A molecular phylogeny and revision of the genus Pyropteron Newman, 1832 (Lepidoptera, Sesiidae) reveals unexpected diversity and frequent hostplant switch as a driver of speciation
Author
Bartsch, Daniel
daniel.bartsch@smns-bw.de
Author
Pühringer, Franz
f.puehringer@sesiidae.net
Author
Milla, Liz
liz.milla@csiro.au
Author
Lingenhöle, Arthur
arthur.lingenhöle@gmx.de
Author
Kallies, Axel
0000-0002-3778-2187
axel.kallies@unimelb.edu.au
text
Zootaxa
2021
2021-05-18
4972
1
1
75
journal article
6247
10.11646/zootaxa.4972.1.1
cdd9688e-9a3f-4602-992e-1f8693caab9b
1175-5326
4771798
E78473FE-5662-409A-90C2-7C4912DC57E8
Pyropteron leucomelaena leucomelaena
(
Zeller, 1847
)
(
Figs 120–122
,
221, 222
,
229, 230
)
Sesia leucomelaena
Zeller, 1847
: Isis, 40 (1): 12.
Lectotype
♂
:
Macri
[
Turkey
, Fethiye] (NHMUK).
Sesia therevaeformis
Lederer, 1853
: Verhandlungen des zoologisch-botanischen
Vereins in Wien
, Abhandlungen 2 [“1852”], 2: 82.
Syntypes
:
2♂
(?),
4♀
,
Spalato
[
Croatia
, Split] and
Brussa
[
Turkey
,
Bursa
] (lost?
Not
in MFNB).
Sesia cretica
Rebel, 1916
: Annalen des k.k. Naturhistorischen Hofmuseums, 30: 143.
Syntypes
:
2♀
,
Greece
,
Crete
,
Kristallenia
and
Mallaes
(NHMW).
Chamaesphecia leucomelaena
f.
cyrnea
Le Cerf, 1920
:
Études de Lépidoptérologie Comparée
, 17: 506.
Syntypes
:
15♂
,
5♀
,
France
, Corsica (MNHN).
Distribution.
The typical subspecies is known from southern
France
in the west across southern Europe to
Turkey
in the east. It also occurs on the Mediterranean islands of
Corsica
, Sardinia, Sicily, Crete and
Cyprus
(
Laštůvka & Laštůvka 1995
,
2001
,
Špatenka
et al.
1999
). Based on Gorbunov (pers. communication), this species is absent from Crimea. Specimens from central and northern
Spain
are here provisionally assigned to the typical subspecies; however, more detailed analyses are needed to map the precise distribution of the two subspecies of
P. leucomelaena
in the Iberian Peninsula.
Specimens examined.
17♂
,
Spain
,
Huesca
,
Graus
,
30.
VI
.–10.
VII
.2009, leg.
Riefenstahl
&
Wagenblass
(
Bartsch
gen. prep. 2020-04) (
Fig. 222
)
;
2♂
,
Spain
,
Teruel
,
Frias
/
Albarracin
,
1400m
,
1.–10.
VII
.1999, leg.
Riefenstahl
&
Wagenblass
(SMNS);
1♂
,
Spain
, Castillia-La-Mancha,
Umgeb. Los Alares
,
26.
VI
.
2012, 580m
, leg. AL;
1♀
, same locality,
19.
VI
.
2012, 580m
, leg. AL;
1♂
,
Spain
, Castilla-La-Mancha,
Horcaja
de los
Montes
,
435m
,
18.
VI
.2012, leg. AL;
1♀
, same locality,
19.
VI
.2012, leg. AL (CAL);
6♂
,
Spain
,
Teruel
,
Portillo de Guadalaviar
(
El Portillo
),
1760–1770 m
,
9.
VII
.2018, leg. FP (CFP);
1♂
,
Spain
,
Teruel
,
Albarracin
,
1200m
,
30.
VI
.1993, leg. DB;
1♂
,
Spain
,
Huesca
,
Graus
,
500m
,
30.
VI
.2009, leg. AL (
Bartsch
gen. prep. 2019-52) (
CDB
)
;
1♂
,
Spain
,
Teruel
(
Aragon
), Albarracin,
25.
VI
.1928, leg. Predota;
1♂
,
Spain
,
Catalonia
,
Vidreras
nr.
Gerona
,
12.
VI
.1993, leg. A. Pürstinger;
1♂
, same data,
15.
VI
.1993 (
CFP
)
.
1♂
,
France
,
Ardeche
, Saint-Maurice-d’Ibie,
350 m
,
6.
VII
.1998, leg. B. Vanholder (CCDB-02110 B07, gen. prep. FP 98/64);
1♂
,
France
, Corsica,
Castirla
NW
Corte
,
950 m
,
10.
VI
.1999, leg.
H. Fischer
(BOX-2219 H11, gen. prep. FP 99/11) (CFP);
1♀
,
France
,
Alpes de Haut Provence
,
Luberon
,
Manosque
,
14.
VI
.2006, leg.
E. Blum
(
Bartsch
gen. prep. 2019-53) (
Fig. 230
) (
CDB
)
.
5♂
,
Provence
,
Montbrun les Bains
, 6.,
7. and 8.
VII
.1991, leg. DB;
1♀
,
France
,
Drome
,
Séderon
,
Col de la Pigiere
,
25.
VII
.1992, leg. DB;
1♂
,
France
,
Provence
,
Esterel
,
Agay
,
25.
VI
.1991, leg. DB (CDB).
3♂
,
Italy
,
Sicily
, PA,
N Castelbuono
,
120 m
,
10.
VI
.2001, leg. FP (DNAtax 02-18, gen. prep. FP 02/08);
12♂
,
Italy
,
Sicily
, SR,
Pedagaggi NW Sortino
,
700 m
,
15.
VI
.2001, leg. FP (DNAtax 02-19, gen. prep. FP 02/06);
2♂
,
Italy
,
Sicily
, SR,
W Sortino
,
530 m
,
13.
VI
.2001, leg. FP;
2♂
, ibid.,
600 m
,
15.
VI
.2001, leg. FP (BOX-2219 H12, gen. prep. FP 02/09) (CFP);
1♂
,
Italy
,
Sicily
,
W of Sortino
,
600m
,
15.
VI
.2001, leg. FP;
1♂
,
Italy
,
Abruzzi
,
Tuffo
,
800m
,
22.
VII
.1991, leg. HR (CRS);
1♂
,
Albania
,
Lezha
,
W Reps
,
260 m
,
8.
VII
.2016, leg. FP;
1♂
,
Bulgaria
,
Burgas
,
Chernomorets
,
30 m
,
9.
VII
.2014, leg. FP;
5♂
,
Bulgaria
,
Slivno, N
Sliven
(
Slivno
),
390 m
,
3.
VII
.2014, leg. FP;
2♂
, ibid.,
400 m
,
10.
VII
.2014, leg. FP;
1♂
, ibid.,
25.
VI
.2016, leg. FP;
1♂
,
Greece
,
Crete
,
Chania
,
Marathi
(
Halbinsel
Akrotiri
),
10 m
,
6.
VI
.2000, leg. FP;
1♂
, ibid.,
50 m
,
7.
VI
.2000, leg. FP (CCDB-02113 F04);
2♂
,
Greece
,
Crete
,
Rethymnon
,
Skaleta E Rethymnon
,
2.
VI
.1996, leg. FP (photo 24/1/40-42, 2/35-41);
1♂
, ibid.,
4.
VI
.1996, leg. FP;
6♂
, ibid.,
6.
VI
.2000, leg. FP (CCDB-02110 B10, photo 41/6/1- 4);
2♂
, ibid.,
7.
VI
.1996, leg. FP;
1♂
,
Greece
,
Fthiotis
,
5 km
N Bralos
,
450 m
,
30.
VI
.2002, leg. FP (DNAtax 01-16);
3♂
, ibid.,
3.
VII
.2003, leg. FP (BOX-2221 F11);
1♂
,
Greece
,
Fthiotis
,
Iti
(
Oiti
),
670 m
,
22.
VI
.2002, leg. FP (photo 51/5/33-41);
1♂
,
Greece
,
Fthiotis
,
Pavliani
,
1000 m
,
22.
VI
.2002, leg. FP;
5♂
,
Greece
,
Ioannina
,
Kipi
(=
Kipoi
,
Timfi Mts
),
780 m
,
28.
VI
.2002, leg. FP (DNAtax 02502, photo 52/5/11-13);
3♂
,
Greece
,
Ioannina
,
Koukouli
,
930 m
,
30.
VI
.2003, leg. FP (CCDB-02110 B08);
1♂
,
Greece
,
Komotini
,
N Drama
,
350 m
,
6.
VII
.2003, leg. FP;
1♂
,
Greece
,
Larissa
,
Ossa Mts
,
SW Stomio
(n),
1000–1090 m
,
20.
VI
.2002, leg. FP;
5♂
, ibid.,
2.
VII
.2003, leg. FP (BOX- 2220 A05) (CFP);
1♂
,
Greece
,
Peloponnese
, inner
Mani
,
Kendro
,
1200–1800m
,
7.
VII
.2005, leg. DB;
1♀
(
Fig. 122
),
Greece
,
Peloponnese
, outer
Mani
,
Kariovouni
,
600m
, e.l. ex.
Sanguisorba minor
, larva
19.IX.2000
adult
30.
V
.2001, leg. DB (CDB);
1♂
,
Bulgaria
,
Vlas
,
27.
VI
.–3.
VII
.2005, leg. Riefenstahl;
1♂
,
Turkey
,
Antalya
,
Saklikent
,
640m
,
28.
VI
.2002, leg. Riefenstahl;
2♂
, ibid.,
1380m
,
28.
VI
.2002, leg. Riefenstahl;
9♂
, ibid.,
900m
,
28.–29.
VI
.2002, leg. Riefenstahl;
2♂
, ibid.,
1610m
,
29.–30.
VI
.2002, leg. Riefenstahl;
10♂
,
Turkey
, Anatolia,
Feke
,
1000m
,
5.
VII
.1992, leg.
Riefenstahl
(SMNS);
7♂
,
Turkey
,
Toros Daglari
, road
Antalya-Saklikent
,
1000m
,
29. and 30.
VI
.1998, leg. DB;
7♂
(
Figs 120–121
), ibid.,
600m
,
16.
VI
. and 3.
VII
.2002, leg. DB;
1♂
,
Turkey
,
Mersin
,
Pusatli Dagi
,
10 km
S Kirobasi,
1260m
,
29.
VI
.2002, leg. AL (
Bartsch
gen. prep. 2019-50) (
Fig. 221
)
;
9♂
, ibid.,
1280m
,
29.
VI
.2002, leg. DB;
1♀
,
Turkey
,
Konya
,
Beysehir
,
1170m
,
27.
VI
.2002, leg. AL (
Bartsch
gen. prep. 2019-51) (
Fig. 230
)
;
2♂
,
Konya
,
Taskent
,
1570m
,
1.
VII
.2002, leg. DB;
2♂
,
Turkey
,
Konya
,
45km
S Beysehir
,
Bakaran
,
1420m
,
18.
VI
. and 2.
VII
.2002, leg. DB;
1♂
,
1♀
,
Turkey
,
Adiyaman
,
25km
W Gölbasi
,
24.
VI
. 1993, leg. G. & M. Novak;
2♂
,
Cyprus
,
Limassol
,
Souni
,
400m
,
19.
V
.2001, leg. G. & M. Novak;
1♂
,
Cyprus
,
Paphos
,
Akamas
,
Ag. Minas
,
23.
V
.2001, leg. G. & M. Novak;
1♂
,
2♀
,
Cyprus
,
Troodos
,
2km
NW Fioni
,
1200m
, larvae
25.
III
.2000, e.l. ex.
P. minor
, leg. DB (CDB).