Order Chiroptera - Family Vespertilionidae Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1 451 529 book chapter 0-8018-8221-4 10.5281/zenodo.7316519 Myotis nigricans subsp. osculatii Cornalia 1849 Synonyms: Myotis nigricans subsp. caucensis Miller and G. M. Allen 1928 ; Myotis nigricans subsp. quixensis Osculati 1854 .