoracic collar is often darker than the dorsal thorax, not concolorous as in 
<taxonomicName id="A4F4C168AE77ABCB1226ADE230F49A7F" authority="frater" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="12" pageNumber="103" phylum="Arthropoda" rank="subSpecies" species="frater" subSpecies="frater">
<emphasis id="3D2AE73328884727009BC7C8099DCDB8" italics="true" pageId="12" pageNumber="103">Raphia frater frater</emphasis>
</taxonomicName>
. The wing facies of subspecies 
<emphasis id="9394CD61BEDDC491F2C051B2488E99B3" italics="true" pageId="12" pageNumber="103">abrupta</emphasis>
is in many ways intermediate between 
<taxonomicName id="24699A12790AF45A71B5215933F8E363" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="12" pageNumber="103" phylum="Arthropoda" rank="subSpecies" species="frater" subSpecies="piazzi">
<emphasis id="CDB64CE363144E18D9EC85B173954D3A" italics="true" pageId="12" pageNumber="103">Raphia frater piazzi</emphasis>
</taxonomicName>
of central and southern Texas and 
<taxonomicName id="2D3E8C0AD42FFAF768B76E42856909B6" authority="frater" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="12" pageNumber="103" phylum="Arthropoda" rank="subSpecies" species="frater" subSpecies="frater">
<emphasis id="406533F7988F9316BFBB194D20687328" italics="true" pageId="12" pageNumber="103">Raphia frater frater</emphasis>
</taxonomicName>
to the north, but the exact nature of the interface between 
<emphasis id="51B03F155343F7DA5CD0B797051E712D" italics="true" pageId="12" pageNumber="103">abrupta</emphasis>
and 
<emphasis id="3681A2F130D9C5BCB34C01CA5BB8DDDA" italics="true" pageId="12" pageNumber="103">piazzi</emphasis>
in Texas remains unstudied.
</paragraph>
</subSubSection>
<subSubSection id="799979E701887B38FC73D42D889C5C60" pageId="12" pageNumber="103" type="biology and distribution">
<paragraph id="8F522BC8E06C31AE98B9748DBEB7F08B" pageId="12" pageNumber="103">Biology and distribution.</paragraph>
<paragraph id="802D1BFCE9CBDC60B477EAAEA887D1C9" pageId="12" pageNumber="103">
Subspecies 
<emphasis id="A55693139829D1315FCBD8C8BED4177D" italics="true" pageId="12" pageNumber="103">abrupta</emphasis>
occurs south of the range of the aspen species favoured by 
<taxonomicName id="419551B2FADDAD8493D5F6A3F19157D1" authority="frater" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="12" pageNumber="103" phylum="Arthropoda" rank="subSpecies" species="frater" subSpecies="frater">
<emphasis id="82BC39AD8EC8FED4DA7BF972AB2E74B0" italics="true" pageId="12" pageNumber="103">Raphia frater frater</emphasis>
</taxonomicName>
larvae, and its riparian haunts suggest it feeds on eastern cottonwood (
<taxonomicName id="F35A09762A35A10CD3FA7F7E4BEEA2A9" authorityName="W.Bartram ex Marshall" authorityYear="1785" class="Dicotyledoneae" family="Salicaceae" genus="Populus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Salicales" pageId="12" pageNumber="103" phylum="Angiospermae" rank="species" species="deltoides">
<emphasis id="FA9AB234DAEF70AD8EF558B266DDE082" italics="true" pageId="12" pageNumber="103">Populus deltoides</emphasis>
</taxonomicName>
), the only 
<taxonomicName id="3215266190B3278245BBDD259BAA10C4" class="Dicotyledoneae" family="Salicaceae" genus="Populus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Salicales" pageId="12" pageNumber="103" phylum="Angiospermae" rank="genus">
<emphasis id="2E5F0FF423E3503556F7F32D3F931B47" italics="true" pageId="12" pageNumber="103">Populus</emphasis>
</taxonomicName>
species in much of its range. Swamp cottonwood (
<taxonomicName id="FF4399A05545C0E203C7820020748A37" class="Dicotyledoneae" family="Salicaceae" genus="Populus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Salicales" pageId="12" pageNumber="103" phylum="Angiospermae" rank="species" species="heterophylla">
<emphasis id="B6A84FF957260DDAFDBF25343C19D5B0" italics="true" pageId="12" pageNumber="103">Populus heterophylla</emphasis>
</taxonomicName>
) and willows (
<taxonomicName id="115B8338298A44402210DF3405F40736" class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Aplousobranchia" pageId="12" pageNumber="103" phylum="Chordata" rank="genus">
<emphasis id="FF6AC5A8E0E1C76A370FC73AD38105B5" italics="true" pageId="12" pageNumber="103">Salix</emphasis>
</taxonomicName>
spp.) may also be suitable hosts. This subspecies is apparently rare on the Atlantic seaboard and absent altogether in the Appalachians. We examined only a single historical specimen from New Jersey (Trenton), with records north of there assignable to 
<taxonomicName id="D0BCDF6463ADBF65202265B6BFC3D6F2" authority="frater" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="12" pageNumber="103" phylum="Arthropoda" rank="subSpecies" species="frater" subSpecies="frater">
<emphasis id="EEB58FA91DD939B1BB3A28C75405A8C9" italics="true" pageId="12" pageNumber="103">Raphia frater frater</emphasis>
</taxonomicName>
. All Ohio records were attributed to 
<taxonomicName id="AFA18CAD94EFC045E2BF3757883F998B" authority="frater" class="Insecta" family="Noctuidae" genus="Raphia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="12" pageNumber="103" phylum="Arthropoda" rank="subSpecies" species="frater" subSpecies="frater">
<emphasis id="1A0F52FE697F855FD60700ED044CBAA4" italics="true" pageId="12" pageNumber="103">Raphia frater frater</emphasis>
</taxonomicName>
by 
<bibRefCitation id="3342315219294447A14C99276017AD9A" author="Rings, RW" journalOrPublisher="Ohio Biological Survey Bulletin (New Series)" pageId="18" pageNumber="109" pagination="1 - 219" refId="B36" refString="Rings, RW, Metzler, EH, Arnold, FJ, Harris, DH, 1992. The owlet moths of Ohio Order Lepidoptera Family Noctuidae. Ohio Biological Survey Bulletin (New Series) 9 (2): 1 - 219" title="The owlet moths of Ohio Order Lepidoptera Family Noctuidae." volume="9" year="1992">Rings et al. (1992)</bibRefCitation>
, although specimens with a pale grey forewing and dusky hindwing, traits of the 
<emphasis id="A6173FAF0DCE25448DC56CC18348DA14" italics="true" pageId="12" pageNumber="103">abrupta</emphasis>
phenotype, rarely occur as far north as southernmost Ontario (Toronto) and southeastern Minnesota (Fillmore Co.).
</paragraph>
</subSubSection>
</treatment>
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