A new species of Succinoraphidia Aspöck & Aspöck, 2004 (Raphidioptera: Raphidiidae) from the late Eocene Rovno amber, with venation characteristics of the genus
Author
Perkovsky, Evgeny E.
Author
Makarkin, Vladimir N.
text
Zootaxa
2019
2019-04-03
4576
3
570
580
journal article
10.11646/zootaxa.4576.3.9
555f42f9-9135-45a6-b7aa-c0b4c064c7db
1175-5326
3715838
4994B0F7-3CB4-4DDE-99CB-BCD061D5730C
Succinoraphidia
and ‘Mesoraphidiidae’
It is now clear that the Mesozoic ‘Mesoraphidiidae’ is a paraphyletic group, which includes in particular the stem lineages leading to extant
Raphidiidae
and
Inocelliidae (
Makarkin & Archibald 2014
)
. The venation of
Succinoraphidia
is more similar to that of some ‘Mesoraphidiidae’ than the venation of any genus of
Raphidiidae
. Moreover, there are a few mesoraphidiids in which all diagnostic character states of
Succinoraphidia
are found (except for the structure of the pterostigma). It is characteristic of some minute species from the early Late Cretaceous, e.g.,
Grimaldiraphidia luzzii
(
Grimaldi, 2000
)
from the Turonian New
Jersey
amber;
Nanoraphidia electroburmica
Engel, 2002
and
Burmoraphidia reni
Liu
et al
., 2016
from the early Cenomanian Burmese amber. These genera are considered to belong to the tribe
Nanoraphidiini
(
Bechly & Wolf-Schwenninger 2011
;
Liu
et al
. 2016
). But the structure of their pterostigma differs from that of
Succinoraphidia
. It lacks the crossvein 2scp-r at the proximal end of the pterostigma (
B. reni
) or incorporated branches of RA within pterostigma (
G. luzzii
and
N. electroburmica
). However, the pterostigma of some other ‘Mesoraphidiidae’ from Burmese amber (e.g.,
Dolichoraphidia engeli
Liu
et al
., 2016
) is very similar to that of
Raphidiidae
with a long pterostigma, including
Succinoraphidia
, i.e., it possesses the crossvein 2scp-r at the proximal end of the pterostigma and the incorporated branch of RA within the pterostigma (see
Liu
et al
. 2016
:
Fig. 4
). The venation of
Dolichoraphidia engeli
differs from that of
Raphidiidae
only by the number of discoidal cells: one in
D. engeli
and normally two in all
Raphidiidae
, except some species, which have one discoidal cell in the hind wings, e.g., those of
Harraphidia
Steinmann, 1963
(see
Monserrat & Papenberg 2006
:
Figs 1–4
).
Judging from great similarity of the venation of
Nanoraphidiini
(and similar genera of the Late Cretaceous ‘Mesoraphidiidae’) and
Succinoraphidia
, among the former may be present a stem group leading to
Raphidiidae
.